Girault, 1913 : 214 Girault, 1913 : 215 Girault, 1935 : 3 Girault, 1913 : 213 Brèthes, 1914 : 8 Girault, 1916 : 41 Dozier, 1933 : 100 Thysanus flavellus : Peck (1951) Thysanus thoreauini : Peck (1951) Signiphora flavella : Rozanov (1965) Signiphora louisianae : Gordh (1979) Taxonomic revision of the flavopalliata species group of Signiphora (Hymenoptera: Signiphoridae) Woolley, J. B. Dal Molin, A. Zootaxa 2017 2017-08-31 4315 1 1 150 4XB7R Girault, 1913 F Girault 1913 [151,557,753,779] Insecta Signiphoridae Signiphora Animalia Hymenoptera 56 57 Arthropoda species flavella  http://eol.org/pages/855954     Signiphora flavella  Girault, 1913: 214. Female. urn:lsid:zoobank.org:act:EF3BFAB6-7CCE-44A6-BE3D-C7326B2B5F19    Signiphora basilica  Girault, 1913: 215. Female. urn:lsid:zoobank.org:act:2415DB5E-ABCE-4C3C-B7ED-0E294E5507CA    Signiphora euclidi  Girault, 1935: 3. Female. NEW SYNONYMYurn:lsid:zoobank.org:act:F263AE05-83ED-443C-A404-3E286A9C1841    Signiphora flava  Girault, 1913: 213. Female. NEW SYNONYMYurn:lsid:zoobank.org:act:73A50AFE-7F26-4CA8-8492-CE593189DEF5    Signiphora caridei  Brèthes, 1914: 8. Female. NEW SYNONYMYurn:lsid:zoobank.org:act:38881A2F-1168-46FD-8FA1-14A76035EB0B    Signiphora thoreauini  Girault, 1916: 41. Female. NEW SYNONYMYurn:lsid:zoobank.org:act:9D229455-B423-49EC-8275-AD3A7272A0BB    Thysanus louisianae  Dozier, 1933: 100. Male. NEW SYNONYMYurn:lsid:zoobank.org:act:D60392F5-FB61-4B3E-89EC-87289659D931   Thysanus flavellus: Peck (1951).   Thysanus thoreauini: Peck (1951).   Signiphora flavella: Rozanov (1965).   Signiphora louisianae: Gordh (1979).    Signiphora thoreauini: Gordh (1979).   Diagnosis.Fore wing without discal seta, marginal vein with seta M1 present (rarely absent); Mt 8 infemale transverse, without a medial emargination; body coloration variable but generally yellow, with or without brown markings on meso- and metasoma; antennal clava commonly dusky brown in distal 1/4–1/3 or entirely dusky brown. Many specimens have a characteristically long pedicel, although this is not always the case.   Signiphora flavellais most similar to  S. aleyrodis,  S. coquillettiand  S. xanthographa;however, these species are parasitoids of Aleyrodidae(  S. flavellais a parasitoid of Diaspididae) and always have brown markings on the mesosoma and metasoma, a uniformly tan or light brown antennal clava, and females have a rounded or v-shaped medial emargination on Mt8. In addition,  S. flavellalacks the distinct reticulate sculpture found on the vertex, frons and mesoscutum of  S. xanthographa.  Signiphora flavellais also similar to  S. perpauca, however the latter species has a discal seta in the fore wing.   Description. Female.Length, anterior margin of pronotum to epiproct apex, 0.31–0.79 mm(n=51). Vertex and frons orange–tan to light brown, face and gena yellow or pale tan, occipital margin ringed with a brown band, clypeus dark brown. Pedicel, anelli and clava dusky brown, distinctly darker than scape or body, or clava yellow or tan with distal 1/4–1/2 darker than proximal portion. Body coloration variable, most commonly body pale yellow except posterior 1/3–2/3 of mesoscutum, scutellum, metanotum and propodeum (particularly the median sclerite) distinctly paler or lighter, almost white; Mt8, epiproct and ovipositor sheaths dusky brown; often more or less extensively marked with brown or dusky as follows: pronotum and anterior 1/4–3/4 mesoscutum light brown, Mt1 yellow or light brown, Mt2 only, or Mt2 and Mt3, or Mt2–Mt4 brown, Mt6 often with brown spots laterally or sometimes entirely dusky brown. Fore wing infuscated from base to below stigmal vein or beyond with hyaline areas behind submarginal vein normal for  flavopalliatagroup.  