Pachyrrhiza pictipennis Philippi, 1865: 704 Reed (1888: 294) Malloch (1932: 248) Stuardo-Ortiz (1946: 86) Webb et al. (2013: 15) Baryphora pictipennis (Philippi) Kertész (1909: 167) Dialineura pictipennis (Philippi) Kröber (1911: 489) Revision of the Patagonian stiletto fly genus Pachyrrhiza Philippi (Therevidae Agapophytinae) Irwin, Michael E. Winterton, Shaun L. Zootaxa 2021 2021-05-25 4975 2 273 305 75HRF Philippi Philippi [151,545,438,465] Insecta Therevidae Pachyrrhiza Animalia Diptera 17 290 Arthropoda species pictipennis  ( Figs 2, 3, 22–25, 30D, G, 31F, 32, 34B, 35)      Pachyrrhiza pictipennis Philippi, 1865: 704—  Reed (1888: 294)[catalogue];  Malloch (1932: 248)[redescription];  Stuardo-Ortiz (1946: 86)[catalogue];  Webb et al.(2013: 15)[catalogue]; Winterton et al.(2016: fig. 3) [phylogeny].    Baryphora pictipennis(Philippi)—  Kertész (1909: 167)[catalogue].    Dialineura pictipennis(Philippi)—  Kröber (1911: 489)[redescription], (1913: 24) [checklist].   Diagnosis. Parafacial with patch of black setae adjacent to gena; hind femur with 3–6 anteroventral macrosetae along segment length; prosternum with setae medially; wing dark infuscate with fenestrations; male without silver velutum pubescence on abdomen; mid coxa without setae on posterior surface; femora brown.   Redescription. Body length: 5.5–8.0 mm. Head with frons silver-grey pubescent with dark brown markings, lighter brown dorsally in female, silver pubescent around antennal base, small patch of setae on frons just above antenna, similar in length to setae on scape, female with short to moderate length setae distributed over frons; occiput grey to brown pubescent; parafacial with small patch of dark setae adjacent to gena; genal setae pale; antennal scape length approximately twice maximum width, brown (male) or dark yellow, brown suffusion dorsally (female), overlain with grey pubescence admixed with numerous large, erect macrosetae; flagellum brown to dark yellow with brown pubescence. Thorax with scutum and scutellum grey pubescent with, narrow dark medial stripe, lateral stripes broken to sub-tessellate, numerous erect black setae (longer and denser anteriorly in male); prosternum with pale setae present; pleuron with silver-grey pubescence, katatergite setae all white, sometimes admixed with black setae; mid coxa without setae on posterior surface; femora dark brown, sometimes with apices yellow, short dark setae on all surfaces admixed with relatively elongate dark setae ventrally, hind femur with 3–6 (sometimes more) anteroventral macrosetae distributed along entire length; tibiae yellow, dark grey-brown apically and basally; tarsi dark yellow with apices brown; wing dark infuscate, fenestrate in some radial and medial cells, costal cell brown infuscate; scutal chaetotaxy: notopleural, 4; supra alar, 1; post alar, 1; dorsocentral, 4–5; scutellar, 2. Abdomen dark brown, slight yellowish suffusion medially, intersegmental membrane pale, narrow male without silver velutum on tergites, male vestiture as short dark setae dorsally, longer pale setae laterally, female setae uniformly distributed, short, dark, erect; terminalia brown. Male and female genitalia as per genus description except: epandrium brownish, darker medially, distiphallus narrow distally.   Comments. Webb et al.(2013)noted that the syntypesdeposited in the National Museum in Santiago, Chilecould not be located yet could not be confirmed as destroyed. A ‘cotype’ determined by Kröber (1911: p. 489) as “  Dialineura pictipennisPhil.” is deposited in the USNM but cannot be part of the syntypeseries as it was collected decades after the original description was published.  