A Revision of the New World Genera Brachylinga Irwin and Lyneborg and Lysilinga Irwin and Lyneborg (Diptera: Therevidae: Therevinae) with the Description of a New Genus, Elcaribe Webb
Webb, Donald W.
Metz, Mark A.
Zootaxa
2006
2006-08-14
1288
1288
1
241
396LV
Webb
Webb
2006
[264,591,1004,1030]
Insecta
Therevidae
Elcaribe
Animalia
Diptera
174
175
Arthropoda
species
guanaensis
sp. nov.
Etymology Ensis(Latin)= a suffix denoting place, locality, country. This species is named for Guana Island from where the majority of specimens of this species were collected and is used as a noun in apposition. Diagnosis Elcaribe guanaensisis taxonomically associated with E. bahamaensis, E. elongatus, E. glabrus, E. laticornis, E. obscurus,and E. scarbroughiin having an anterodorsal patch of dark, short setae on the gena. Elcaribe guanaensisdiffers from E. bahamaensis, E. elongatus, E. glabrus, E. laticornis, E. obscurus,and E. scarbroughiin having anteroventral and posteroventral macrosetae on the midfemur. Description of holotypemale (MEI 048133) Body length 4.7 mm. Head.Length 0.68 mm. Ommatidia smaller ventrally and laterally. Frons pruinescence gray; setae dark reddish brown, elongate. Antenna/head length 0.69; black, pruinescence on scape gray; scape length 0.16 mm, width 0.12 mm, length/width 1.3, scape/pedicel length 2.3, scape/flagellum width 1.0, setae reddish brown dorsally, dark yellow ventrally; pedicel length 0.07 mm, length/width 0.7; first flagellomere oval, attenuate apically, flagellum length 0.24 mm, width 0.12 mm, length/width 2.0, flagellum/ scape length 1.5. Parafacial pruinescence blackish gray. Maxillary palpus dark yellow; length 0.32 mm, length/width 8.0; setae white. Genal setae white. Occipital setae white becoming pale whitish yellow, appressed lanceolate dorsally.
Thorax.Macrosetae 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Mesonotum pruinescence dull gray; setae black, elongate intermixed with white, lanceolate, subappressed setae. Pleuron pruinescence gray; setae white on anepisternum, katepisternum, laterotergite, metanepisternum, and scutellum. Wing.Membrane pale brownish gray; veins pale brown; length 3.5 mm, length/width 2.9; pterostigma indistinct; cell m 3closed, petiolate. Halter knob pale yellow, stalk dark brown. Legs.Femora dark brown, pruinescence white; ventral setae white on forefemur; av 3:1:5, pv 2:2:2. Tibiae dark yellow, apex dark reddish brown. Tarsomeres 1–2 dark yellow, apex dark reddish brown; 3–5 dark reddish brown. Abdomen.Dark reddish brown, pruinescence white; dorsal setae white, lanceolate, appressed; lateral setae white elongate. Terminalia(MEI 037178) pale dark reddish brown. Tergite 8 and epandrium (Fig. 225), setae pale yellow. Sternite 8 ( Fig. 239), sides converging posteriorly, posterior margin emarginate. Hypoproct (Fig. 225) quadrate, posterior margin emarginate; apicoventral surface flat. Gonocoxite ventral view ( Fig. 239), basomedial lobe rounded, posterior twothirds tapered to narrow apex, setae pale yellow (occasionally with several