Head. Mandibular ducts enlarged apically. Pedicel length:scape length 0.33–0.86; 3 anelli, the second from subequal to 3× length of first, the third 1.5–4.0× length of first; clava length:scape length 1.44–1.96. Vertex and frons minutely and transversely striate, frons with 4 longitudinal rows of minute punctations.  Mesosoma. Pronotum and mesoscutum transversely imbricate. Scutellum, metanotum and medial sclerite of propodeum imbricate or weakly so. Scutellum with 6 or 7 setae (rarely 4 or 5) and 2 campaniform sensilla; medial propodeal sclerite rounded, process on medial sclerite rounded or pointed apically. Fore wing 2.9–4.1× as long as wide; fore wing LMS:fore wing width 1.3–2.1; marginal vein 2.1–3.3× stigmal vein; marginal vein with 6 dorsal setae and without ventral setae; seta M1 rarely absent (see discussion); seta M3 length:marginal vein length 0.43– 0.81; apical end of costal cell most commonly at seta M2 but may be from proximal to seta M2 to seta M3. Hind wing with subparallel margins, 6.4–10.9× as long as wide, 0.28–0.50× fore wing width, fore wing LMS:fore wing width 2.62–6.14. Mesofemur with 1 long spine and 1 short spine in posteroapical margin; mesotibial spur with 5–8 teeth, mesotibial spur length:basitarsus length 0.63–1.30; basitarsus length:mesotibia length 0.47–0.77.  Metasoma. Mt1 strongly bilobed with medial portion transverse or occasionally with medial portion rounded; Mt1 length:Mt2 length 0.5–3.0 (most commonly 0.5, see discussion). Ovipositor in dorsal view with anterior-most portion under propodeum to Mt4; ovipositor length:metasoma length 0.45–0.95; ovipositor sheath length:ovipositor length 0.15–0.30; Ms3–Ms6 with anterior projections short to long; Ms 6 inposterior 1/4 metasoma and with 6–10 setae; Mt8 with transverse anterodorsal margin, without a medial emargination ( Fig. 206).   FIGURES 197–200.  Signiphora flavella: 197, head (BMNH(E) 990140); 198, female antenna (BMNH(E) 990140); 199, mandibles (BMNH(E) 990140); 200, male antenna (BMNH(E) 990134).   FIGURES 201–206.  Signiphora flavella: 201, fore wing, female (BMNH(E) 990140); 202, venation of fore wing (UCRC ENT 299381); 203, hind wing, female (UCRC ENT 299381); 204, venation of hind wing (BMNH(E) 990153); 205, middle leg, female (UCRC ENT 299346); 206, Mt8 of metasoma, female (BMNH(E) 990153).   Male.Length, anterior margin of pronotum to epiproct apex, 0.29–0.61 mm(n=8). As described for females except the following. Male coloration highly variable: as in female but without the dusky brown area at apex of metasoma. Clava length: scape length 1.40–1.79. Genitalia normal for  flavopalliatagroup, digitus with 1 apical denticle and 1 seta at its midpoint, length of digitus approximately 2× width. Ms8 a thin, transverse strip, extending to cerci laterally.   Discussion.The species we treat as synonyms of  S. flavellawere described primarily on the basis of differences in coloration. For example,  S. basilicawas distinguished primarily on the basis of lateral brown markings on Mt6 (the typeof  S. basilicais from the same rearing as the typesof  S. flavella). Peck (1951)synonymized  S. basilicawith  S. flavellaand most authors ( e.g. De Santis 1968; Gordh 1979) have followed this interpretation. This particular color pattern is most common in material from Argentinaand Brazil, but with one possible exception, discussed below, we have no other evidence to indicate that it represents another species. In fact, the range of coloration in long series of reared material from California, Argentina, Brazil, Israeland Greecemore than encompasses the patterns of coloration in the typespecimens of  S. basilica, S. flava, S. thoreauini,and  S. caridei.  