Chthonius (C.) agazzii Beier, 1966: 175 Chthonius (s. str.) ischnocheles ischnocheles : Lazzeroni 1969b: 390 C. densedentatus C. ischnocheles Chthonius (s. str.) malatestai Callaini, 1980a: 220 Chthonius (s. str.) agazzii : Callaini 1980a: 226 Gardini 1991: 172 Chthonius (C.) jugorum : Minelli & Negrisolo 1993: 65 The Italian species of the Chthonius ischnocheles group (Arachnida, Pseudoscorpiones, Chthoniidae), with reference to neighbouring countries Gardini, Giulio Zootaxa 2021 2021-06-17 4987 1 1 131 V39C Beier, 1966 Beier 1966 [151,514,367,394] Arachnida Chthoniidae Chthonius Animalia Pseudoscorpiones 5 6 Arthropoda species agazzii  ( Figs 1–25, 402)      Chthonius (C.) agazzii Beier, 1966: 175, fig. 1 (in part: ♀only).    Chthonius(s. str.) ischnocheles   ischnocheles: Lazzeroni 1969b: 390(in part: see  C. densedentatusand  C. ischnocheles).   Chthonius(s. str.) malatestai Callaini, 1980a: 220, figs 1a–c, 2a–d (n. syn.).     Chthonius(s. str.) agazzii: Callaini 1980a: 226;  Gardini 1991: 172.     Chthonius (C.) jugorum: Minelli & Negrisolo 1993: 65.     Typelocality: Italy, Veneto, Treviso Province, Pederobba, La Bislonga 1001 V/ TV( 45°52’48”N, 11°57’41”E).   Distribution. Italy( Friuli, Venetoand Trentino).   Diagnosis ( ♂ ♀).A microphthalmic epigean or hypogean  Chthoniusthat differs from other species of the  ischnochelesgroup in the following combination of characters: anterior margin of carapace with 1 (rarely 2) preocular microseta; posterior margin of carapace with 2 macrosetae, without lateral microsetae; chelicerae with 1 (rarely 2) lateral microseta; chela length 0.56–1.08 mm; movable chelal finger length 0.39–0.74 mm; chelal fingers with reclined, pointed, widely spaced teeth; movable chelal finger with coupled sensilla pclodged on a noticeable tubercle.   Type material of  Chthonius malatestaiexamined.  ITALY—  Veneto: Treviso Prov.— 1 ♂( holotype), “Cansiglio [Fregona],  12.XI.1976, MG, [Mount] Pizzoc, faggeta” ( MSNV).  Other material examined.  ITALY—  Friuli: Pordenone Prov.— 1 ♂, Aviano, Cansiglio, Col Piova,  21. VI.1973, R. Poggileg., beech wood; 4 ♂ 1 ♀, Aviano, near Madonna del Monte,  340 ma.s.l.,  17.IV.1995, G. Governatorileg.; 3 ♂, Aviano, Piancavallo, 1980–1990, M. Seriani leg., beech wood; 2 ♂, id., 15.XI.1981, M. Seriani leg.; 5 ♂ 4 ♀, id., 1250 ma.s.l., 3.VII.1983, M. Seriani leg.; 1 ♂, id., 1500 ma.s.l., 15. V.1994, G. Governatori leg., beech wood; 2 ♂, Barcis, Lago di Barcis,  409 ma.s.l.,  7.X.1979, M.G. Paolettileg.;  1 ♀, Barcis, Nslope of Montelonga,  800 ma.s.l.,  11.IX.2006, G. Governatorileg., beech wood; 1 ♀, Budoia, San Tomè di Dardago,  450 ma.s.l.,  19.X.1993, F. Gasparoleg., mixed wood; 1 ♂, Claut, Alta Val Cellina, Pian de Cea,  930 ma.s.l.,  30. V.2002, A. Dall’Astaleg.,  pitfall trapin beech wood; 2 ♂, Claut, near Stalle Parentonia,  750 ma.s.l.,  22. VI.1996, G. Governatorileg.;  1 ♂ 1 ♀, Claut, Val Settimana, between Stalle Pariginaand Stalla Pustrin,  650 ma.s.l.,  3. V.2002, A. Dall’Astaleg., sieved in beech wood; 4 ♂, Clauzetto, Gerchia,  540 ma.s.l.,  29.IX.1990, F. Gasparoleg.;  3 ♂, Tramonti di Sotto, BoscoSelvaz,  21. VI.1987, L. Dreonleg.;  4 ♂, Tramonti di Sotto, near Campone,  2.X.1985, S. Zoialeg.   