New Colombian Goja Navás (Psocodea: ‘ Psocoptera’: Epipsocidae) with peculiar genitalia, and the first Goja with brachypterous male, from Oaxaca, Mexico Carrejo, Nancy Obando, Ranulfo González Casasola-González, José Arturo García Aldrete, Alfonso N. Zootaxa 2021 2021-09-23 5040 4 451 481 9J49Q Carrejo & Obando & Casasola-González & García Aldrete, 2021 Carrejo & Obando & Casasola-González & García Aldrete 2021 [151,377,151,178] Insecta Epipsocidae Goja Animalia Psocodea 21 472 Arthropoda species vallecaucana sp. nov.  ( Figs 40–45)   Diagnosis.It differs from  G. andina  n. sp., and  G. risaraldensis  n. sp., in the head pattern, wing venation, in the apex of the aedeagal arch, in the endophallic sclerites, and mainly in that the external parameres are long, slender, curved inward and distally acuminate (compare Figs 1-2, 19-20and 40-41, Figs 3, 21and 42, and Figs 6, 24and  45inthis paper).  Color(in 80% ethanol). Body brown with light spots. Head ( Fig. 42): vertex dark brown, cream laterally on boundaries with the inner margin of the compound eyes; forehead brown, with dark brown transverse bands. Compound eyes dark brown to black; ocelli hyaline, with ochre centripetal crescents, postclypeus brown, with ochre striae converging mesally, as illustrated; genae cream, with dark brown spots at midline, postgenae and subgenae cream; labrum light brown, with dark brown labral sclerites, anteclypeus brown. Maxillary palps light brown to cream, Mx 1-2 cream, Mx 3-4 brown.Antennae: scape and pedicel light brown, flagellomeres brown. Thorax brown to cream, pronotum light brown, tergal lobes of meso- and metathorax brown, pleura cream. Legs brown to cream, trochanters, femora and tibiae cream, tarsi light brown. Wings hyaline, veins light brown; forewings ( Fig. 40) amber, with abundant brown setae. Abdomen brown, sternum light brown to cream. Clunium, hypandrium, epiproct and paraprocts light brown. Phallosome brown, with aedeagal arch and endophallic sclerites dark brown, external parameres brown.  Morphology.Head: H/MxW: 1.62; compound eyes small, H/d: 3.84; IO/MxW: 0.64. Outer cusp of lacinial tips broad, with eight denticles. Mx4/Mx2: 1.10. Labrum elongate, lower margin with a pair of rounded projections. Forewings ( Fig. 40): FW/W: 2.79; pterostigma elongate, wider in the middle: lp/wp: 5.53; Rs 2-3 branched; M five to six branched. Areola postica elongate: al/ah: 4.89, open in some specimens. Hindwings ( Fig. 41): HW/w: 3.21. Rs 2-branched, M simple or 2-branched. Legs: trochanters with two long setae on dorsal surface; fore- tarsi without ctenidiobothria, mid- and hind- tarsi with ctenidiobothria. Hypandrium ( Fig. 44), broad, setose, posterior border concave in the middle. Phallosome ( Fig. 45), almost straight anteriorly, side struts slender, straight; aedeagal arch robust, apex with two short mesal projections, distally acuminate, curved outward; endophallic sclerites ( Fig. 45) long, slender. Clunium projected over the area of the epiproct. Paraprocts broad ( Fig. 43), oval, with setae and macrosetae on posterior border, sensory fields large, elliptic, with 25-28 trichobothria in basal rosettes. Epiproct broadly trapeziform ( Fig. 43), with three setae mesally, other setae as illustrated.   