Signiphora louisianaewas described from several male specimens. Although males of this species are not common in North America, the structural features and coloration of Dozier's typespecimens do not differ from that noted in the male specimens from Argentina,  Mexicoand Peru.  Signiphora euclidiwas described from a single female specimen in balsam. Although this is a new record for the species from Australia,  S. euclidiappears to fall well within the limits of  S. flavellaas here defined.   FIGURES 207–212.  Signiphora flavella: 207, female habitus (USNM type 14195, holotype of  flavaGirault); 208, mesosoma of female (BMNH(E) 990153); 209, metasoma of female (BMNH(E) 990153); 210, male habitus (BMNH(E) 990134); 211, male genitalia (BMNH(E) 990144); 212, Ms8 of metasoma, male (BMNH(E) 990144). However, two different series of specimens may represent cryptic species. First, specimens from certain collections from four localities in Argentina [Buenos Aires, Saenz-Peña, 20-IV-1976, coll. Rose, ex armored scale on ivy, one ♀ in Hoyer's (UCRC ENT 299612); La Rioja, Aimogasta (Plaza), 30-XI-1968, coll. A. Teran, on olive or ex  Chrysomphaluson olive 8 ♀ and 3 ♂ in Hoyer's (SHYM0006 (IFML); UCRC ENT 299613); La Rioja, Mazán, 17-XI-1978, coll. A. Teran, ex  Acutaspis paulistaon olive, 3 ♀ and 2 ♂ in Hoyer's (SHYM0007- SHYM0008, IFML) Tucumán, Ticucho, 31-III-1969, coll. Teran and Guyot, on leaves of  Aspidosperma quebracho-blancoSchltdl., 5 ♀ and 2 ♂ in Hoyer's (SHYM0009 (IFML); UCRC ENT 299614)] differ in minorbut consistent ways from  S. flavellaas here interpreted. Males are relatively common in these collections (7 out of 24) but are rare in  S. flavella. These specimens have short antennal clava (clava length:scape length 1.25–1.45 for females and 1.00–1.31 for males) and 4 or occasionally 5 setae on the scutellum, but agree with  S. flavellawith respect to other structural features. The coloration of the metasoma of females in these is consistent: Mt1 and Mt2 brown, Mt3–Mt5 yellow, Mt6 yellow with brown lateral spots. As noted above, metasomal coloration in  S. flavellais quite variable but includes this pattern. We suspect that this material may represent a distinct species, but additional collections from Argentina and biological or molecular information will be required to confirm this. Second, many specimens in the long series from Nova Teutonia, Brazil, may represent another cryptic species. Males are also common in these collections, and specimens in of both sexes the frons and vertex are darker than typical  S. flavellaspecimens, and the sculpture on the vertex is weakly but distinctly reticulate (transversely striate in typical  S. flavella). In addition, in these female specimens Mt8 is not transverse, but u-shaped and many specimens have a distinct medial incision on Mt8. Mt1 is bilobed with the medial portion either transverse or rounded in the material examined from California, but in other material Mt1 is bilobed with the medial portion rounded. Most specimens from California have 6 or 7 setae on the scutellum, but specimens from South America typically have 4. The ratio Mt1 length:Mt2 length is 1.0 in most specimens, rarely the ratio is 0.5 or from 2.0–3.0. Seta M1 is absent from the marginal vein of the fore wing in some paralectotypes and from one fore wing of the lectotype of  S. flavella. Otherwise, 6 setae are generally present on the marginal vein of the fore wing in this species.   