Friuli: Udine Prov.— 1 ♂ 2 ♀, Amaro,  350 ma.s.l.,  8. V.1991, F. Gasparoleg., wet meadows; 4 ♂ 5 ♀, Bordano, 260 ma.s.l., 12. VI.1991, F. Gasparo leg., pitfall traps; 1 ♂ 2 ♀, Forgaria nel  Friuli, Monte Prat,  750 ma.s.l.,  22.VII.1991, F. Gasparoleg.;  1 ♀, Raveo,  430 ma.s.l.,  21. VIII.1991, F. Gasparoleg.;  1 ♂, near Tolmezzo,  23.VII.1978, S. Vitleg.;  2 ♀, Venzone, Val Verzonassa, Rio Bruschienear Casera Frassin,  630 ma.s.l.,  29. V.2008, L. Dorigoleg., beech wood ( MFNB); 2 ♂ 1 ♀, Villa Santina,  400 ma.s.l.,  29.IV.1994, F. Gasparoleg.,  Carpinuswood; 1 ♂, id., 5.VII.1994, F. Gasparo leg.,   Carpinuswood.  Trentino: Trento Prov.— 5 ♀, Vallarsa, Passo di Pian delle Fugazze,  1140 ma.s.l.,  2.VII.2013, D. Biancoleg., digging.   Veneto: Belluno Prov.— 1 ♂, Belluno, N slope of Monte Faverghera,  1520 ma.s.l.,  5.XI.2000, G. Peretto& E. Pivaleg.; 1 ♀, id., 20. V.2001, E. Piva leg.;  1 ♂ 1 ♀, Mel, Praderadego,  750 ma.s.l.,  2.XI.1990, L. Latella& S. Zoialeg.; 1 ♂, Pedavena, Le Vette Feltrine, 870 ma.s.l., 18. VI.1994, E. Piva leg., digging in canyon; 1 ♀, Schievenin,  350 ma.s.l.,  29.III.1978, I. Löblleg.;  1 ♂, Sovramonte, Grotta Faller3189 V/BL,  25.IV.2010, D. Bianco& G. Martinileg.   Veneto: Treviso Prov.— 5 ♂(  C. jugorum, det. M. Beier), Vittorio  Veneto, Fadalto,  X.1992, A. Minellileg.;  1 ♂ 1 ♀, id., Sella di Fadalto,  500 ma.s.l.,  3. VIII.1991, F. Gasparoleg.;  3 ♂ 2 ♀, Fregona, near Pian del Cansiglio,  1000 ma.s.l.,  4.X.1985, S. Zoialeg., beech wood; 1 ♂ 2 ♀, Mansuè, Basalghelle,  10.VII.1980, M.G. Paolettileg.;  3 ♂ 6 ♀, Milies,  30.III.1978, I. Löblleg.;  1 ♂, San Pietro di Feletto, Busdei Tedeschi  1818 V/ TV,  100 ma.s.l.,  11.XII.1993, E. Pivaleg.;  1 ♂, id., La Bora 1252 V/ TV,  110 ma.s.l.,  18.XI.2013, D. Bianco& E. Pivaleg.;  1 ♂ 3 ♀, Segusino, Buco delle Salamandre 1387 V/ TV,  400 ma.s.l.,  8.XII.1996, E. Pivaleg.;  1 ♀, Segusino, digging near Buco delle Salamandre 1387 V/ TV,  400 ma.s.l.,  25. VI.2008, D. Bianco& E. Pivaleg.; 11 ♂ 5 ♀, Susegana, Castello San Salvatore, 100 ma.s.l., 23.IX.2013. G. Gardini & C. Giusto leg., mixed wood.   Veneto: Vicenza Prov.— 1 ♀, Arcugnano, near Trattoria della Moreieta,  350 ma.s.l.,  11.IV.1992, E. Pivaleg.;  1 ♀, Arcugnano, grove near Trattoria della Moreieta,  335 ma.s.l.,  31.III.1996, G. Peretto& E. Pivaleg.;  2 ♀, Arcugnano, Val Scaranti, digging near Grotta del Lupo5524 V/ VI,  230 ma.s.l.,  24. VI.2016, E. Pivaleg.;  1 ♂ 1 ♀, Arcugnano, Eslope of Monte Zattolo,  120 ma.s.l.,  18.III.2012, E. Pivaleg.;  1 ♀, Arcugnano, Villabalzana, small valley near Casa Rappo,  230 ma.s.l.,  30.X.2011, E. Pivaleg.;  1 ♂, Montecchio Maggiore, Covolo Basso, digging near the cave L’Infangatoion.c. V/ VI,  19. V.2015, D. Biancoleg.;  1 ♂ 2 ♀, Schio, Colletto Grande, military gallery at  880 ma.s.l.,  15.IX.2005, D. Biancoleg.;  1 ♀, Schio, Munari,  200 ma.s.l.,  22.VII.2010, D. Biancoleg.;  