Measurements.FW: 4150, HW: 3050, F: 990, T: 1790, t1: 860, t2: 223, ctt1: 38, f1: 940, f2: 810, f3: 670, f4: 430, f5: 235, f6: 190, f7: 195, Mx4: 325, IO: 273, D: 215, d: 265, IO/d: 1.51, PO: 0.79.  Specimens studied.  Holotypemale. COLOMBIA. Valle del Cauca. Santiago de Cali, Los Andes, Pueblo Nuevo, 3°25’57.3”N: 76°37’15.4”W,  1682 m.,  20.I.2011, R. González& N. Carrejo, on tree trunk.  Paratypes: 1 male, same locality as the holotype,  15. VI.2011, R. González.  2 males, Santiago de Cali, Los Andes, Charco Azul, 3°25’20.3”N: 76°36’50.3”W,  1658 m.,  20.I.2013, O. Sáenz& N. Calderón.  1 male, same locality,  23.I.2013, R. González& N. Calderón.  1 male, Santiago de Cali, K18, ca. Zingara, 3°30’38.3”N: 76°37’13.8”W,  1990 m.,  25.XI.2012, O. Sáenz& N. Calderón.  1 male, Santiago de Cali, Pance, Quebrada El Pato,  18. VI.2011, J. A. Mendivil.  2 males, Yumbo, Dapa, Palo Alto, 3°34’9.0”N: 76°34’13.6”W,  1866 m.,  19.III.2011, R. González.  1 male, same locality,  17. VI.2011, J. A. Mendivil. 1 male, 10.IX.2011, J. A. Mendivil.  1 male, La Cumbre, Chicoral, 3°34’30.3”N: 76°35’36.2”W,  1722 m.,  27. VI.2011, J. A. Mendivil.  1 male, Yotoco, Bosque Yotoco, 3°52’N: 76°23W,  1609 m.,  12.I.2011, R. González& A. N. García Aldrete.  1 male, Darién, Puente Tierra, 3°53’21.5”N: 76°24’W,  1600 m.,  6.XI.2010, R. González.  1 male, Roldanillo, Vía Bélgica, 4°26’57.7”N: 7°13’21.9”W,  1781 m.,  10.III.2013, O. Sáenz& N. Calderón, on tree trunk.   Etymology.The specific epithet refers to the department of Valle del Cauca, Colombia. 2011-01-20 R Gonzalez & N. Carrejo Colombia Santiago de Cali 1682 3.4325833 Pueblo Nuevo 1 -76.62095 Los Andes 21 472 1 Valle del Cauca holotype 2011-06-15 VI, R Gonzalez Colombia Santiago de Cali 1682 3.4325833 Pueblo Nuevo 1 -76.62095 Los Andes 21 472 1 1 Valle del Cauca paratype 2013-01-20 O. Saenz & N. Calderon Colombia Charco Azul 1658 3.4223056 Santiago de Cali 1 -76.61397 Los Andes 21 472 2 2 Valle del Cauca paratype 2013-01-23 R Gonzalez & Calderon Colombia Charco Azul 1658 3.4223056 Santiago de Cali 1 -76.61397 Los Andes 21 472 1 1 Valle del Cauca paratype 2012-11-25 O. Saenz & N. Calderon Colombia 1990 3.510639 Santiago de Cali 1 -76.6205 Zingara 21 472 1 1 Valle del Cauca paratype [335,1235,1442,1468] 2011-06-18 VI Quebrada El Pato & J. A. Mendivil Colombia Santiago de Cali Pance 21 472 1 1 Valle del Cauca paratype 2011-03-19 R Gonzalez Colombia Palo Alto 1866 3.5691667 Yumbo 1 -76.57044 Dapa 21 472 2 2 Valle del Cauca paratype 2011-06-17 VI Mendivil Colombia Palo Alto 1866 3.5691667 Yumbo 1 -76.57044 Dapa 21 472 1 1 Valle del Cauca paratype 2011-06-27 VI La Cumbre & J. A. Mendivil Colombia 1722 3.5750833 Chicoral 1 -76.59339 21 472 1 1 Valle del Cauca paratype 2011-01-12 R Gonzalez & A. N. Garcia Aldrete Colombia 1609 3.8666668 Yotoco 1307 -76.38333 Bosque Yotoco 21 472 1 1 Valle del Cauca paratype 2010-11-06 R Gonzalez Colombia 1600 3.8893054 Darien 923 -76.4 Puente Tierra 21 472 1 1 Valle del Cauca paratype 2013-03-10 O. Saenz & N. Calderon Colombia 1781 4.4493613 Roldanillo 1 -7.22275 Via Belgica 21 472 1 1 Valle del Cauca paratype