Type material.  Signiphora flavellaGirault—LECTOTYPE♀ [here designated]: in balsam, Florida, Miami, coll. E.A. Bessey, bred 8-VI-1908, ex  Aspidiotus lataniae[now  H. lataniae] on Ochras sapota(sapodilla); USNM Type 14196. PARALECTOTYPES:data as lectotype, 3 ♀ in balsam on same slide, USNM Type 14196. The specimen at the lower right (red USNM type labels to right) is here designated lectotype and the slide has been labeled accordingly.   Signiphora basilicaGirault—HOLOTYPE♀ [examined]:in balsam on slide with lectotype and paralectotypes of  S. flavella. Data as types of  flavella(see above); USNM Type 14197. The specimen that matches the original description of  S. basilicais to the left and slightly above the lectotype of  S. flavella(red USNM type labels to right).   Signiphora euclidiGirault—HOLOTYPE♀ [examined]:in balsam, Indooroopilly, Feb. 3, 1935, QM Holotype T.8826. As noted in Dahms (1983); Girault provided the following information in his unpublished ms.: “The type was a single female taken from a window in forest country”.   Signiphora flavaGirault—HOLOTYPE♀ [examined]:in balsam, Peru, Lima, coll. C.H.T. [Townsend], with following data: 192o3a, Nov. Gen. 2d, sp. 1, sec 31-09, T., USNM Type 14195.   Signiphora thoreauiniGirault—HOLOTYPE♀ [examined]:in balsam, USNM Type 19209, California, Santa Barbara, coll. P.H. Timberlake, 14-XI-1911, ex  Aspidiotus hederae[now  Aspidiotus nerii(Bouché)] on ivy, 14594c.   Signiphora louisianaeDozier— HOLOTYPE♂ [examined]:in balsam, USNM Type 44819, Louisiana, New Orleans, coll. H.L. Dozier, 12-I- 1932, ex  Aspidiotus lataniae[now  H. lataniae] or C. [ Chrysomphalus]  dictyospermi(Morgan)on oleander. PARATYPES:2 ♂ in balsam, USNM Type 44819 (2 slides, one slide with holotype also); data as holotype. The slide with the holotype and paratype is not marked to indicate which specimen is the holotype. Dozier's (1933) description states only that the "type male and paratype male on single slide is deposited in the U.S. National Museum collection." Both specimens on the slide agree with Dozier's description.   Signiphora carideiBrèthes— SYNTYPES [examined]:regarding material from which this species was described, Brèthes (1914)describes female and male, and then states only: “Dr. Pedro Caride Massini, to whom I dedicate this species, has sent me in the middle of winter a few branches of a palm tree attacked by  Diaspis pentagona, from which I obtained more specimens of this interesting wasp.” Dr. Luis De Santis (personal communication) kindly informed us that the Brèthes collection was conserved in the Museo Argentino de Ciencias Naturales Bernadino Rivadavia (MACN). Following my request for the type(s) of this species, the MACN sent two slides. One bears three labels, as follows: "S17", "  S. caridei" on a red paper strip, and “  S. caridei” written in ink on the slide and covered with transparent tape. This slide has one female specimen mounted in Faure's or a similar medium, which has mostly dried out. The specimen is crushed and in poor condition, although the body and head are present and in adequate condition to allow observation of certain relevant details. This is one presumed female syntype. The other slide bears the following labels: "S18", a red paper strip with no writing, and “  S. caridei” written in ink on the slide and covered with transparent tape, and "Museo Argentino de Ciencias Naturales, Signiphora caridei Br., 187, det. In'tulo Semisman [the latter not clear]." This slide contains one female, condition as above, but more badly dried out. This specimen is a second syntype.    