1 ♀, Valdagno, San Quirico, Pellichero,  475 ma.s.l.,  15. VI.2006, D. Biancoleg.;  1 ♂, Valdagno, Grendene,  610 ma.s.l.,  11.IV.2015, D. Biancoleg., soil washing; 1 ♀, id., 620 ma.s.l., 14.IV.2017, E. Piva leg.;  4 ♀, Valdagno, Valle del Boia,  440 ma.s.l.,  11. V.2009, D. Bianco& G. Martinileg., soil washing; 1 ♀, id., 28. V.2009, D. Bianco leg., digging; 1 ♀, Valdagno, SEslope of Monte Scandolara,  745 ma.s.l.,  14.IV.2017, D. Biancoleg., soil washing; 1 ♀, Valdagno, id.,  18.IV.2017, D. Bianco& E. Pivaleg., soil washing; 1 ♀, id., 22.IV.2017, D. Bianco leg., soil washing; 1 ♀, Velo d’Astico, Pozzo Cima Altan.c. V/ VI,  1629 ma.s.l.,  27.IX.2015, D. Bianco, P. Falsirollo& E. Filippileg.   Veneto: Verona Prov.— 2 ♂ 2 ♀(  C. ischnocheles, det. G. Lazzeroni; together with 1 ♂ 1 ♀of  C. densedentatus), Grezzana, slope N of Vajo del Paradiso,  27.III.1968, G. Osellaleg. ( MSNV);  1 ♀, Marano di Valpolicella, near Marezzane,  550 ma.s.l.,  3. VI.2007, A., M. Zanettileg., mixed wood; 1 ♀, Selva di Progno, Giazza, Vallo di Rivolto,  1100 ma.s.l.,  13.VII.2007, D. Biancoleg.;  1 ♂(  C. ischnocheles,det. G. Lazzeroni), Tregnago, Giazza,  8. V.1968, G. Lazzeronileg.( MSNV).   Description of adults ( ♂ ♀).Integument slightly pigmented, carapace, tergites, chelicerae and pedipalps pale brown; weak hispid granulation on lateral surface of carapace, cheliceral palm, base of fixed chelal finger and on dorso-distal surface of chelal hand. Carapace ( Fig. 4) almost square, moderately constricted posteriorly, 0.8–0.9 times longer than broad; anterior margin between median macrosetae strongly denticulate, rarely with prominent epistome in both sexes ( Figs 1–3); ocular area as in fig. 4, anterior eyes reduced, with flat lens and tapetum, posterior ones absent or as vestigial remains; standard chaetotaxy m 4m:6:4:2:2(20), anterior row rarely with mm 4mor mm 4mm; length of anteromedian macrosetae  0.095 –0.145mm. Chaetotaxy of tergites 4:4:4:4:6:6:6:6:1T2T1:4:1 T2T1:0. Standard chaetotaxy of sternites II–X: 10:(3)8–10(3):(2)7–9(2):8:6–8:6:6:6:2T1T2, sternite II rarely with 9 or 11 setae; genital opening of males flanked by 6–7 (rarely 5) setae on each side. Chelicerae ( Fig. 5) 2.1–2.3 ( ♂ ♀) times as long as broad, palm with 6 setae and 1 (rarely 2) lateral microseta; fixed finger with 2 distal large teeth followed by 7–10 small teeth proximally reduced in size and 3–5 proximal microtubercles; movable finger with an isolated subapical tooth ( di) at level of the spinneret, followed by a large tooth and 7–12 teeth proximally reduced in size; gsratio 0.52–0.59; spinneret weakly raised and rarely rounded in males ( Figs 5, 7), more prominent and rounded in females ( Fig. 6); rallum with 11 blades ( 12 in ♀from Piancavallo, 3.VII.1983); serrulae interior and exterior with 14 and 15 blades respectively. Coxal setae: pedipalp 5 (including 2 on manducatory process), I 3 + 3 marginal microsetae, II 4, III 5, IV 6 (coxa I with 2 + 4 marginal microsetae in ♂from Arcugnano, Mt. Zattolo); coxa II with 5–11 (mainly 6–8) coxal