Pachyrrhiza argentata Oldroyd, 1968: 383 Webb et al. (2013: 15) Revision of the Patagonian stiletto fly genus Pachyrrhiza Philippi (Therevidae Agapophytinae) Irwin, Michael E. Winterton, Shaun L. Zootaxa 2021 2021-05-25 4975 2 273 305 6THRF Oldroyd Oldroyd [151,535,735,762] Insecta Therevidae Pachyrrhiza Animalia Diptera 4 277 Arthropoda species argentata  ( Figs 4–8, 30A, E, 31E, 33A, 35)      Pachyrrhiza argentata Oldroyd, 1968: 383—  Webb et al.(2013: 15)[catalogue].   Diagnosis. Parafacial with distinct patch of black setae adjacent to gena; hind femur with one or two subapical anteroventral macrosetae; prosternum with setae present or absent; wing mottled; male abdomen with silver velutum pubescence present; mid and hind coxal pile absent on posterior surface (rarely present on mid coxa).   Redescription. Body length: 5.0– 6.8 mm. Head with frontal pubescence light brown dorsally, silver around antennal base, dark brown band above antenna in female; parafacial with small dark patch of setae adjacent to gena; genal setae pale; male frontal vestiture consisting of patch of short setae above antenna, shorter than setae on scape; female frontal vestiture consisting of scattered dark, short to moderate length setae; occiput grey with brown pubescence; antennal scape length slightly longer than maximum width, brown, overlain with grey pubescence, numerous erect, large black macrosetae; flagellum brown with grey pubescence. Thorax with scutum pubescent grey with broken, dark brown stripes, some setal bases dark brown, two dark medial stripes anteriorly joining posteriorly, lateral stripes irregular, vestiture as numerous erect black setae, longer and denser in male; prosternum with setae present or absent medially; pleuron dark, overlain with grey pubescence; katatergite setae all white; posterior surface of mid coxa without setae, rarely with a few white setae present; femora dark brown (usually males) to dark yellow, suffused with brown-black dorsally (female), short dark setae admixed with elongate pale setae ventrally; hind femur with 1 or 2 subapical anteroventral macrosetae; tibiae yellow, dark grey-brown apically; tarsi dark yellow with apices brown; wing mottled dark infuscate, fenestrate in some cells; costal cell brown infuscate basally, yellow infuscate distally; scutal chaetotaxy (macrosetae pairs): notopleural 3, supra alar 1, post alar 1, dorsocentral 3–5, scutellar 2. Abdomen of male abdomen base colour dark brown dorsally, yellow laterally, silver velutum on tergites 2–7, male vestiture mostly elongate pale setae, denser laterally; terminalia dark yellow; female abdominal markings mostly dark brown, posterior margin pale, broader laterally, intersegmental membrane distinctly pale coloured, well defined. Male and female genitalia as per genus description except male with epandrium with dark medial stripe; aedeagus with distiphallus narrow apically.   Comments.  Pachyrrhiza argentatais easily distinguished from other  Pachyrrhizaspeciesby the mottled wing pattern, 1–2 subapical anteroventral macrosetae on the hind femur and male abdomen with silver velutum. It appears closely related to  P. parargentata  sp. n.and  P. argentatoides  sp. n.and while their respective distributions overlap,  P. argentatais more widely distributed along the Argentinian eastern slopes of the southern Andes mountain range ( Fig. 35).   Type material.   Holotypemale, ARGENTINA:  Río Negro Province: San Carlos de Bariloche, San Ramon[ -41.1373, -71.2482],  22.iv.1966, [reared] pupated  14.xi.1966, emerged  8.xii.1966( NHM).   Paratypes. ARGENTINA:  Río Negro Province: 6 females, 2 males, same data as holotype [various dates, also reared] ( NHM).  Neuquen Province: 1 male, Estancia Fortin Chacabuco[approximated as -41.0044, -71.1638],  3.iv.1966, [reared] pupated  25.xi.1966, emerged  23.xii.1966( NHM). (Types examined)  Other material examined.  