Edessa offuscata Breddin, 1907: 332 Kirkaldy, 1909: 368 Edessa rimata Breddin, 1907: 338 Kirkaldy, 1909: 368 Edessa offuscata Revision of Hypoxys Amyot & Serville, 1843 stat. rest. (Heteroptera Pentatomidae) Nunes, Benedito Mendes Campos, Lourival Dias Mendonca, Maria Thayane Da Silva Cunha, Eduardo Victor De Paiva Fernandes, Jose Antonio Marin Zootaxa 2020 2020-02-21 4742 3 401 441 (Breddin, 1907) Nunes & Campos & Mendonca & Cunha & Fernandes 2020 Breddin 1907 [151,590,1196,1223] Insecta Pentatomidae Hypoxys GBIF Animalia Hemiptera 15 416 Arthropoda species offuscatus comb. nov.  ( Figs. 7A–E; 19 A–B; 21)      Edessa offuscata Breddin, 1907: 332;  Kirkaldy, 1909: 368(catalog)     Edessa rimata Breddin, 1907: 338;  Kirkaldy, 1909: 368(catalog) syn. nov.   Lectotypemale.  Edessa offuscata. Brasil, Pebas( SDEI). Designated by Gaedike (1971:94). Examined.    Lectotypefemale.  Edessa rimata. Peru, Iguapo( SDEI). Designated by Gaedike (1971:97, by use of “ Holotypus”). Examined.   Material examined. PERU. Loreto: ♀Iquitos, Amazon River, II–21–1927, H. Bassler ( USNM); ♂ ♀Confl. De los Rios Zamur e Yahuasyacu, II–80, J. Becker leg.  Edessa rimata Breddin, 1907Comp.w. TYPEFernandes, JAM 1999,  Edessa offuscata Breddin, 1907Comp.w. TYPEFernandes, JAM 1999. ( RMNH).   Measurements.total length: 17.2–19.0; head length: 2.3; head width: 3.4–3.6; pronotal width: 11.0–11.3; abdominal width: 9.6–10.3; length of antennal segments. I: 0.9–1.0; II: 2.4; III: 1.3–1.4; IV: 5.7; V: 6.3; interocular distance: 1.6–1.7; scutellum length: 8.4–9.0; scutellum width: 6.2–6.6; pronotal length: 3.6–3.9.   Diagnosis.Anterolateral margin of pronotum with solid black stripe reaching all its extension. Cicatrices with dark brown punctures delimiting anterior sulci ( Fig. 19A). Anterolateral margin with few dark punctures. Male ( Fig. 7A–C). Pygophore with dorsal rim widely excavated ( Fig. 7A). Superior process of genital cup small, arrow-headshaped, adjacent to dorsal rim ( Fig. 7B). Paramere elongated, shank triangular (cross section) due to a conspicuous carina on inner side ending before enlarged part; head of the paramere spatulate ( Fig. 7B). Proctiger with laterodorsal tufts of setae widely separated by medial ridge; posterior face pentagonal, flat ( Fig. 7B). Ventral rim with expansions rounded, wide, tumid, well-projected, clearly surpassing level of posterolateral angle in ventral view ( Fig. 7C); base of each expansion with a conspicuous transverse carina. Female ( Fig. 7D). Gonocoxites 8 with posterior margin sinuous, inner angle rounded projected posteriorly reaching gonocoxites 9; sutural border with elliptic, short excavation on posterior half. Gonapophyses 8 exposed.   Comments.  Edessa offuscataand  E. rimatawere described by Breddin (1907). The author considered  E. rimatarelated to  E. trabeculaand  E. offuscataclose to  E. quadridensand  E. boerneri. However, these similarities are based only on the external morphology of the species. Females of  H. offuscatushave inner angles of gonocoxites 8 more protruding and sutural border less excavated ( Fig. 7D) than  H. boerneri( Fig. 2D); both species have posterior margin of gonocoxites 8 sinuous while   H. quadridenshas posterior margin arched ( Fig. 1D). Males of  H. offuscatusand   H. quadridensare very similar and can be separated by subtle characteristics like paramere more robust and dorsal rim more excavated in  H. offuscatus( Fig. 7Band 1B); more obvious distinctive characteristic is the expansion of ventral rim more developed and rounded in  H. offuscatus( Fig. 7C) than in   H. quadridens( Fig. 1C).   Distribution ( Fig. 21). PERU: Loreto. 2570056042 [151,1194,1361,1387] SDEI Brazil Pebas 15 416 1 lectotype 2570056012 [151,1426,1392,1417] SDEI Peru Iguapo 15 416 2 lectotype