Pachyrrhiza pictipennisis distinctive compared with other species in the genus based on the dark body colouration, extensive wing infuscation with small hyaline fenestrations distally and lack of silver velutum on the male abdomen; these are all evident in the original description and the figure plate provided by Philippi (1865: fig. 25); the identity of the species is not in question and a neotypeis not necessary at this time. This species appears to be locally abundant in central Chile.  Pachyrrhiza pictipennisappears closely related to  P. subpictipennis  sp. n.and  P. elegantissima  sp. n.based on wing pattern and presence of a series of anteroventral macrosetae along the hind femur.   Type material.  three syntypefemales, CHILE:  Santiago Province: Region Metropolitana de Santiago, Cordillere(location unknown, presumed lost).   FIGURE 25.  Pachyrrhiza pictipennis Philippi, 1865, adult female, lateral view.  Other material examined.  CHILE:  Arauco Province: 9 males, 1 female, Naluelbuta National Park,  2 kmE Cerro Anai[ -37.801, -73.0153], Malaisein Araucariaforest,  1–8.XII.2003, M.E. Irwin,  1253 m( MEI166325, 166319–20, 166321, 166323, 166328–9, 166336, 166342, 167019; CSCA);  1 male, Parque Nacional Nahuelbuta, Centro de Informaciones[-37.8166, -73.00],  8.XII.1994, S.L. Heydon, E. Arias( MEI116832; UCDC).   Biobío Province: 4 males, 2 females,  El Albanico, Ross& Michelbacher,  30.XII.1950[ -37.3322, -71.5225] ( MEI167452, 167463–5, 167467, 167476; CAS).   Choapa Province: 1 male, Quilimari[ -32.1136, -71.5035],  13.XII.1950, Ross& Michelbacher( MEI167475; CAS).   Concepción Province: 1 female, Concepción, Parque Bot. Hualpen[ -36.7845, -73.1021],  10–15.II.1970 Malaise( MEI112369; UCCC).   Curicó Province: 3 females, Caminoal Relvo, 5 km E. Potrero Grande[ -35.2061, -70.9627], Malaisein  Nothofagusforest,  14–24.X.2004, J.E. Barriga( CSCA);  10 males, 15 km E. Curicó, CerroHueca-Huecan [ -35.0665, -71.1226], Malaise,  400 m,  10–22.I.1998, J.E. Barriga( CSCA);  6 females, Fundo la Montana Esterola Palma at Rio Teno,  6 kmE Los Quenes[ -34.9669, -70.9606],  14.I.1967, M.E. Irwin( MEI167015, 167020–1, 167023, 167030, 167032; CSCA);  9 females, Estero la Jaula, Curicó[ -35.0145, -70.7465],  I.1964. L. Peña,  Nothofagus( MEI167180–88; CNC).   Elqui Province: 1 male, 1 female, Quebrada El Arrayán, 33 km S. La Villa[ -30.1951, -70.9338], Malaisein vegetated ravine,  13–21.XI.2003, M.E. Irwin,  653 m( CSCA);  1 male, 1 female, Quebrada El Arrayán, 15 km S. La Villa[ -30.1045, -70.9901], pan trapin damp wash,  1.XI.2003, F.D. Parker, M.E. Irwin,  472 m( CSCA).   Limari Province: 2 males,  8 kmN San Pedro de Quile[ -30.7506, -71.2531],  16.XI.1961, L. Peña( MEI167417, 167480; CNC).   Milipilla Province: 2 males, El Bollenar, 2 km N. Pichi Alhue[ -33.5729, -71.0215], Malaisein dry wash,  100 m,  1–10.I.2000, ME Irwin, E.I. Schlinger( CSCA).   Ñuble Province: 12 females, 7.5 km E. Recinto[ -36.8773, -71.5785], Malaisein dry riverbed,  760 m,  4–6.I.2000, M.E. Irwin, E.I. Schlinger( MEI118861, 118867, 121224; CSCA);  4 males, 2 kmN. Las Trancas[ -36.903, -71.481], Malaisein  Nothofagusforest,  6–9.I.2000, M.E. Irwin, E.I. Schlinger( MEI12195–8; CSCA);  3 males, 2 kmN. Las Trancas( -36.903, -71.481), hand netted,  4.I.2000, M.E. Irwin( MEI121264–6; CSCA);  1 male, 1 female, Termas de Chíllan, 8 km E. Las Trancas( -36.906, -71.410), hand netted,  9.I.2000, M.E. Irwin( MEI121215, 121267; CSCA).   Quillota Province: 9 males, Palma de Ocoa, Parque Nacional La Campana[ -32.9324, -71.0781], Malaisein hillside draw,  6–13.XI.1997, M.E. Irwin, E.I. Schlinger( MEI107556–64; CSCA).   Santiago Province: 10 males, 10 females, Queb. de la Plata, Rinconada, Maipú[ -33.561, -70.783],  510m, [various dates]  26.X.1966–  1.I.1967, M.E. Irwin( CAS);  2 males, 26 females, El Canelo[ -33.5788, -70.4499],  26.XI.1954, L.E. Peña( CNC);  5 males, 2 km E. Corral Quemado[-33.35,-70.3],  2250 m,  11.XII.1994, S.L. Heydon, E. Arias( MEI116834–8; UCDC);  2 females, Pilay, Rio Peuco, ca. 45 kmS. Santiago[ -33.9481, -70.6413],  23–24.XI.1981, D.R. Davis,  800 m( MEI167275, 167277; USNM).   