ARGENTINA:  Chubut Province: 2 females, El Maitén[ -42.0517, -71.1667],  18.XI.1966, L. Pena( MEI167328, 167352; CSCA);  1 male, Esquel[ -42.9099, -71.2849],  590 m, hand netted on walkway ( CSCA);  7 males, 4 females, Cholila, Chacra of Stephen Bussey, Malaisein steep gorge [-42.4503, -71.45],  23–31.XII.2005, M.E. Irwin,  560 m( CSCA);  2 males, 13 females, Sierra de San Berardo, 57 km W. Sarmiento[ - 45.4555, -69.682],  540 m, Malaisein small wash,  21–31.XII.2005, M.E. Irwin( CSCA);  2 males, 1 female, Parque Nacional Alerces,  530 m, Lago Futalaufquen, nr. Hosteria Quime Quipan[ -42.8611, -71.6026],  22.XII.2005, Malaise,  Nothophagus, M.E. Irwin( CSCA).   Mendoza Province: 1 male, 1 female, Ref. Gral. Alvarado[approximated as -32.895, -68.842], Col. Duret, 13.XII.53 ( MNHN).   Rio Negro Province: 45 males, 57 females, Nahuel Huapi National Park, 2 km S. Rincon Chico, Malaise in Patagoniansteppe [ -41.0025, -71.1268],  24–31.XII.2005, M.E. Irwin,  890 m( CSCA).   Neuquen Province: 2 males, Parque Nacional Lanin, 4 km E. Estancia Los Helechosat Arroyo Blanco[ -39.7836, -71.0345], Malaisein steppe prairie,  25.XII.2005–  2.I.2006, M.E. Irwin,  990 m( CSCA).  12 males, 20 females, 9 kmW. highway 23 on highway 60, Estancia Mamuil Malal Lolen[ -39.7555, -71.1248], Malaisein sand prairie,  26.XII.2005–  3.I.2006, M.E. Irwin,  890 m( CSCA);  24 females, Dept. Collon Cura, 54 kmS. La Rinconada, Malaisein dry wash [ -40.4038, -70.6538],  25–27.XII.2005, M.E. Irwin,  670 m( CSCA).  CHILE:  Aysén Province: 1 male, 1 female, 5.8 kmW. Chile Chico[ -46.5483, -71.8047],  450 m,  23.XI.1966, E.I. Schlinger, M.E. Irwin( MEI167363, 167323; CSCA);  7 males, 2 females, 8 kmW. Chile Chico[ -46.5517, -71.8355],  540 m,  22.XI.1966, E.I. Schlinger, M.E. Irwin( MEI167366–7, 167371, 167373–6, 167327, 167329; CSCA).   Los Rios Province: 1 female, Panguipulli[approximated as -39.638, -72.335],  Jan.1945( MEI167474; CSCA).   Malleco Province: 1 female, Marimenuco[approximated as -38.712, -71.107],  10.XII.1959, L.E. Peña,  1100 m( MEI167460; CSCA). 3128592327 1966-04-22 1966-12-08 1966-04-22 NHM Argentina -41.1373 San Ramon 6 -71.2482 San Carlos de Bariloche 4 277 1 Rio Negro holotype 3128592353 1966-04-03 1966-12-23 1966-04-03 NHM Argentina 5 278 -41.0044 Estancia Fortin Chacabuco 6 -71.1638 San Carlos de Bariloche 4 277 9 6 3 Rio Negro paratype 3128592308 1966-11-18 CSCA Pena Argentina -42.0517 El Maiten 6 -71.1667 5 278 MEI167328, 167352 2 2 Chubut 3128592351 CSCA Argentina 590 -42.9099 Esquel 6 -71.2849 5 278 1 1 Chubut 3128592345 2005-12-23 2005-12-31 2005-12-23 CSCA M. E. Irwin Argentina Cholila 560 Malaise Chacra of Stephen Bussey 5 278 11 4 7 Chubut 3128592303 2005-12-21 2005-12-31 2005-12-21 CSCA W. Sarmiento & M. E. Irwin Argentina 540 -45.4555 Malaise 39 -69.682 Sierra de San Berardo 5 278 15 13 2 Chubut 3128592306 2005-12-22 CSCA M. E. Irwin Argentina Parque Nacional Alerces 530 -42.8611 Hosteria Quime Quipan 6 -71.6026 Lago Futalaufquen 5 278 3 1 2 Chubut 3128592313 MNHN Col. Duret Argentina -32.895 Ref. Gral. Alvarado 72 -68.842 5 278 2 1 1 Mendoza 3128592312 2005-12-24 2005-12-31 2005-12-24 CSCA S. Rincon Chico & Irwin Argentina 890 -41.0025 Malaise in Patagonian 6 -71.1268 Nahuel Huapi National Park 5 278 102 57 45 Rio Negro 3128592346 2005-12-25 2006-01-02 2005-12-25 CSCA E. Estancia Los Helechos & M. E. Irwin Argentina Parque Nacional Lanin 990 -39.7836 Malaise 6 -71.0345 Arroyo Blanco 5 278 2 2 Neuquen 3128592304 2005-12-26 2006-01-03 2005-12-26 CSCA M. E. Irwin Argentina 890 -39.7555 Malaise 6 -71.1248 Estancia Mamuil Malal Lolen 5 278 32 20 12 Neuquen 3128592335 2005-12-25 2005-12-27 2005-12-25 CSCA Irwin Argentina Dept. Collon Cura 670 -40.4038 Malaise 6 -70.6538 La Rinconada 5 278 24 24 Neuquen 3128592333 1966-11-23 CSCA Schlinger & M. E. Irwin Chile 450 -46.5483 Chile Chico 6 -71.8047 5 278 MEI167363, 167323 2 1 1 Aysen 3128592328 1966-11-22 CSCA Schlinger & Irwin Chile 540 -46.5517 Chile Chico 6 -71.8355 5 278 MEI167366-7, 167371, 167373-6, 167327, 167329 9 2 7 Aysen 3128592331 1945-01 CSCA Chile -39.638 Panguipulli 69 -72.335 5 278 MEI167474 1 1 Los Rios 3128592316 1959-12-10 CSCA Pena Chile 1100 -38.712 Marimenuco 70 -71.107 5 278 MEI167460 1 1 Malleco Province