Edessa tragelaphus Breddin, 1903: 139 Kirkaldy, 1909: 165 Edessa pachyacantha Breddin, 1904a: 136 Kirkaldy, 1909: 161 Edessa tragelaphus Revision of Hypoxys Amyot & Serville, 1843 stat. rest. (Heteroptera Pentatomidae) Nunes, Benedito Mendes Campos, Lourival Dias Mendonca, Maria Thayane Da Silva Cunha, Eduardo Victor De Paiva Fernandes, Jose Antonio Marin Zootaxa 2020 2020-02-21 4742 3 401 441 (Breddin, 1903) Nunes & Campos & Mendonca & Cunha & Fernandes 2020 Breddin 1903 [151,608,837,864] Insecta Pentatomidae Hypoxys GBIF Animalia Hemiptera 31 432 Arthropoda species tragelaphus comb. nov.  ( Figs. 16A–E; 20 G–H; 23)      Edessa tragelaphus Breddin, 1903: 139; Bergroth, 1908: 179;  Kirkaldy, 1909: 165(catalog); Froeschner, 1981: 67.     Edessa pachyacantha Breddin, 1904a: 136; Bergroth, 1908: 179;  Kirkaldy, 1909: 161(catalog) syn. nov.   Lectotypefemale.  Edessa tragelaphus. Ecuador, Coca, leg. Haensch( SDEI). Designated by Gaedike (1971: 101, by use of “ Holotypus”). Examined.    Lectotypemale.  Edessa pachyacantha. Peru, Marcapata( SDEI). Designated by Gaedike (1971: 94). Examined. Gaedike also indicated one paralectotypefemale from the same locality.   Material examined.   PERU. Loreto: 2♀ ♂ Confl. de Los Rios Zumune Yahuasyaw, II–80, J. Beckerleg.,  Edessa tragelaphus Breddin, 1903Comp.w. TYPE Fernandes, JAM 1999.  Edessa pachyacanthaBreddin, 1904 Comp.w. TYPE Fernandes, JAM 1999 ( RMNH);  ♂ E. Le Moult. IquitosApril. S. Roque.  Edessa pachyacanthaBreddin, 1904 Comp.w. TYPE. Fernandes, JAM 1999 ( RMNH);  ♀  80kmNE Iquitos, 1kmfrom Amazon River on RioYana- mono, Explorama Lodge,  1–5–IX–1992, Castner, Skelley, et al. ( JEE)   Measurements.total length: 17.5–18.3; head length: 2.2–2.3; head width: 3.3–3.4; pronotal width: 12.1–13.6; abdominal width: 9.0–9.7; length of antennal segments. I: 0.5–0.6; II: 1.1–2.3; III: 1.4–1.7; IV: 3.7–5.0; V: 3.6–5.0; interocular distance: 1.4–1.7; scutellum length: 7.6–8.8; scutellum width: 5.6–6.1; pronotal length: 3.6–4.2.   Diagnosis.Anterolateral margin of pronotum without solid black stripe, more concave than the other species, impunctate. Humeral angle with apex globose and solid black ( Fig. 20G). Connexivum with small green spots. Anterolateral margin more concave than the other species. Anterolateral margin not puncture. Scutellum with anterior margin concolorous ( Fig. 20G). Ventrally apex of pseudosutures without rounded dark spots ( Fig. 20H). Male ( Fig. 16A–C). Pygophore with dorsal rim shallowly concave; lateral strongly projected posteriorly, surpassing posterolateral angles and expansions of ventral rim ( Fig. 16A,B). Paramere with anterior lobe shorter than posterior lobe; posterior lobe tongue-like. Proctiger with tufts of setae narrowly separated dorsally; posterior face subrectangular ( Fig. 16B). Ventral rim with expansion little developed, barely tumid, and acuminate ( Fig. 16C). Female ( Fig. 16D). Gonocoxites 8 inner angle reaching middle of gonocoxites 9. Laterotergites 9 narrow, clearly surpassing sclerite uniting laterotergites 8.   Comments. Breddin (1903)related  E. tragelaphusto  E. nigrispina. When analyzing the general external morphology,  H. tragelaphushas humeral angles of pronotum well developed, globose and slightly posteriorly directed ( Fig. 20G), while  E. nigrispinahas apex acute and strongly curved posteriorly. The shape of the humeral angles also distinguishes this species from the others of the group  H. oxyacanthus.   Distribution ( Fig. 23). PERU: Loreto. 2570056114 SDEI Haensch Ecuador Ecuador 31 432 2 lectotype 2570056091 SDEI Peru Marcapata 31 432 2 lectotype 2570056034 RMNH Yahuasyaw & J. Becker & Fernandes Peru Confl. de Los Rios Zumun 31 432 3 2 1 Loreto 2570056095 RMNH E. Le Moult. Iquitos April. S. Roque. & Fernandes Peru Iquitos 31 432 1 1 Loreto 2570056122 1992-09-01 1992-09-05 1992-09-01 JEE Explorama Lodge & Castner & Skelley Peru Amazon River on Rio 31 432 1 1 Loreto