Taxonomic revision of the Canthidium Erichson, 1847 species of the gigas group (Coleoptera, Scarabaeidae, Scarabaeinae) Carvalho De Santana, Edrielly C. Pacheco, Thaynara L. Vaz-De, Fernando Z. European Journal of Taxonomy 2019 2019-06-06 530 1 24 5WXGC Carvalho De Santana & Pacheco & Vaz-De, 2019 Carvalho De Santana & Pacheco & Vaz-De 2019 [525,965,535,562] Insecta Scarabaeidae Canthidium Animalia Coleoptera 12 13 Arthropoda species stofeli sp. nov. Neocanthidium   urn:lsid:zoobank.org:act: 1A3F5B17-8091-4ACC-B7DE-4311ADA3400D  Figs 7, 9    Diagnosis   Canthidium stofeli sp. nov.differs from the other species in the group by having two anterior medial projections, one on each side of the pronotum.    Etymology Named after Roberto Stofel, for the field support during collections in Cotriguaçu.    Type material    Holotype BRAZIL•  ♂; “BRASIL: Rondônia: Guajará/ Mirim: Proprietário: Fandinho/ 10°44 ′57.62 ″S, 65°19 ′0.32 ″W/  138m.flor. Amaz. Isca: baço bov/  24.II.2010 F.Coletti”; “22366”; CEMT.    Paratypes BRAZIL• 1 ♂; Acre, Rio Branco, Fazenda [“Faz”] Catuaba; 9°57 ′19.77 ″S, 65°51 ′55.72 ″W;  Feb. 1997; F. Vaz-de-Melloleg.; CEMT•  1 ♀; Rondônia, Guajará-Mirim, Sítio Coletti; flight interception trap[“FIT”]; 10°19 ′47.11 ″S, 4°45 ′12.5 ″W;  170 ma.s.l.;  28 Jan. 2010; F. Colettileg.; CEMT•  1 ♀; Pará, Tailândia, Empresa Agropalma;  9–16 Jul. 2016; P. Silvaleg.; CEMT•  1 ♀; Belterra, forest; hum/ pig trap; 3°17 ′47 ″S, 54°57 ′48 ″W;  132 ma.s.l.;  24 Mar. 2017; F. Françaleg.; T2P3; CEMT•  1 ♀; same locality as for preceding; T15P20; CEMT•  1 ♀; Mato Grosso, Querência, Fazenda São Luiz, Amazon Forest; flight interception trap[“FIT”]; 12°39 ′3.21 ″S, 52°21 ′9.37 ″W;  Feb. 2009; R. Andradeleg.; CEMT•  2 ♂♂; Cláudia, Fazenda [“Faz”] Continental; fungus; 11°34 ′54 ″S, 55°17 ′15 ″W;  20 Feb. 2010; M.F. Souzaleg.; CEMT•  1 ♂; Cotriguaçu, Matinha; pitfall baited with fungus; 9°50 ′19 ″S, 58°15 ′3 ″W;  11–13 Jul. 2009; F.Z. Vaz-de-Melloleg.; BMNH•  1 ♂; same data as for preceding; CEMT•  2 ♀♀; Cotriguaçu, Fazenda [“Faz”] São Nicolau; flight interception trap[“FIT”]; 9°49 ′9 ″S, 58°15 ′47 ″W;  12–14 Jul. 2009; F.Z. Vaz-de-Melloleg.; CEMT•  1 ♀; same data as for preceding; BMNH•  1 ♂; Cotriguaçu, Fazenda [“Faz”] São Nicolau; pitfall baited with fungus; 9°49 ′8 ″S, 58°15 ′40 ″W;  17–21 May 2012; F.Z. Vaz-de-Melloleg.; CEMT•  1 ♂; Cotriguaçu, Fazenda [“Faz”] São Nicolau, primary forest; flight interception trap[“FIT”]; 9°50 ′24 ″S, 58°15 ′10 ″W;  250 ma.s.l.;  6 May 2009; Vaz-de-Melloleg.; CEMT•  1 ♂, 1 ♀; Fazenda São Nicolau(site 6), primary forest; flight int. trap; 9°49 ′16 ″S, 58°15 ′33 ″W;  220 ma.s.l.;  1 Dec. 2013; F. Génierleg.; CMNC 2013-118•  1 ♀; Fazenda São Nicolau(site 5), riparian primary forest; flight int. trap; 9°51 ′56 ″S 58°13 ′09 ″W;  220 ma.s.l.;  3 Dec. 2013; F. Génierleg.; CMNC 2013-129•  1 ♀; Fazenda São Nicolau(site 6), primary forest; flight int. trap; 9°49 ′16 ″S, 58°15 ′33 ″W;  220 ma.s.l.;  3 Dec. 2013; F. Génierleg.; CMNC 2013-131.   