Canthon kästneri Balthasar, 1939: 193
Vulcano & Pereira 1967: 552
Canthon kaestneri : Blackwelder 1944: 199
Pereira & Martínez 1956: 114
Krajcik 2006: 28
Carvajal et al . 2011: 314
Krajcik 2012: 63
Scybalocanthon kästneri : Vulcano & Pereira 1964: 616
Scybalocanthon kaestneri: Halffter & Martínez 1977: 68
Carvajal et al . 2011: 314
Bezdek & Hajek 2011: 363
Chamorro et al . 2018: 87
Chamorro et al. 2019: 229
A taxonomic revision of the genus Scybalocanthon Martínez, 1948 (Coleoptera Scarabaeidae: Scarabaeinae: Deltochilini)
Silva, Fernando A. B.
Valois, Marcely
Zootaxa
2019
2019-07-08
4629
3
301
341
Chamorro et al. 2019: 229
4W3LK
(Balthasar, 1939)
Balthasar
1939
[151,683,1197,1224]
Insecta
Scarabaeidae
Scybalocanthon
Animalia
Coleoptera
21
322
Arthropoda
species
kaestneri
( Figs. 1K, 3E, 4K, 5J, 6E)
Canthon kästneri Balthasar, 1939: 193, 227 (original description); Vulcano & Pereira 1967: 552(distribution, identification key). Canthon kaestneri: Blackwelder 1944: 199(checklist); Pereira & Martínez 1956: 114(comment); Krajcik 2006: 28(catalog); Carvajal et al. 2011: 314(cited for Ecuador); Krajcik 2012: 63(catalog). Scybalocanthon kästneri: Vulcano & Pereira 1964: 616(distribution). Scybalocanthon kaestneri: Halffter & Martínez 1977: 68(checklist); Carvajal et al. 2011: 314(cited for Ecuador); Bezdek & Hajek 2011: 363(catalog of types); Chamorro et al. 2018: 87(fig. 10B), 98 (cited for Ecuador); Chamorro et al.2019: 229, 230 (catalog of dung beetles species from Ecuador).
Diagnosis. Specimens of S. kaestneriare readily distinguishable from all other species by pronotum with four black spots (two central spots with triangular shape and two lateral spots rounded) ( Fig. 2K); femora almost completely yellow or brown, with black spots only on the tips ( Fig. 3G); parameres of aedeagus slightly asymmetrical, dorsal margin of left paramere strongly curved outward at apical portion ( Fig. 4K); endophallus with bristles right beside the FLP sclerite ( Fig. 5J); and additional sclerite ( AS) absent ( Fig. 5J). A specimen from Napo, Ecuador, has only three black spots on pronotum (one spot at posterior margin with triangular shape and two lateral spots rounded).
Description. Body.Oval-elongate, with variable metallic sheen. Surface completely microgranulate. Color. Head and elytra opaque, pronotum with metallic sheen. Most of pronotum, hypomera, prosternum, mesoventrite, central portion of metaventrite, metaepisternae, lateral portion of abdominal ventrites, pygidium, and legs yellow or light brown. Head, some spots of pronotum, elytra, and central portion of abdominal ventrites black. Length.9.0–10.0 mm. Thorax. Anterior angles of pronotum approximately 90°. Lateral margin strongly curved outward at the middle portion. Elytra. Striae thin and impressed, with elliptical punctures. Eighth stria with a thin carina at the anterior portion. Aedeagus.Parameres slightly asymmetrical, obliquely truncate apically ( Fig. 4K). Dorsal margin of parameres curved inward from the basal to medial portions, and curved outward at the apical portion. Ventral margin of parameres curved inward at the basal portion and substraight from the medial to apical portions. SRP circular, with rounded handle-shaped extension ( Fig. 5J). FLP subelliptical, with dense set of bristles right beside it ( Fig. 5J). A+SA with two superposed and elongate sclerites ( Fig. 5J).