dark reddish brown macrosetae medially and apically) with anterior patch of short, dark yellow setae; dorsal view ( Fig. 253) gonocoxal bridge moderately large; lateral view ( Fig. 267) oblong, tapered evenly toward apex; ventral lobe ( Fig. 239) lacking spines. Gonostylus (Fig. 281), basodorsal lobe absent; ventral margin curved, expanded apically with dark yellow apical macrosetae, not extended apically in distinct tuft; fine, short spines over apical area. Aedeagus with dorsal apodeme ( Fig. 295) quadrate, sides sinuate; ventral apodeme (Fig. 309) large, broadened anteriorly; distiphallus dorsal view ( Fig. 295) broad basally, tapered to rounded point; lateral view ( Fig. 323) narrow basally, curving ventrally at 90° angle; ejaculatory apodeme ( Fig. 295) slightly expanded anteriorly, not extending beyond anterior margin of dorsal apodeme, anterior margin truncate, in crosssection flattened dorsoventrally. Variation (n=10). Body length 4.3–5.2, 4.7 mm. Thorax. Macrosetae 2–3 np, 2 sa, 1 pa, 2 dc, 2 sc. Legs.Anteroventral macrosetae 3–6:1–3:4–7, pv 1–3:1–2:2–3. Female Similar to male except for following. Variation (n=8). Body length 4.2–5.0, 4.5 mm. Head.Frons pruinescence pale brownish gray; setae dark reddish brown, elongate. Antenna dark brown, base of first flagellomere yellow, pruinescence on scape gray; setae on scape dark reddish brown, grading into macrosetae. Thorax.Macrosetae 3 np, 2 sa, 1 pa, 2 dc, 2 sc. Legs.Anteroventral macrosetae 1–5:1–5:4–8, pv 1–2:1–2:1–2. Abdomen.Black, glossy, pruinescence gray on posterior margin of tergites 1–7, becoming wider on posterior tergites; setae black, appressed with white, elongate setae laterally on tergite 1 and across posterior margin of tergites 1–3. Terminalia(MEI 037198). Anterior margin of sternite 8 ( Fig. 337) with large medial projection. Furca (Fig. 349) triangular, posterior margin pointed; length 0.36mm, width 0.26 mm; posteromedial projection absent, anterolateral and anteromedial projections short. Common duct (Fig. 349) shorter than furca. Spermathecal duct (Fig. 349) saclike posteriorly then becoming threadlike. Distribution Elcaribe guanaensishas been collected in the British Virgin Islands, Puerto Ricoand the U. S. Virgin Islands( Fig. 358). Habitats and phenology Adults of Elcaribe guanaensishave been handnetted and collected at blacklights in a Coccolobaforest and on a beach. Specimens have been collected from 24 March to 23 July. Specimens examined Type specimen. The holotypemale of Elcaribe guanaensisWebb(MEI 048109) is labeled "BRIT. VIRGIN ISL. Guana Island, 0–80m, 6–23 July 1985, S. E. & P. M. Miller" and is deposited in the USNM. PARATYPES. BRITISH VIRGIN ISLANDS. ANEGADAISLAND. Setting Point[ 18.717, 64.383], 0 m, 17–19.VII.1985, S. Miller, P. Miller, 1♂ME I 037183 ( USNM), blacklight. GUANA ISLAND. North Bay[ 18.483, 64.567], 0–80 m, 5–23.VII.1985, S. E. Miller, P. M. Miller, 14♂MEI 037166 ( BMNH), 037171 (CNC), 037182 (FSCA), 037185 (INHS), 037188 (MEIC), 037173–9, 037190, 109133 (USNM), 5♀037191 ( BMNH), 037194 (CNC), 037195 (FSCA), 037196 (INHS), 037197 (USNM), blacklight; 15–23.VII.1985, 14♂MEI 037167–70, 048109–12, 048114, 048116, 048119–22 ( USNM), 1♀037198 ( MEIC), blacklight 1–14.VII.1984, 3♂MEI 048188–9, 048201, 2♀048187, 048190 ( USNM); 18–19.VII.1988, S. E. Miller, C. O’Connell, 1♂MEI 106344 ( BPBM). PUERTO RICO. Guánica Insular Forest, 6.VII.1953, J. R. Ramos, J. Maldonado, 1♂MEI 040245 ( USNM); 6.VI.1953, J. A. Ramos, J. Maldonado, 1♂MEI 040259 ( USNM); Route334, 26–29.VII.1969, H. & A. Howden, 2♀MEI 039717–8 ( CNC); Lajas[ 18,052, 67.060], 6.VII.1952, J. A. Ramos, J. Maldonado, 1♀MEI 040246 ( USNM); Salinas[ 17.9833, 66.3000], 16l. VII.1945, 1♀, MEI 039714 ( CNC); Guanica Bosque, Estatalde Guanica, 3.6 kmE Guanica[ 17.8669, 66.8744], 1000m, 12.VI1996, J. Rawlins, R. Davidson, C. Young, S. Thompson, 3♂CMNH 65204–6, 3♀65203–3, 65207 ( CMNH). UNITED STATES VIRGIN ISLANDS. SABAISLAND. St. Thomas County, 3.2 kmS Saint Thomas[18.331, 64.93], 24.III.1979, M. A. Ivie, L. L. Ivie, 1♂MEI 040863 ( OSU). St. Croix. Sandy Point[17.68933, 64.90], 6.II.1969, H. E. Evans, 5♂MEI 077648–52 ( MCZ); 14.II.1996, W. E. Steiner, M. Evans, M., J. Swearingen, 2♀MEI 115592–3 ( USNM), blacklight on beach; East Point, 10.XI.1967, W. H. Pierce, 5♂MEI 040208–10, 040212–3 ( FSCA).
1985-07-06
1985-07-23
1985-07-06
USNM
S. E. & P. M. Miller & S. Miller & P. Miller
British Virgin Islands
40
18.717
Setting Point
76
-64.383
The
176
177
1
1
holotype
1985-07-05
1985-07-23
1985-07-05
BMNH
S. E. Miller & P. M. Miller
British Virgin Islands
40
18.483
North Bay
76
-64.567
176
177
14
14
holotype
[264,511,1323,1350]
BMNH
British Virgin Islands
176
177
5
5
holotype
1985-07-15
1985-07-23
1985-07-15
USNM
British Virgin Islands
176
177
14
14
holotype
[525,762,1403,1430]
MEIC
British Virgin Islands
176
177
1
1
holotype
1984-07-01
1984-07-14
1984-07-01
USNM
British Virgin Islands
176
177
5
2
3
holotype
1988-07-18
1988-07-19
1988-07-18
BPBM
Miller & O'Connell
British Virgin Islands
176
177
1
1
holotype
1953-07-06
USNM
J. R. Ramos & J. Maldonado
Puerto Rico
Insular Forest
176
177
1
1
Guanica
holotype
1953-06-06
USNM
Ramos & Maldonado
Puerto Rico
176
177
1
1
Guanica
holotype
1969-07-26
1969-07-29
1969-07-26
CNC
H. & A. Howden
Puerto Rico
Route
176
177
2
2
Guanica
holotype
1952-07-06
USNM
J. A. Ramos & J. Maldonado
Puerto Rico
18.052
Lajas
76
-67.06
176
177
1
1
Lajas
holotype
[387,1209,1643,1670]
1945-07
CNC
Puerto Rico
17.9833
Salinas
7
-66.3
176
177
17
1
Salinas
holotype
1996-06-12
CMNH
J. Rawlins & R. Davidson & C. Young & S. Thompson
Puerto Rico
1000
17.8669
Estatal
7
-66.8744
Bosque
176
177
6
3
3
Guanica
holotype
1979-03-24
OSU
M. A. Ivie & L. L. Ivie
United States Virgin Islands
St. Thomas County
3.2 km S Saint Thomas
176
177
1
1
holotype
1969-02-06
MCZ
H. E. Evans
United States Virgin Islands
177
178
St. Croix. Sandy Point
176
177
5
5
holotype
1996-02-14
USNM
Steiner & Evans & M., J. Swearingen
United States Virgin Islands
177
178
2
2
holotype
1967-11-10
FSCA
W. H. Pierce
United States Virgin Islands
East Point
177
178
5
5
holotype