spines, coxa III with 3–6 (mainly 4–5) coxal spines; intercoxal tubercle bisetose. Pedipalp: femur 4.5–5.6 ( ♂ ♀) times as long as broad, femoral chaetotaxy 3:5:2:5(6):1; chela ( Figs 10, 14) 5.0–6.1 ( ♂) or 4.55–5.6 ( ♀) times as long as deep; hand of chela 1.55–1.9 ( ♂) or 1.5–1.75 ( ♀) times as long as deep, with elongated oval outline in dorsal view ( Figs 8–9); fixed chelal finger with 34–55 teeth inclined backwards: 7–12 small, contiguous and pointed distal teeth, proximally increasing in size, followed by large, sharp, reclined and widely spaced teeth ( Figs 18–25), decreasing in size from trichobothrium esttowards finger base; 6–20 proximal teeth low, more contiguous, rounded, gradually reduced in size ( Figs 11–13, 15–17); all teeth with dental canal; fixed finger at level of est-itwith 5–9 ( ♂) or 4–7 ( ♀) teeth occupying 0.1 mm, distance between successive apices 0.012 –0.027( ♂) or 0.011 –0.032( ♀) mm; tip of fixed chelal finger with apical sensilla af  1-2, distal paraxial seta gradually curved and thin; movable chelal finger with 30–47 teeth, slightly scalene triangle-shaped, reclined and apically pointed proximally up to trichobothrium st,rounded and increasingly reduced from sttowards finger base, reaching back near b(between sband bthey are mainly recognizable only by the presence of dental canal) ( Figs 11–13, 15–17); movable finger at level of st-twith 6–9 ( ♂) or 5–8 ( ♀) teeth occupying 0.1 mm, distance between successive apices 0.011 –0.019( ♂) or 0.011 –0.023( ♀) mm; coupled sensilla pcjust distad of sband on a noticeable tubercle; tip of movable chelal finger with apical sensilla am  1-2; trichobothria as in figs 10, 14; ratio sb -st/sb -b= 1.5–1.9; ratio of movable finger/hand of chela 2.0–2.3 ( ♂ ♀); ratio of pedipalpal femur/movable finger 0.9–1.0 ( ♂ ♀); ratio of pedipalpal femur/carapace 1.2–1.5 ( ♂ ♀). Measurements (in mm). Body length 1.0–1.5 ( ♂) or 1.1–1.7 ( ♀). Carapace 0.30–0.43 × 0.35–0.47 (0.32–0.45 anteriorly) ( ♂) or 0.32–0.50 × 0.40–0.53 (0.37–0.51 anteriorly) ( ♀). Chelicerae 0.31–0.43 × 0.14–0.20 ( ♂) or 0.36–0.50 × 0.17–0.23 ( ♀); movable finger length 0.16–0.23 ( ♂) or 0.18–0.26 ( ♀). Pedipalp: femur 0.36–0.60 × 0.08–0.11 ( ♂) or 0.40–0.65 × 0.09–0.12 ( ♀); chela 0.56–0.93 × 0.105–0.16 ( ♂) or 0.605–1.08 × 0.125 –0.215( ♀); hand length 0.175 –0.295( ♂) or 0.195–0.35 ( ♀); movable finger length 0.39–0.62 ( ♂) or 0.42–0.74 ( ♀).   Remarks.  Chthonius agazziiwas described by Beier (1966)from four femalesand one malefrom Veneto(Cave La Bislonga 1001 V/TV, Pederobba, and Bus de le Fave 1272 V/TV, Refrontolo, respectively). The types of  C. agazziiare still not traceable (see also Gardini 1998). Beier (1966)highlighted the presence of one microseta on each side of carapacal basal series in male, that are absent in females. The examination of five femalesfrom the cave Bus de le Fave ( 4.X.1985, S. Zoia, R. Sciaky & E. Piva leg.; 5.I.1991, E. Piva leg.) allows me to attribute the same to  C. ischnocheloides Beier, 1973, a hypogean species that is widespread in the same geographical area as  C. agazziiand belongs to the  C. orthodactylus-tenuisgroup according to the arrowhead-shaped distoparaxial seta on the fixed chelal finger. It is therefore almost certain that the male paratypeof  C. agazziibelongs to  C. ischnocheloides.   