PERU• 1 ♀; Madre de Dios, CICRA Los Amigos B.S; 12°34 ′1 ″S, 70°06 ′0 ″W;  250 ma.s.l.; 2005; T. Larsenleg.; TLC•  1 ♀; CICRA Field Station, trail 6, research plot; blue pan trap; 12.55207° S, 70.10962° W;  295 ma.s.l.;  7–9 Jun. 2011; Chabooteam leg.; SEMC PER-11-PTB-009•  1 ♀; Rio Tambopata, Jorge Chavez; flight intercept trap; 12°38 ′59 ″S, 69°06 ′24 ″W,  230 ma.s.l.;  28–29 Sep. 1999; T. Larsenleg.; CMNC•  1 ♀; Rio Palma Real Grande, Limon Camp; flight intercept trap; 12°32 ′20 ″S, 68°51 ′40 ″W;  220 ma.s.l.;  9–10 Oct. 1999; T. Larsenleg.; CMNC•  1 ♀; Rio Palma Real Grande, Limon Camp; flight intercept trap; 12°32 ′20 ″S, 68°51 ′40 ″W;  220 ma.s.l.;  13–14 Oct. 1999; T. Larsenleg.; CMNC.   Type locality Guajará-Mirim, Rondônia, Brazil.    Description ( holotype) MEASUREMENTS. Length: 10 mm. HEAD. Surface densely and distinctly punctated; with rectangular frontoclypeal carina, length greater than four-fifths of the interocular distance. PRONOTUM. Completely punctate, with dense and distinct punctation, becoming more accentuated towards the centre of the disc, regularly convex with two medial projections anteriorly, one on each side. HYPOMERON. Hypomeral carina slightly curved toward edge.   Fig. 7.  Canthidium stofeli sp. nov., holotype (CEMT) A. Dorsal view. B. Lateral view of aedeagus. C. Dorsal view of aedeagus. Scale bars: 1 mm. METAVENTRITE. With simple and shallow punctation, laterobasal region with stronger punctation, lateral region with microsculpture obscurely variolate, almost semicircular. MESEPIMERON AND MESEPISTERNUM. With shallow punctation, stronger microsculpture obscurely variolate, almost semicircular. HIND LEG. Metafemora with a carina along posterior edge, anterior edge with setal fringe, surface with fine punctation. Metatibiae elongated and gradually dilated towards the apex; external margin serrated. AEDEAGUS. In dorsal view, parameres with a strong concave angulation and abruptly curved apex.   Variation and sexual dimorphism Length ranging from 8.87 to 11.11 mm. The projections on the pronotal disc vary according to the size of the specimen, being allometrically developed as the size of the body increases. Males with ventrite VI wider medially, female ventrite VI with the same length throughout. Males also have protibial spur flattened, while in females the protibial spur is narrowed and pointed.    Geographic distribution The specimens examined came from Brazil( Mato Grosso, Rondônia, Pará) and Peru( Madre de Dios). The species occurs in the southern region of the Amazon biome, corresponding to the Southeastern Amazonian dominion and Boreal Brazilian dominion. 