Chrysotus excavatus Van Duzee, 1924: 48 Taxonomic revision of Achradocera Becker (Diptera: Dolichopodidae), with description of two new species Quevedo, Lucas Capellari, Renato S. Lamas, Carlos José E. Zootaxa 2024 2024-10-09 5519 3 301 344 C2NJT (Van Duzee, 1924) Van Duzee 1924 [151,654,870,897] Insecta Dolichopodidae Achradocera Animalia Diptera 19 320 Arthropoda species excavata  ( Figs 11, 12, 26)      Chrysotus excavatus Van Duzee, 1924: 48, figs 45–46.  Type-locality: St. Thomas, Virgin Island. Holotype male, USNM.   Diagnosis (male).Postpedicel longer than arista-like stylus; lower postoccipital setae dense. Legs mostly yellow, except for FIII, dorsal surface of FI and FII, TI on apical 3/4, TII on apical half, TIII on basal 1/4 and apical 1/5, and distal tarsi, which are all brown infuscated; TI slightly bent, bearing ventral row of conspicuous setae; TII bulged ventrally in middle, with two ventral rows of setae along swelling, then constricted at apical 1/3 to apex, forming an excavation; IIt 1arched, with ring of setae at basal 1/3 and anteroventral tubercule within ring; FIII bulged on basal half and concave below, bearing clutch of strong, long setae near middle of anterior surface; IIIt 3–4concave ventrally. Hypopygium small; surstylus subtriangular, tapered and bearing 1 apical and 1 subapical strong seta; postgonite enlarged and covered by small setae.   Redescription.Similar to  A. angustifacies  stat. rev., except as noted. Male.Body length: 2.6–2.9 mm. Wing: 2.1–2.3 mmlong, 0.7–0.8 mmwide. Antenna.Postpedicel longer than arista-like stylus; FAr, 1.2. Wing.CuAx: 0.5; RMx: 1.36. Leg I.35, 32, 12, 7, 4, 3, 4. T brownish infuscated pv on apical 2/3, slightly bent beyond basal half, bearing ventral row of setae, ending with distinct apical seta; t 1entirely yellow. Leg II.32, 40, 18, 9, 7, 4, 4. T rusty yellow on apical half of dorsal side, bearing anterior seta at basal 1/4, and 4 setae apically (1 av, 1 pv, 1 ad and 1 pd); T ( Fig. 11C) bulged ventrally in middle, with two ventral rows of short setae along swelling, then constricted at apical 1/3 to apex, forming excavation; t 1arched with small av tubercule on basal 1/5 with some spines apically; bearing ventral and posterior row of strong decreasing setae, other row ad making circlet of conspicuous setae. Leg III.42, 46, 15, 9, 5, 4, 3. F bulged on basal half and concave posteriorly, bearing cluster of strong, long setae near middle of anterior surface; T yellow; t 3–4ventrally concave. Hypopygium.As in Fig. 12A–D, similar to that of  A. angustifacies  stat. rev.  Female.Unknown.   Distribution. Bahamas, British Virgin Islands, Cuba, Dominican Republic, Jamaica, and Puerto Rico( Fig. 26).   Type material examined.   HOLOTYPE ♂, labelled: “ St. Thomas, Vir. Ids. |  June 5, 1917. (129) | Harold Morrison”, “ Type”, “  Chrysotus excavatus| V.D.” [ USNM].   FIGURE 11.  Achradocera excavata. A.Habitus, left lateral. B.Legs, left lateral. C.Legs, focused on TII excavation. D.Head, frontal. E.Thorax, dorsal. Scale bar: A, 1 mm; B–E, 0.2 mm.  Additional material examined.   Bahamas. 12♂, “ GRAND BAHAMAISLAND. | Freeport|  20–27 June 1987| W. E. Steiner, | M. J. & R. Molineaux”, “ Yellow pan trapin | Caribbean pine | and palmetto scrub” [ 4♂, MZUSP].   British Virgin Islands. 1♂, “BVI: Eustatia Island| N18º30.59′ W64º21.58′| pan traps/ garden area |  28– 30OCT2016, J Runyon”;  1♂, “BVI: Virgin Gorda| Biras Hill,  90 m| N18º29.74′ W64º21.30′| Guy’s Trail, pan traps|  11NOV2016, JB Runyon”;  3♂, “BVI: Virgin Gorda| Bitter End Yacht Club| small ghut,  30melev | N18º29.74’ W64º21.38’|  10NOV2016, JB Runyon”.  Cuba .  