Othermaterial examined. ALGERIA: Wilaya d'Alger: MHNG ENTO0 0 0 0 9849 ( MHNG).   ARGENTINA: Buenos Aires: 1 ♀, UCRC ENT 299612( UCR); 1 other (host material); UCRC ENT 299611 (UCR).   ARGENTINA: La Rioja: 3 ♀, 1 ♂, SHYM0006-SHYM0008( IFML); UCRC ENT 299613 (UCR).   ARGENTINA: Salta: 1 ♀, USNM ENT 763158( USNM).   ARGENTINA: Tucumán: 2 ♀, SHYM0009( IFML); UCRC ENT 299614 (UCR).   AUSTRALIA: Queensland: 1 ♀, BMNH(E) 991087 ( BMNH).    BRAZIL: Pernambuco: 1 ♀, UCRC ENT 299351( UCR).    BRAZIL: Rio de Janeiro: 2 ♀, UCRC ENT 299083, 299087( UCR).    BRAZIL: Santa Catarina: 43 ♀, 10 sex unknown, 16 ♂, BMNH(E) 1038934–1038943, 990125–990152, 990154–990182, 991088 ( BMNH); UCRC ENT 299088 (UCR).   CHILE: 4 ♀, UCRC ENT 299077, 299079– 299081( UCR).   CHILE: Santiago: 1 ♀, UCRC ENT 299084( UCR).   CHILE: Valparaíso: 1 ♀, UCRC ENT 299078( UCR).   GREECE: 11 ♀, UCRC ENT299106–299116 (UCR).   HONDURAS: Yoro: 1 ♀, USNM ENT 763049( USNM).   INDIA: 1 ♀, UCRC ENT 299344( UCR).   ISRAEL: 19 ♀, 13 sex unknown. TAUZM 165462– 165475, 165479–165487, 165492–165500 ( TAUI).    MEXICO:1 mixed series. INHS 72494( INHS).    MEXICO: Michoacán: 1 ♀, UCRC ENT 299082( UCR).    MEXICO: Morelos:1 mixed series. INHS 72508( INHS).    MEXICO: Mexico:1 mixed series. USNM ENT 763118( USNM).    MEXICO: Veracruz: 1 ♀, USNM ENT 763044( USNM).   MOROCCO: Rabat-Sale-Zemmour-Zaer: 1 ♀, MHNG ENTO0 0 0 0 9853 ( MHNG).    NEW  ZEALAND: 1 ♀, BMNH(E) 990153 ( BMNH).    PERU: 1 ♂, USNM ENT 763068( USNM).    PERU: Callao:8 sex unknown. BMNH(E) 1038945–1038952 ( BMNH).    PERU: Lima: 1 ♀, USNM ENT 763043( USNM).    PERU: Piura: 1 ♀, 5 ♂, INHS 72509( INHS); USNM ENT 763067, 763069–763072 (USNM).   PUERTO RICO: USNM ENT 763046( USNM).   SOUTH AFRICA: 3 ♀, USNM ENT 763051( USNM); TAMU–ENTO x0616172, x0616176 (SANC).   SOUTH AFRICA: Cape Province: 12 ♀, UCRC ENT299089–299099 (UCR); TAMU– ENTO x0616168 (SANC).   SPAIN: 2 ♀, UCRC ENT299345–299346 (UCR).   TRINIDAD & TOBAGO: 3 ♀, CNCHYMEN  122361( CNC); UCRC ENT 299085–299086 (UCR).   USA: California: 58 ♀, 1 sex unknown. BMNH(E) 1038944 ( BMNH); USNM ENT 299384; USNM ENT 763047 (USNM); UCRC ENT 299061–299076, 299100–299104, 299361, 299347–299350, 299352–299383 (UCR).   USA: Florida: 1 ♀, 2 ♂, TAMU-ENTO X0852781, X0852782, x0853048 ( TAMU).   USA: Louisiana: 4 ♀, USNM ENT 763045, 763050, 763052–763053 ( USNM).   USA: Texas: 1 ♀, USNM ENT 763048( USNM).   VENEZUELA: Mérida: 3 ♀, 1 ♂, CNCHYMEN  122464–  122467( CNC).   Countrynot specified: 1 ♀, UCRC ENT 299105( UCR).   Biology. DeBach et al.(1958)mentioned that this species (cited as  Thysanus thoreauini) develops as a primary parasitoid of  Hemiberlesia rapax(Comstock)and  H. lataniae(Diaspididae). It is commonly reared from these scales in southern Californiaand elsewhere, often in sympatry with  S. merceti. The species appears to be a common and cosmopolitan parasitoid of armored scales. The host range includes many species of Diaspididaeincluding:  A. aurantii,  Aonidiella ensiferaMcKenzie,  A. nerii, A. spinosus,  Aulacaspis rosae(Bouché);  A. paulista, L. beckii,  Hemiberlesia palmae(Cockerell);  H. rapax,  Oceanaspidiotus spinosus(Comstock);  Parlatoria pergandii(Comstock);  Parlatoria pittospori(Maskell);  P. trilobitiformis, and  C. perniciosa. Three specimens from South Africafit the diagnosis of  S. flavellaperfectly, but were reared from soft scales [Durban, Natal, iii.1964, C.J. Villiers, ex: soft scale on  Grewiasp. [TAMU-ENTO x0616172 and x0616176, SANC]; Cape Province, Port Elizabeth, xii.1963, J.F. de Villiers with  Ceroplastessp. on  Dovyalis caffra[TAMU-ENTO x0616168, SANC], perhaps an indication of another cryptic species. Interestingly, 3 additional specimens with identical label data as the Durban, Natal specimens appear to be  S. perpauca, a similar species that has a discal seta in the fore wing, a feature which is not known to be polymorphic in  Signiphora, despite the long series of reared specimens of each species. This species is generally uniparental. Males are unknown in Californiaand rare in collections from the southeastern USA,  Mexicoand Argentina. 1587279799 [199,1433,223,248] MHNG, ENTO Algeria Other 61 62 1 Wilaya d'Alger 1587279787 [151,910,260,285] UCRC, UCR Argentina Buenos Aires 61 62 ENT 299612 1 1 Buenos Aires 1587279791 [246,1086,295,320] IFML Argentina La Rioja 61 62 SHYM0006-SHYM0008 4 3 1 La Rioja 1587279780 [151,816,332,357] USNM Argentina Salta 61 62 ENT 763158 1 1 Salta 1587279785 [826,1432,332,357] IFML Argentina Tucuman 61 62 SHYM0009 2 2 Tucuman 1587279782 [518,1299,367,392] BMNH Australia Queensland 61 62 1 1 Queensland 1587279798 BRAZIL, UCRC, UCR Brazil Pernambuco 61 62 ENT 299351 1 1 Pernambuco 1587279792 BRAZIL, UCRC, UCR Brazil Rio de Janeiro 61 62 ENT 299083, 299087 2 2 Rio de Janeiro 1587279803 BRAZIL, BMNH Brazil Santa Catarina 61 62 59 43 16 Santa Catarina 1587279783 UCRC, UCR Chile CHILE 61 62 ENT 299077, 299079- 299081 4 4 1587279801 [339,961,511,536] UCRC, UCR Chile Santiago 61 62 ENT 299084 1 1 Santiago 1587279794 UCRC, UCR Chile Valparaiso 61 62 ENT 299078 1 1 Valparaiso 1587279797 [336,778,548,573] UCRC, ENT Greece GREECE 61 62 11 11 1587279800 USNM Honduras Yoro 61 62 ENT 763049 1 1 Yoro 1587279786 [356,830,583,608] UCRC, UCR India INDIA 61 62 ENT 299344 1 1 1587279784 TAUI Israel ISRAEL 61 62 TAUZM 165462- 165475, 165479-165487 19 19 1587279781 [738,1296,620,645] INHS Mexico MEXICO 61 62 INHS 72494 1 MEXICO 1587279822 UCRC, UCR Mexico MEXICO 61 62 ENT 299082 1 1 MEXICO 1587279824 [726,1432,655,680] INHS Mexico MEXICO 61 62 INHS 72508 1 MEXICO 1587279812 [151,941,692,717] USNM Mexico MEXICO 61 62 ENT 763118 1 MEXICO 1587279811 USNM Mexico MEXICO 61 62 ENT 763044 1 1 MEXICO 1587279826 [362,1355,727,752] MHNG, ENTO Morocco Rabat-Sale-Zemmour-Zaer 61 62 1 1 Rabat-Sale-Zemmour-Zaer 1587279813 NEW, ZEALAND, BMNH New Zealand ZEALAND 61 62 1 1 ZEALAND 1587279817 [692,1182,764,789] PERU, USNM Peru PERU 61 62 ENT 763068 1 1 1587279835 PERU, BMNH Peru Callao 61 62 1 Callao 1587279821 [742,1337,799,824] PERU, USNM Peru Lima 61 62 ENT 763043 1 1 Lima 1587279810 PERU, INHS Peru Piura 61 62 INHS 72509 6 1 5 Piura 1587279815 USNM Puerto Rico PUERTO RICO 61 62 ENT 763046 1 1587279805 [438,1116,871,896] USNM South Africa SOUTH AFRICA 61 62 ENT 763051 3 3 1587279814 [385,1137,908,933] UCRC, ENT South Africa Cape Province 61 62 12 12 Cape Province 1587279807 [465,859,943,968] UCRC, ENT Spain SPAIN 61 62 2 2 1587279823 61-12-23 CNC Trinidad and Tobago TOBAGO 61 62 3 3 TOBAGO 1587279820 BMNH United States of America California 61 62 58 58 California 1587279832 TAMU United States of America Florida 61 62 3 1 2 Florida 1587279809 USNM United States of America Louisiana 61 62 ENT 763045, 763050 4 4 Louisiana 1587279827 [267,840,1124,1149] USNM United States of America Texas 61 62 ENT 763048 1 1 Texas 1587279816 64-12-24 67-12-24 64-12-24 CNC Venezuela Merida 61 62 4 3 1 Merida 1587279825 [430,1076,1159,1184] UCRC, UCR Venezuela Country 61 62 ENT 299105 1 1 Merida