Additions to Neotropical species of the genera Lithocharodes Sharp, 1881 and Somoleptus Sharp, 1881 (Coleoptera, Staphylinidae) Irmler, Ulrich Contributions to Entomology 2024 2024-01-31 74 1 13 34 https://zoobank.org/CFE8C1AE-3C95-4449-8C63-D6E699F84AB9 Irmler, 2024 Irmler 2024 Insecta Staphylinidae Somoleptus CoL Animalia Somoleptus paramocola Coleoptera 0 13 Arthropoda species paramocola sp. nov.   Figs 16a-d , 21D  Material examined.    Holotype: COSTA RICAmale; Cartago; Cerro Chirripo; Vallede los Conelos; 9°28'37"N, 83°29'23"W;  3600 melevation; paramo shrub litter; 26 VI 1999; R. Andersonleg.; KNHM CR1A99 104D.    Paratypes: COSTA RICA 3 females; same data as holotype; KNHM;  1 male; same data; UIC;  1 male, San Jose/Cart.; km 71 Int-Am. Hwy; nr. Tres de Junio; 9°37'44"N, 83°50'13"W;  2900 melevation; wet cloud forest litter; 23 VI 1999; R. Andersonleg.; KNHM CR1A99 101A.  Diagnosis. Together with  S. alajuelaesp. nov.,  S. paramocolaforms a new sub-group. The species resembles a group of other species with short elytra and extremely short eyes,  S. brunneusIrmler, 2022,  S. triangulusIrmler, 2022,  S. longicepsIrmler, 2022 and  S. ovatusIrmler, 2022 from the same high montane region of Central America. It can be differentiated from those species by the extremely short eyes with PS:E ratio 6.4 and the unique structure of the aedeagus.  Description. Length: 6.1 mm; Colouration: completely black; legs and antennae dark brown. Head: 1.06 mm long, 0.91 mm wide; eyes not prominent; extremely short; PS:E ratio 6.4; postocular sides slightly divergent; posterior angles sub-rectangular; posterior edge straight; inter-antennal furrows short; not reaching front edge of eyes; setiferous punctation deep and dense; on average, interstices between punctures half as wide as diameter of punctures; surface without microsculpture; polished; antennae with first antennomere elongate; half as long as head; second and third antennomeres longer than wide; conical; combined half as long as first antennomere; following antennomeres wider than long; anteriad increasing in width. Pronotum: 1.86 mm long, 0.78 mm wide; widest at anterior third; anteriad, convexly narrowed to neck; posteriad slightly convergent; posterior angles sub-rectangular; centre of posterior margin straight; setiferous punctation deep and moderately dense; on average, interstices between punctures as wide as diameter of punctures; a long wide impunctate mid-line denser; adjacent to mid-line, 17-18 punctures; surface without microsculpture; polished. Elytra: 0.78 mm long, 0.87 mm wide; humeral angles oblique; posteriad, sides divergent; posterior angles sub-rectangular; posterior margin deeply retreated to suture; hind wings absent; setiferous punctation slightly finer than on pronotum, but as dense; on average, interstices between punctures 0.8 times as wide as diameter of punctures; surface without microsculpture; polished. Abdomen with setiferous punctation still finer than on elytra, but as dense; surface with transverse reticulate microsculpture; shiny; sternite and tergite VII of male with short central emargination; meso- and meta-tibia with one ctenidium each. Aedeagus nearly circular; dorsal plate covering nearly total central lobe; apical edge slightly prominent; straight; cones at apical orifice translocated laterally; combined forming a shape like cattle horns; parameres bilobed; inner lobe wide at base; abruptly narrowed in apical half to acute top; covered by several setae; two setae in apical half thicker than at base.  Etymology. The species name is a combination of the words Paramothat names the habitat in high Neotropical mountains and the Latin word colomeaning "dwelling"and refers to the habitat where the species was collected.  Geography. Central Costa Rica.  Ecology. Floor of high cloud forests and in Paramo between approximately 2900 and 3600 m elevation in June. 4515812345 1999-06-26 R. Anderson COSTA RICA Cerro Chirripo 3600 9.476944 Conelos 21 -83.489716 Valle 1 Cartago Holotype 4515812324 2024-01-01 2024-12-31 2024-01-01 R. Anderson COSTA RICA Cerro Chirripo 3600 9.476944 COSTA RICA 21 -83.489716 Valle 3 3 Cartago Paratypes 4515812321 2024-01-01 2024-12-31 2024-01-01 R. Anderson Costa Rica Cerro Chirripo 3600 9.476944 Conelos 21 -83.489716 Valle 1 1 Cartago paratype 4515812367 1999-06-23 de Junio, R. Anderson Costa Rica 2900 9.628889 Tres de Junio 21 -83.836945 Hwy 1 1 San Jose paratype