Additions to the knowledge of the land snails of Sabah (Malaysia, Borneo), including 48 new species Vermeulen, Jaap J. Liew, Thor-Seng Schilthuizen, Menno ZooKeys 2015 531 1 139 3FTBP http://zoobank.org/A3969A3E-5C3D-4A25-BAE8-13E53AAD5FD8 Vermeulen, Liew & Schilthuizen Gastropoda Trochomorphidae Geotrochus CoL Animalia Geotrochus meristotrochus Stylommatophora 110 111 Mollusca species meristotrochus  Taxon classification Animalia Stylommatophora Trochomorphidae     Holotype. Malaysia, Sabah, Interior Province, Sepulut valley, Batu Punggul(leg. J.J. Vermeulen, RMNH.5003932).  Examined material from Sabah. Interior Province. Upper Padas valley, Long Pa Sia (leg. T.S. Liew & Meckson, BOR/MOL 4343). Crocker Range N.P., near the km 59 marker on the road Kota Kinabalu-Tambunan (leg. J.J. Vermeulen, V 1186); West of the km 10 marker on the road Tambunan-Ranau, Mahua Waterfall (leg. J.J. Vermeulen & M. Schilthuizen, V 9743; leg. J. Schilthuizen, BOR/MOL 896; leg. M. Schilthuizen, BOR/MOL 2420, BOR/MOL 2423, BOR/MOL 2424, BOR/MOL 3143, BOR/MOL 3345). Gunung Trusmadi slopes, Gua Loloposon (leg. J.J. Vermeulen, V 13236). Pinangah valley, Batu Urun (= Bukit Sinobang) (leg. J.J. Vermeulen, V 8008, V 1163, BOR/MOL 891); Pun Batu c. 30 km West of Sepulut (leg. J.J. Vermeulen, V 1302); Batu Punggul (leg. J.J. Vermeulen, V 1979; leg. T.S. Liew, M. Schilthuizen & S. Chiba, BOR/MOL 4317); Batu Pungiton (leg. M. Schilthuizen, BOR/MOL 890); Batu Temurung (leg. J.J. Vermeulen, V 8054, BOR/MOL 892; leg. M. Schilthuizen, BOR/MOL 893); Gua Sanaron (leg. J.J. Vermeulen & M. Schilthuizen, V 8073; leg. T.S. Liew, M. Schilthuizen & S. Chiba, BOR/MOL 4280; leg. M. Schilthuizen, BOR/MOL 887); Kampung Labang (leg. J.J. Vermeulen & M. Schilthuizen, V 7727, BOR/MOL 895). Sandakan Province. Beluran, Meliau Range (leg. T.S. Liew, M. Schilthuizen & A. van Til, BOR/MOL 3207, BOR/MOL 3208, BOR/MOL 3209, BOR/MOL 3210). Tawau Province. Batu Baturong, North slope (leg. J.J. Vermeulen, V 7603). Segama valley, limestone hill on North bank Segama River, near bridge of road Sandakan to Lahad Datu (leg. J.J. Vermeulen, V 7517). Tawau Hills Park (leg. J.P. King, BOR/MOL 2425; leg. T.S. Liew, J.J. Vermeulen & M. Schilthuizen, BOR/MOL 4272). West Coast Province. Kinabalu N.P., Serinsim-Numbuyukon trail between 548 and 1204 m (leg. T.S. Liew, BOR/MOL 4474, BOR/MOL 4475, BOR/MOL 4477). Kota Kinabalu, Kiansom (leg. M. Schilthuizen, BOR/MOL 889, BOR/MOL 2935; leg. UMS Tropical Malacology Course participants, BOR/MOL 3481).  Description. Shell rather small, rather thin, about opaque, yellowish to pale brown, moderately low-conical with flat to slightly convex sides; apex slightly protruding or not. Surface with a silky luster. Whorls: Apical whorls moderately convex, outer slightly convex; suture impressed, between the outer 4-5 whorls slightly below the periphery; last whorl acutely angular, slightly compressed at the periphery, rounded below the periphery. Protoconch 2 1/8-2 1/2 whorls, smooth. Teleoconch. Radial sculpture: above the periphery some growth lines, locally grading into inconspicuous riblets; below the periphery indistinct growth lines only. Spiral sculpture: last whorl with a sharp, slightly pinched peripheral keel; start of fifth whorl with 6-9 thin, widely spaced spiral threads: 1-2 distinct close to the periphery, as well as 4-8 inconspicuous ones above these; no spiral sculpture below the periphery. Umbilicus closed. Peristome thickened and reflexed. Dimensions: Height 6.0-7.0 mm; width 9.3-10.8 mm; h/w 0.59-0.67; diameters of the first 4 whorls 1.0-1.4 mm, 1.8-2.2 mm, 2.7-3.4 mm, 4.0-5.0 mm respectively; number of whorls 6 3/4-7 3/8, height aperture 2.8-3.5 mm; width aperture 4.8-5.5 mm.  Habitat in Sabah and distribution. Primary, secondary forest and abandoned agricultural land on limestone and sandstone soil, up to 1300 m alt. Sabah: West part, East coast (lower Segama River valley and further South). Endemic to Sabah.  Cross diagnosis. Shell less wide than Geotrochus subscalaris, compare shell width and aperture width of both species. Usually, the h/w-ratio is also higher than in Geotrochus subscalaris, but here a slight overlap exists.  Etymology. The name refers to the slightly protruding peripheral keel, creating a small notch in between the whorls in lateral view [meristos (Gr.) = divided; trokhos (Gr.) = wheel, often used for the gastropod spire]. 2015-01-01 2015-12-31 2015-01-01 RMNH J. J. Vermeulen Malaysia http://data.biodiversitydata.nl/naturalis/specimen/RMNH.5003932 Interior Province Sabah RMNH.5003932 1 holotype