The cave Buco delle Salamandre 1387 V/ TV, Segusino, from which the illustrated male and female were collected, is only 5.5 kmaway from the cave La Bislonga, the typelocality of  C. agazzii .   FIGURES 18–25.  Chthonius agazzii Beier, 1966, detail of dentition at the chelal fingers tip, antiaxial view (setae and sensilla af 1-2, am 1-2omitted). 18, male holotypeof  C. malatestai, Mt. Pizzoc (Fregona, Treviso); 19, male, Milies (Treviso); 20, female, Pozzo Cima Alta (Velo d’Astico, Treviso); 21, female, near Trattoria della Moreieta (Arcugnano, Vicenza); 22, female, near Marezzane (Marano di Valpolicella, Verona); 23, male, Bus dei Tedeschi (San Pietro di Feletto, Treviso); 24, female, slope of Mt. Faverghera (Belluno); 25, female, near Buco delle Salamandre (Segusino, Treviso).   Chthonius malatestaiwas described by Callaini (1980a)from a male and two femalesfrom Veneto[ ♂ ♀, Monte Pizzoc, Fregona (loc. typ.); ♀, Passo di San Boldo, Cison di Valmarino] and compared with  Chthonius ressli Beier, 1956and  C. agazzii. Following Callaini (1980a),  Chthonius malatestaishould differ from  C. agazziiin having smaller size, less slender pedipalps and in the shape of chelal teeth. Fig. 18reproduces dentition of the distal half of chelal fingers in male holotypeof  C. malatestai. The specimens examined share, with rare exceptions (see the above Description of adults), the same chaetotaxy of carapace (m 4m:6:4:2:2), the presence of one lateral microseta on the cheliceral palm, the chelal fingers with the same tooth pattern (fixed finger distally with small, reclined and contiguous teeth, becoming more widely spaced proximally), coupled sensilla pcalways distad of sb(between the sixth and tenth teeth) and placed on a noticeable tubercle; but at the same time they have wide inter-populational variability in size, which does not seem to follow any geographical or adaptive gradient. Large specimens usually have more widely spaced chelal teeth while smaller specimens have closer up teeth, as evident in specimens from Buco delle Salamandre ( Figs 10–17) and Mt. Pizzoc ( Fig. 18) respectively. The continuum of variability in the dentition of the chelal fingers is represented in figs 18–25. In view of the above,  Chthonius malatestaiCallaini, 1980is proposed here as a junior subjective synonym of  Chthonius agazzii Beier, 1966 (n. syn.). Pedipalpal measurements of specimens of  Chthonius agazziifrom Cansiglio, Veneto( Callaini 1980a) fall within the range of variability reported in the above description of the species. The doubtful quotation of  C. agazziifrom the cave Cevola dei Battei 245 Fr/PN, Friuli( Gardini 1991) must be referred to  C. ischnocheloides. Among the species of the  Chthonius ischnochelesgroup,  C. agazziiis morphologically very close to  C. paolettii Beier, 1973, a hypogean species from Veneto.  