Talca Province: female, Alto de Vilches,  26–28.I.1964, L. Peña,  Nothofagus( MEI167161; CNC);  1 male, El Radal[ -35.459, -71.024],  900 m,  26.XI.1957, L. Peña, ( MEI079761; CNC). 3128592341 Chile Cordillere 20 293 1 1 Region Metropolitana syntype 3128592320 2003-12-01 2003-12-08 2003-12-01 CSCA Malaise & M. E. Irwin Chile 1253 -37.801 Naluelbuta National Park 55 -73.0153 20 293 MEI166325, 166319-20, 166321, 166323, 166328-9, 166336, 166342, 167019 10 1 9 Araucania 3128592347 1994-12-08 UCDC S. L. Heydon & E. Arias Chile Centro de Informaciones Parque Nacional Nahuelbuta 20 293 MEI116832 1 1 Araucania 3128592319 1950-12-30 CAS Chile El Albanico -37.3322 Ross 6 -71.5225 El Albanico 20 293 MEI167452, 167463-5, 167467, 167476 6 2 4 Biobio 3128592349 1950-12-13 CAS Ross & Michelbacher Chile -32.1136 Quilimari 6 -71.5035 20 293 MEI167475 1 1 Choapa Province 3128592301 1970-02-10 1970-02-15 1970-02-10 UCCC Malaise Chile -36.7845 Parque Bot. Hualpen 6 -73.1021 Concepcion 20 293 MEI112369 1 1 Concepcion Province 3128592336 2004-10-14 2004-10-24 2004-10-14 CSCA J. E. Barriga Chile -35.2061 Relvo 6 -70.9627 Camino 20 293 3 3 Curico Province 3128592305 1998-01-10 1998-01-22 1998-01-10 CSCA J. E. Barriga Chile 400 -35.0665 Cerro 6 -71.1226 20 293 10 10 Curico Province 3128592318 1967-01-14 CSCA M. E. Irwin Chile 21 294 -34.9669 Rio Teno 6 -70.9606 Fundo la Montana Estero 20 293 MEI167015, 167020-1, 167023, 167030, 167032 6 6 Curico Province 3128592344 1964-01 CNC L. Pena Chile -35.0145 Curico 6 -70.7465 Estero la Jaula 21 294 MEI167180-88 9 9 Curico Province 3128592310 2003-11-13 2003-11-21 2003-11-13 CSCA M. E. Irwin Chile 653 -30.1951 Quebrada El Arrayan 6 -70.9338 21 294 2 1 1 Elqui Province 3128592307 2003-11-01 CSCA F. D. Parker & M. E. Irwin Chile 472 -30.1045 Quebrada El Arrayan 6 -70.9901 21 294 2 1 1 Elqui Province 3128592321 1961-11-16 CNC L. Pena Chile -30.7506 8 km N San Pedro de Quile 6 -71.2531 21 294 MEI167417, 167480 2 2 Limari Province 3128592322 2000-01-01 2000-01-10 2000-01-01 CSCA ME Irwin & E. I Schlinger Chile 100 -33.5729 El Bollenar 6 -71.0215 21 294 2 2 Milipilla Province 3128592311 2000-01-04 2000-01-06 2000-01-04 CSCA M. E. Irwin & E. I. Schlinger Chile 760 -36.8773 Nuble Province 6 -71.5785 21 294 MEI118861, 118867, 121224 12 12 Nuble Province 3128592342 MEI12195 2000-01-06 2000-01-09 2000-01-06 CSCA M. E. Irwin & E. I. Schlinger Chile -36.903 Las Trancas 70 -71.481 21 294 MEI12195-8 4 4 Nuble Province 3128592329 2000-01-04 CSCA M. E. Irwin Chile -36.903 Las Trancas 70 -71.481 21 294 MEI121264-6 3 3 Nuble Province 3128592348 2000-01-09 CSCA E. Las Trancas & M. E. Irwin Chile -36.906 Termas de Chillan 70 -71.41 21 294 MEI121215, 121267 2 1 1 Nuble Province 3128592350 1997-11-06 1997-11-13 1997-11-06 CSCA M. E. Irwin & E. I. Schlinger Chile -32.9324 Parque Nacional La Campana 6 -71.0781 Palma de Ocoa 21 294 MEI107556-64 9 9 Quillota Province 3128592338 1966-10-26 1967-01-01 1966-10-26 CAS M. E. Irwin Chile Queb. de la Plata 510 -33.561 Maipu 72 -70.783 Rinconada 21 294 20 10 10 Region Metropolitana 3128592315 [481,1432,690,717] 1954-11-26 CNC L. E. Pena Chile -33.5788 El Canelo 6 -70.4499 21 294 28 26 2 Region Metropolitana 3128592339 1994-12-11 UCDC S. L. Heydon & E. Arias Chile 2250 21 294 MEI116834-8 5 5 Region Metropolitana 3128592324 1981-11-23 1981-11-24 1981-11-23 USNM D. R. Davis Chile Pilay 800 -33.9481 Santiago 6 -70.6413 Rio Peuco 21 294 MEI167275, 167277 2 2 Region Metropolitana 3128592323 1964-01-26 1964-01-28 1964-01-26 CNC L. Pena Chile Alto de Vilches 21 294 MEI167161 1 Talca Province 3128592314 [392,1361,834,861] 1957-11-26 CNC L. Pena Chile 900 -35.459 El Radal 71 -71.024 21 294 MEI079761 1 1 Talca Province