2252301152 2010-02-24 CEMT F. Coletti Brazil Guajara 138 -10.749339 Guajara / Mirim: Proprietario: Fandinho 1 -65.31676 Mirim 12 13 139 1 Rondonia holotype 5006659324 1997-02 CEMT F. Vaz-de-Mello Brazil Rio Branco -9.955491 Fazenda [" Faz "] Catuaba 1 -65.86548 Rio Branco 12 13 1 1 Acre paratype 2252301106 2010-01-28 flight interception trap CEMT F. Coletti Brazil 170 -10.329753 Sitio Coletti 1 -4.7534723 Guajara-Mirim 12 13 1 1 Rondonia paratype 2252301161 2016-07-09 2016-07-16 2016-07-09 CEMT P. Silva Brazil Empresa Agropalma Tailandia 12 13 1 1 Para paratype 2252301181 2017-03-24 pig trap CEMT F. Franca Brazil 132 -3.2963889 Belterra 21 -54.963333 12 13 1 1 paratype 2252301209 2017-03-24 pig trap CEMT F. Franca Brazil 132 -3.2963889 Belterra 21 -54.963333 12 13 1 1 paratype 2252301135 2009-02 flight interception trap CEMT R. Andrade Brazil Querencia -12.650891 Fazenda Sao Luiz, Amazon Forest 1 -52.3526 Querencia 12 13 1 1 Mato Grosso paratype 2252301154 2010-02-20 CEMT M. F. Souza Brazil Claudia -11.581667 Claudia, Fazenda [" Faz "] Continental 21 -55.2875 Fazenda 12 13 2 2 paratype 2252301113 2009-07-11 2009-07-13 2009-07-11 BMNH F. Z. Vaz-de-Mello Brazil -9.838611 Matinha 21 -58.25083 Cotriguacu 12 13 1 1 paratype 2252301133 [805,1292,1464,1492] 2009-07-11 2009-07-13 2009-07-11 CEMT F. Z. Vaz-de-Mello Brazil -9.838611 Matinha 21 -58.25083 Cotriguacu 12 13 1 1 paratype 2252301191 2009-07-12 2009-07-14 2009-07-12 flight interception trap CEMT F. Z. Vaz-de-Mello Brazil Cotriguacu -9.819166 Fazenda [" Faz "] Sao Nicolau 21 -58.263054 Cotriguacu 12 13 2 2 paratype 2252301098 [809,1310,1534,1563] 2009-07-12 2009-07-14 2009-07-12 flight interception trap BMNH F. Z. Vaz-de-Mello Brazil Cotriguacu -9.819166 Sao Nicolau 21 -58.263054 Fazenda 12 13 1 1 paratype 2252301097 2012-05-17 2012-05-21 2012-05-17 CEMT F. Z. Vaz-de-Mello Brazil Cotriguacu -9.818889 Fazenda [" Faz "] Sao Nicolau 21 -58.261112 Cotriguacu 12 13 1 1 paratype 2252301195 2009-05-06 flight interception trap CEMT Vaz-de- Mello Brazil Cotriguacu 250 -9.84 Fazenda [" Faz "] Sao Nicolau 21 -58.252777 Cotriguacu 12 13 1 1 paratype 2252301180 2013-12-01 CMNC F. Genier Brazil 220 -9.821111 Fazenda Sao Nicolau 21 -58.259167 12 13 CMNC 2013-118 2 1 1 paratype 2252301210 2013-12-03 CMNC F. Genier Brazil 220 -9.865556 Fazenda Sao Nicolau 21 -58.219166 12 13 CMNC 2013-129 1 1 paratype 2252301117 2013-12-03 CMNC F. Genier Brazil 220 -9.821111 Fazenda Sao Nicolau 21 -58.259167 12 13 CMNC 2013-131 1 1 paratype 2252301111 TLC T. Larsen Peru 250 -12.566944 CICRA Los Amigos B. S 21 -70.1 12 13 1 1 Madre de Dios 2252301142 2011-06-07 2011-06-09 2011-06-07 blue pan trap SEMC Chaboo Peru 295 -12.55207 CICRA Field Station 1 -70.10962 12 13 PER-11-PTB-009 1 1 2252301116 1999-09-28 1999-09-29 1999-09-28 flight intercept trap CMNC T. Larsen Peru 230 13 14 -12.649722 Jorge Chavez 21 -69.10667 Rio Tambopata 12 13 1 1 2252301233 1999-10-09 1999-10-10 1999-10-09 flight intercept trap CMNC T. Larsen Peru 220 -12.538889 Limon Camp 21 -68.86111 Rio Palma Real Grande 13 14 1 1 2252301149 1999-10-13 1999-10-14 1999-10-13 flight intercept trap CMNC T. Larsen Peru 220 -12.538889 Limon Camp 21 -68.86111 Rio Palma Real Grande 13 14 1 1