Chthonius agazziidiffers from  C. paolettiiin the following characters (comparison among females): chela length 0.605–1.08 mm( 1.10–1.15 mmin  C. paolettii); movable chelal finger length 0.42–0.74 mm( 0.74–0.77 mmin  C. paolettii); fixed and movable chelal fingers at level of est–istand st–t,respectively, with 4–7 and 5–8 teeth occupying 0.1 mm, distance between successive apices 0.011 –0.032and  0.011 –0.023mmrespectively (with 3 and 5 teeth occupying 0.1 mm, distance between successive apices 0.035 and 0.025 mm, respectively, in  C. paolettii); movable chelal finger with coupled sensilla pcon a noticeable tubercle (coupled sensilla pclodged not on a tubercle in  C. paolettii).   Chthonius agazziiis also morphologically similar to  C. comottii Inzaghi, 1987, an epigean or locally hypogean species from the western Alps.  C. agazziidiffers from  C. comottiiin the following characters: anterior margin of carapace rarely with a prominent epistome (mostly with prominent epistome in  C. comottii), posterior margin of carapace with 2 macrosetae, without lateral microsetae (with one microseta on each side in  C. comottii); chelicerae with 1 (rarely 2) lateral microseta (with 2 lateral microsetae in  C. comottii).   Chthonius agazziiand  C. comottiiare vicariant in the Alps: the first occurs from Friulito Trentino and the latter is found from Lombardyto western Liguria. 3323445933 [151,1394,664,691] V, TV La Bislonga Italy 45.879997 Treviso Province 18 11.961389 5 6 1 Veneto holotype 3323445628 1976-11-12 MG, MSNV Italy Treviso Prov. 5 6 1 1 Veneto holotype 3323445669 1973-06-21 VI, R Poggi Italy Pordenone Prov. Cansiglio Aviano 5 6 1 1 Friuli-Venezia Giulia 3323445600 [308,1432,1060,1087] 1995-04-17 Aviano & G. Governatori Italy 340 Madonna del Monte 5 6 5 1 4 3323445386 1979-10-07 Barcis & M. G. Paoletti Italy 409 Lago di Barcis 5 6 2 2 3323445719 2006-09-11 Barcis, N & G. Governatori Italy 800 Montelonga 5 6 1 1 3323445505 1993-10-19 Budoia & F. Gasparo Italy 450 San Tome di Dardago 5 6 1 1 3323445415 [335,1432,1240,1267] 2002-05-30 V Claut & A. Dall'Asta Italy 930 Pian de Cea Val Cellina 5 6 1 1 3323445598 [151,1331,1276,1303] 1996-06-22 VI Claut & G. Governatori Italy 750 Stalle Parentonia 5 6 2 2 3323445513 2002-05-03 V Claut & A. Dall'Asta Italy Val Settimana 650 Stalla Pustrin Stalle Parigina 5 6 2 1 1 3323445478 1990-09-29 Clauzetto & F. Gasparo Italy 540 Gerchia 5 6 4 4 3323445397 1987-06-21 VI L. Dreon Italy Bosco Tramonti di Sotto 5 6 3 3 3323445674 [505,1189,1384,1411] 1985-10-02 S. Zoia Italy Campone Tramonti di Sotto 5 6 4 4 3323445470 1991-05-08 V Amaro & F. Gasparo Italy 350 Udine Prov. 5 6 3 2 1 Friuli-Venezia Giulia 3323445567 [727,1410,1456,1483] 1991-07-22 F. Gasparo Italy 750 Monte Prat 5 6 1 Friuli-Venezia Giulia 3323445972 1991-08-21 VIII F. Gasparo Italy 430 Raveo 5 6 1 1 3323445708 [762,1265,1492,1519] 1978-07-23 S. Vit Italy Tolmezzo 5 6 1 1 3323445427 2008-05-29 V Venzone & L. Dorigo Italy Val Verzonassa 630 Casera Frassin Rio Bruschie 