Afrotropical Melambiina: revision of the genus Tragardhus Koch, 1956 and redescription of Pseudemmallus Koch, 1956 (Tenebrionidae: Pedinini) Kamiński, Marcin Jan Zootaxa 2017 2017-11-10 4347 1 71 93 Koch 1956 [454,596,1042,1068] Insecta Tenebrionidae Tragardhus GBIF Animalia Coleoptera 2 73 Arthropoda genus    1. Body length: 4.5–6.0 mm. Center of pronotal disc with two pairs of rounded carinae ( Fig. 2A). Epipleuron dilated apically, with roundish alveola at the level of fourth ventrite ( Fig. 2C). Abdomen with fine punctures. Fifth abdominal ventrite simple ( Fig. 2E). Second lobe of coxites trapezoidal, much smaller in size than paraproct ( Fig. 5A)......... subgenus  Tragardhus  -. Body length: 3.0– 4.5 mm. Pronotal disc simple, without rounded carinae ( Fig. 2B). Epipleuron evenly narrowed towards apex, without any additional structures ( Fig. 1N). Abdomen with coarse punctures. Fifth abdominal ventrite raised in apical third ( Fig. 2D). Second lobe of coxites rounded laterally, nearly as large as paraproct ( Fig. 5B)......... subgenus Mitragardhus