1♂, “ Columbia| Isle of Pines| CubaVI-27-50 | C.S. 464 | 50- 9283”, “  Chrysotus excavatusVD| Wirth”;  1♂, “ Aguada Pas.| Cuba Feb.’15”, “ A. H. Sturtevant| Collection, 1970”.  Dominican Republic .  1♂, “DOMINICAN REPUBLIC: | San Cristobal, Centro| Sur DesarrolloAgro-”, “pecuaria ( Sanidad| Vegetal station) 24. |  vii.1987 A. L. Norrbom”;  3♂, “DOM. REP.: Prov. Santo| Domingo, Centrode | Tecnologias Agricolas| N18.5400 ºW69.9934º|  22OCT2021, J.B. Runyon”.  Jamaica .  1♂, “ Battersea| Jamaica|  Feb. 1910| F. Thaxter”, “ALMelander | Collection | 1961”;  1♂, “JAMAICA | 5 mi.E. Negril|  13 March 1970| W. W. Wirth| fresh march”;  3♂, “JAMAICA | AnnottoBay|  25 Feb. 1969| W. W. Wirth| marsh”;  2♂, “JAMAICA | WagWaterR. |  25 Feb. 1969| W. W. Wirth | stream”;  1♂, “JAMAICA | Pt. Henderson|  24 Feb. 1969| W. W. Wirth| Bay shore”;  1♂, “JAMAICA | Kingston| Fresh River|  24 Feb. 1969| W. W. Wirth”;  1♂, “JAMAICA | Runaway Bay|  16-28 Feb 1969| W. W. Wirth”;  2♂, “JAMAICA | Savanna La Mar|  13 March 1970| W. W. Wirth”.  Puerto Rico .  1♂, “Grape Fruit | III-1-32” [handwritten], “ AreciboP. R. | Anderson| T Faxon”, “ San Juan| No- 1967”, “1967” [all labels handwritten];  4♂, “ PUERTO RICO| Rio Piedras| Agr. Exp. Sta.|  15–18 Oct. 1968”, “S. Medina | Gaud.coll.”;  2♂, “ PUERTO RICO| Arecibo| Cambalache For.|  15–18 Feb. 1969”, “S. Medina | Gaud.coll.”;  1♂, “ PUERTO RICO| Cayey|  15–24 Nov. 1968| S. Medina Gaud”. [ Mostspecimens housed in USNM, except those collected by J. Runyon, housed in MTEC].   Remarks.It is interesting to note that  A. excavataand  A. apicalisare the only species of  Achradocerathat can be found in West Indies, and the distribution of these two species are very well delimitated:  A. excavatais restricted to the Greater Antilles while  A. apicalisto the Lesser Antilles ( Fig. 26). 4953412372 1917-06-05 V, USNM Type St. Thomas Harold Morrison 19 320 1 1 holotype 4953412340 1987-06-20 1987-06-27 1987-06-20 R, MZUSP W. E. Steiner Bahamas GRAND BAHAMA 20 321 16 16 West Grand Bahama 4953412439 2016-10-28 2016-10-30 2016-10-28 British Virgin Islands 18.509834 Eustatia Island 13 -64.359665 20 321 1 1 4953412463 2016-11-11 British Virgin Islands Virgin Gorda 90 18.495667 Guy's Trail 13 -64.355 Biras Hill 20 321 1 1 4953412347 2016-11-10 Cuba 30 18.495667 Bitter End Yacht Club 13 -64.35633 Virgin Gorda 20 321 3 3 4953412445 Cuba Columbia Cuba Isle of Pines 20 321 1 1 4953412448 A. H. Sturtevant Dominican Republic Cuba Feb. Aguada Pas. 20 321 1 1 4953412361 1987-07 A. L. Norrbom Dominican Republic Centro 21 322 Sanidad Sur Desarrollo 20 321 1 1 San Cristobal 4953412368 2021-10-22 J. B. Runyon Jamaica Prov. Santo 18.54 Centro 7 -69.9934 Domingo 21 322 3 3 4953412444 1910-02 F. Thaxter Jamaica Jamaica Battersea 21 322 1 1 4953412406 1970-03-13 W. W. Wirth Jamaica 22 323 Negril 21 322 1 1 4953412418 [439,1224,150,177] 1969-02-25 W. W. Wirth Jamaica AnnottoBay 22 323 3 3 4953412334 1969-02-25 Jamaica 22 323 2 2 4953412436 1969-02-24 Pt. Henderson & W. W. Wirth Jamaica 22 323 1 1 4953412304 [306,1113,223,249] 1969-02-24 W. W. Wirth Jamaica Fresh River 22 323 1 1 Kingston 4953412381 1969-02-16 1969-02-28 1969-02-16 W. W. Wirth Jamaica Runaway Bay 22 323 1 1 4953412353 1970-03-13 W. W. Wirth Puerto Rico Savanna La Mar 22 323 2 2 4953412320 R, T Puerto Rico Anderson San Juan Faxon 22 323 III-1-32 1 1 Arecibo 4953412431 1968-10-15 1968-10-18 1968-10-15 PUERTO, RICO Gaud. Puerto Rico Agr. Exp. Sta. Rio Piedras 22 323 4 4 4953412455 [313,1384,366,393] 1969-02-15 1969-02-18 1969-02-15 PUERTO, RICO Gaud. Puerto Rico Cambalache For. 22 323 2 2 Arecibo 4953412338 1968-11-15 1968-11-24 1968-11-15 PUERTO, RICO, USNM, MTEC Most & J. Runyon Puerto Rico Cayey 22 323 1 1 Cayey