5 6 2 2 3323445704 1994-04-29 MFNB Villa Santina & F. Gasparo Italy 400 5 6 3 1 2 3323445308 2013-07-02 Vallarsa & D. Bianco Italy Trentino 1140 Passo di Pian delle Fugazze Trento Prov. 5 6 5 5 3323445687 2000-11-05 G. Peretto & E. Piva Italy 1520 Monte Faverghera Belluno Prov. 5 6 1 1 Veneto 3323445655 1990-11-02 L. Latella & S. Zoia Italy 750 Praderadego 5 6 2 1 1 3323445315 1978-03-29 I. Lobl Italy 350 Schievenin 5 6 1 1 3323445523 2010-04-25 V D. Bianco & G. Martini Italy Grotta Faller Sovramonte 5 6 1 1 3323445327 [436,1152,1780,1807] Vittorio Italy M. Beier Treviso Prov. 5 6 5 5 Veneto 3323445625 1992-10 A. Minelli Italy Fadalto 5 6 1 Veneto 3323445518 [328,1115,1816,1843] 1991-08-03 VIII F. Gasparo Italy 500 Sella di Fadalto 5 6 2 1 1 3323445378 1985-10-04 Fregona & S. Zoia Italy 1000 Pian del Cansiglio 5 6 5 2 3 3323445734 1980-07-10 Mansue & M. G. Paoletti Italy Basalghelle 5 6 3 2 1 3323445938 [303,776,1888,1915] 1978-03-30 I. Lobl Italy Milies 5 6 9 6 3 3323445741 1818-05 1993-12-11 1818-05 V, TV E. Piva Italy San Pietro di Feletto 100 Tedeschi Bus 5 6 1 1 3323445918 2013-11-18 V, TV D. Bianco & E. Piva Italy 110 La Bora 5 6 1 1 3323445814 [208,1257,1960,1987] 1996-12-08 V, TV Segusino & E. Piva Italy 400 Buco delle Salamandre 5 6 4 3 1 3323445313 2008-06-25 V, TV, VI D. Bianco & E. Piva Italy 400 Buco delle Salamandre Segusino 5 6 1 1 3323445454 1992-04-11 Arcugnano & E. Piva Italy 350 Trattoria della Moreieta Vicenza Prov. 6 7 1 1 Veneto 3323445392 1996-03-31 G. Peretto & E. Piva Italy 335 Trattoria della Moreieta 6 7 1 1 3323445934 2016-06-24 V, VI Arcugnano & E. Piva Italy 230 Grotta del Lupo Val Scaranti 6 7 2 2 3323445484 2012-03-18 Arcugnano, E & E. Piva Italy 120 Monte Zattolo 6 7 2 1 1 3323445673 2011-10-30 Arcugnano & E. Piva Italy 230 Casa Rappo Villabalzana 6 7 1 1 3323445873 2015-05-19 V, VI Montecchio Maggiore & D. Bianco Italy cave L'Infangatoio Covolo Basso 6 7 1 1 3323445693 2005-09-15 Schio & D. Bianco Italy 880 Colletto Grande 6 7 3 2 1 3323445301 [293,981,403,429] 2010-07-22 Schio & D. Bianco Italy 200 Munari 6 7 1 1 3323445945 2006-06-15 VI Valdagno & D. Bianco Italy 475 Pellichero San Quirico 6 7 1 1 3323445738 [624,1383,438,465] 2015-04-11 Valdagno & D. Bianco Italy 610 Grendene 6 7 1 1 3323445605 2009-05-11 V D. Bianco & G. Martini Italy 440 Valle del Boia 6 7 4 4 3323445545 2017-04-14 Valdagno, SE & D. Bianco Italy 745 Monte Scandolara 6 7 1 1 3323445803 2017-04-18 D. Bianco & E. Piva Italy Valdagno 6 7 1 1 3323445884 2015-09-27 V, VI D. Bianco, P. Falsirollo & E. Filippi Italy 1629 Pozzo Cima Alta Velo d'Astico 6 7 1 1 3323445764 1968-03-27 MSNV Grezzana & G. Osella Italy G. Lazzeroni Verona Prov. 6 7 6 3 3 Veneto 3323445566 2007-06-03 VI A., M. Zanetti Italy 550 Marezzane Marano di Valpolicella 6 7 1 1 3323445460 [291,1431,727,753] 2007-07-13 D. Bianco Italy Selva di Progno 1100 Vallo di Rivolto Giazza 6 7 1 1 3323445944 [151,1138,763,790] 1968-05-08 V G. Lazzeroni Italy G. Lazzeroni Giazza Tregnago 6 7 1 1 3323445463 V, TV cave La Bislonga The cave Segusino 8 9 1