Type material. Canthon kaestneriBalthasar, lectotype ♀( here designated; NMPC): “ Equador” // “TY- PUS” // “ Kastneri / m.” // “ Scybalocanthon/ kaestneri (Balth.)/ Det. B. D. Gill’96” // “ LECTOTYPE/ Canthon ♀/ kaestneri/ Balth/ des. F. Z. Vaz-de-Mello/ 2013”. Paralectotypes 2♀( NMPC): “Mera / Ecuador” // “Typus” // “ PARALECTOTYPE/ Canthon ♀/ kaestneri Balth./ des. F. Z. Vaz-de-Mello”. Non-type material [ 18 males, 36 females]: ECUADOR: MORONA-SANTIAGO, Untsuantssitio 4, 1100 m, 20.i.2002, J. Celi& M. Ortega, pitfall trap, human feces— 1♂ 1♀( CMNC); Untsuantssitio 6, 1100 m, 23.i.2002, J. Celi& M. Ortega, pitfall trap, fish— 1♀( CMNC); Untsuantssitio 7, 900 m, 23.i.2002, J. Celi& M. Ortega, pitfall trap, human feces— 1♀( CMNC); Bosque Domono, 22.viii.1997, 1650 m, C. Carpio— 1♂ 1♀( CEMT); PASTA- ZA, 22 kmSE Puyo, 900 m( 1°37’11’’S, 77°50’40’’W), 12–16.vii.1976, S. Peck, forest, dung traps( dung traps39–41)— 1♀( CMNC); 22 kmSE Puyo, 900 m( 1°37’11’’S, 77°50’40’’W), 12–16.vii.1976, S. Peck, forest, carrion traps( carrion traps42–43)— 1♂( CMNC); 25 kmNNE Puyo, 1000 m( 1°18’19’’S, 77°52’59’’W), 4–13.vii.1976, S. Peck, forest, dung traps( dung trap25-27)— 3♂ 16♀( CMNC, MZUFPA); 9 kmESE Veracruz, 900 m( 1°33’49’’S, 77°53’8’’W), 22–24.viii.1975, R. Webster, forest, dung traps— 1♂ 1♀( CMNC); Llandia, 17 kmN Puyo, 1000 m( 1°21’3’’S, 77°58’4’’W), 16.vii.1994, F. Génier, remnant rainforest, hand collecting— 1♂( CMNC); Llandia, 17 kmN Puyo, 1000 m( 1°21’3’’S, 77°58’4’’W), 19.vii.1994, F. Génier, remnant rainforest, feces trap— 4♂ 3♀( CMNC); Llandia, 17 kmN Puyo, 1000 m( 1°21’3’’S, 77°58’4’’W), 20.vii.1994, F. Génier, remnant rainforest, feces trap— 1♀( CMNC); Oglán River, 29.x. 2008, 809 m, F. Gallo& E. Camillo— 1♀( CEMT); Mera, Estação Biológica Pinto MiradorUTE [Usina Termelétrica] ( 78°06’29”W, 1°27’25”S), 22.iii.2012, pitfallpig feces, 1000 m, C. Luzuriaga— 5♂ 9♀( CEMT); Mera, Estação Biológica Pinto MiradorUTE [Usina Termelétrica] ( 78°1’00”W, 2°31’59”S)— 1♂( CEMT); NAPO: Puerto Napo, 480 m—2 [unsexed specimens] ( MQCAZ).
Distribution.Known from Ecuador( Fig. 6E). Endemism areas: Brazilian sub-region: Boreal Brazilian dominion: Napo province(see Morrone 2014; fig. 12).
2847261339
NMPC
Det. B. D. Gill' & F. Z. Vaz-de-Mello
Ecuador
Det. B. D. Gill'
Equador
22
323
2
2
lectotype
2847261510
NMPC
F. Z. Vaz-de-Mello
Ecuador
Ecuador
22
323
3
3
paralectotype
2847261343
2002-01-20
CMNC
J. Celi & M. Ortega
Ecuador
1100
Untsuants
22
323
56
37
19
Morona-Santiago
2847261416
2002-01-23
CMNC
J. Celi & M. Ortega
Ecuador
1100
Untsuants
22
323
1
1
Morona-Santiago
2847261450
2002-01-23
CMNC
J. Celi & M. Ortega
Ecuador
7900
Untsuants
22
323
1
1
Morona-Santiago
2847261549
[546,1329,548,575]
1997-08-22
CEMT
Ecuador
Bosque Domono
22
323
2
1
1
Morona-Santiago
2847261344
1976-07-12
1976-07-16
1976-07-12
CMNC
S. Peck
Ecuador
900
-1.6197222
Puyo
21
-77.844444
22
323
1
1
Morona-Santiago
2847261518
1976-07-12
1976-07-16
1976-07-12
CMNC
S. Peck
Ecuador
900
-1.6197222
Puyo
21
-77.844444
22
323
1
1
Morona-Santiago
2847261608
1976-07-04
1976-07-13
1976-07-04
CMNC, MZUFPA
S. Peck
Ecuador
1000
-1.3052777
Puyo
21
-77.88306
22
323
19
16
3
Morona-Santiago
2847261614
1975-08-22
1975-08-24
1975-08-22
CMNC
R. Webster
Ecuador
900
-1.563611
Veracruz
21
-77.88555
22
323
2
1
1
Morona-Santiago
2847261472
1994-07-16
CMNC
F. Genier
Ecuador
1000
-1.3508334
Llandia
21
-77.96778
22
323
1
1
Morona-Santiago
2847261316
1994-07-19
CMNC
F. Genier
Ecuador
1000
-1.3508334
Llandia
21
-77.96778
22
323
7
3
4
Morona-Santiago
2847261356
1994-07-20
CMNC
F. Genier
Ecuador
1000
-1.3508334
Llandia
21
-77.96778
22
323
1
1
Morona-Santiago
2847261494
[276,1075,873,899]
2008-10-29
CEMT
F. Gallo
Ecuador
Oglan River
22
323
1
1
Morona-Santiago
2847261320
2012-03-22
CEMT
Ecuador
1000
-1.4569445
Estacao Biologica Pinto Mirador
21
-78.108055
Mera
22
323
14
9
5
Morona-Santiago
2847261321
CEMT
Ecuador
-2.5330555
Estacao Biologica Pinto Mirador
21
-78.01667
Mera
22
323
1
1
Morona-Santiago
2847261446
[262,993,980,1007]
MQCAZ
Ecuador
Puerto Napo
22
323
1
Napo