Helix conicoides Trochomorpha conicoides Nigritella conicoides Dendrotrochus conicoides Trochonanina conicoides Eurybasis conicoides Geotrochus conicoides Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species Vermeulen, Jaap J. Liew, Thor-Seng Schilthuizen, Menno ZooKeys 2015 531 1 139 3FTBC Metcalfe, 1851 Metcalfe 1851 Gastropoda Trochomorphidae Geotrochus CoL Animalia Geotrochus conicoides Stylommatophora 103 104 Mollusca species conicoides  Taxon classification Animalia Stylommatophora Trochomorphidae    HelixconicoidesMetcalfe, 1851: 71. Trochomorpha conicoides(Metcalfe) Wallace, 1865: 407. Nigritella conicoides(Metcalfe) Godwin Austen, 1891: 42. Dendrotrochus conicoides(Metcalfe) Kobelt, 1897: 50. Trochonanina conicoides(Metcalfe) Von Martens, 1908: 284. Eurybasis conicoides(Metcalfe) Van Benthem Jutting, 1941: 23. Geotrochus conicoides(Metcalfe) Van Benthem Jutting, 1959: 143. Type from Borneo (unspecified).  Examined material from Sabah. West Coast Province. Pulau Gaya, 8 km Northwest of Kota Kinabalu (leg. M. Schilthuizen, BOR/MOL 2937). Pulau Tiga (leg. UMS students, BOR/MOL 899). Tawau Province. Tawau Hills N.P., path up to Bukit Bombalai (leg. J.J. Vermeulen, V 13167).  Description. Shell medium-sized, rather thin, opaque, uniformly pale to dark brown (some material from Sarawak a paler brown just below the suture), (low-)conical with almost flat to distinctly concave sides, apex slightly to distinctly protruding. Surface more or less dull or with a silky luster above the periphery, shiny below. Whorls: Apical whorls moderately convex, next whorls approx. flat to slightly convex, outer whorlsoften flat; suture impressed in the inner whorls, not so in the outer, coinciding with the periphery; last whorl acutely angular, (distinctly) compressed at the periphery, slightly to rather distinctly concave just below the periphery, slightly to moderately convex towards the umbilical region. Protoconch 1 7/8-2 1/2 whorls, smooth. Teleoconch. Radial sculpture: above and below the periphery inconspicuous growth lines, somewhat raised locally. Spiral sculpture: last whorl with a sharp, distinctly pinched peripheral ridge; start of fifth whorl with 5-17 rather densely placed to somewhat spaced, low and thin spiral threads, the 2-5 threads close to the periphery usually slightly more distinct, the others very inconspicuous or (almost) absent; no spiral sculpture below the periphery. Umbilicus closed. Peristome thickened and reflexed. Dimensions: Height 7.7-10.0 mm; width 16-19.8 mm; h/w 0.42-0.53; peripheral keel of the last whorl at 0.57-0.66 of the shell height, measured from the apex; diameters of the first 4 whorls 1.3-1.5 mm, 2.0-2.5 mm, 2.5-4.0 mm, 4.2-5.6 mm respectively; number of whorls 7-8, height aperture 2.8-3.8 mm; width aperture 8-9.5 mm.  Figure 86-88. 86 Geotrochus conicoides(Metcalfe, 1851) 86A Frontal view, shell 8.5 mm high 86B Frontal view, shell 11.0 mm high 86C Apical view (86A, 86C Malaysia, Sarawak, 4th Division, Niah Caves N.P., N and NW side of limestone massif, V 1573 86B Malaysia, Sabah, Tawau Province, Tawau Hills N.P., path up to Bukit Bombalai, V 13167) 87 Geotrochus spilokeiriasp. n. 87A Frontal view, shell 8.5 mm high 87B Umbilical view (Malaysia, Sabah, West Coast Province, Crocker Range N.P., Ulu Kimanis, along Keningau-Kimanis road, BOR/MOL 2784, holotype, aperture slightly reconstructed) 88 Geotrochus scolopssp. n. 88A Frontal view, shell 10.0 mm high 88B Umbilical view (Malaysia, Sabah, Tawau Province, Tawau Hills N.P., BOR/MOL 2432, holotype).  Habitat in Sabah and distribution. Primary and old secondary forest on sandstone and volcanic bedrock, lowland conditions. Living on decaying wood. Sabah, scattered localities: Labuan, Pulau Tiga; Pulau Gaya, Tawau Hills. Also in Sarawak. Distribution: Sumatra. A record from Morotai, Moluccas ( Van Benthem Jutting 1941: 23) is probably erroneous.  Cross diagnosis. Generally characterized by the conical spire, but low-conical specimens occur. These approach the Geotrochus labuanensisgroup in general shape but have a more distinctly protruding apex and more distinctly concave sides. Also, relative to the shell height, the peripheral keel is positioned closer to the base of the shell, and the lower surface of the shell is often rather distinctly concave towards the periphery. See also the remarks under Geotrochus scolops.