Pachyrrhiza pictipennis Philippi, 1865: 704
Reed (1888: 294)
Malloch (1932: 248)
Stuardo-Ortiz (1946: 86)
Webb et al. (2013: 15)
Baryphora pictipennis (Philippi)
Kertész (1909: 167)
Dialineura pictipennis (Philippi)
Kröber (1911: 489)
Revision of the Patagonian stiletto fly genus Pachyrrhiza Philippi (Therevidae Agapophytinae)
Irwin, Michael E.
Winterton, Shaun L.
Zootaxa
2021
2021-05-25
4975
2
273
305
75HRF
Philippi
Philippi
[151,545,438,465]
Insecta
Therevidae
Pachyrrhiza
Animalia
Diptera
17
290
Arthropoda
species
pictipennis
( Figs 2, 3, 22–25, 30D, G, 31F, 32, 34B, 35)
Pachyrrhiza pictipennis Philippi, 1865: 704— Reed (1888: 294)[catalogue]; Malloch (1932: 248)[redescription]; Stuardo-Ortiz (1946: 86)[catalogue]; Webb et al.(2013: 15)[catalogue]; Winterton et al.(2016: fig. 3) [phylogeny]. Baryphora pictipennis(Philippi)— Kertész (1909: 167)[catalogue]. Dialineura pictipennis(Philippi)— Kröber (1911: 489)[redescription], (1913: 24) [checklist].
Diagnosis. Parafacial with patch of black setae adjacent to gena; hind femur with 3–6 anteroventral macrosetae along segment length; prosternum with setae medially; wing dark infuscate with fenestrations; male without silver velutum pubescence on abdomen; mid coxa without setae on posterior surface; femora brown.
Redescription. Body length: 5.5–8.0 mm. Head with frons silver-grey pubescent with dark brown markings, lighter brown dorsally in female, silver pubescent around antennal base, small patch of setae on frons just above antenna, similar in length to setae on scape, female with short to moderate length setae distributed over frons; occiput grey to brown pubescent; parafacial with small patch of dark setae adjacent to gena; genal setae pale; antennal scape length approximately twice maximum width, brown (male) or dark yellow, brown suffusion dorsally (female), overlain with grey pubescence admixed with numerous large, erect macrosetae; flagellum brown to dark yellow with brown pubescence. Thorax with scutum and scutellum grey pubescent with, narrow dark medial stripe, lateral stripes broken to sub-tessellate, numerous erect black setae (longer and denser anteriorly in male); prosternum with pale setae present; pleuron with silver-grey pubescence, katatergite setae all white, sometimes admixed with black setae; mid coxa without setae on posterior surface; femora dark brown, sometimes with apices yellow, short dark setae on all surfaces admixed with relatively elongate dark setae ventrally, hind femur with 3–6 (sometimes more) anteroventral macrosetae distributed along entire length; tibiae yellow, dark grey-brown apically and basally; tarsi dark yellow with apices brown; wing dark infuscate, fenestrate in some radial and medial cells, costal cell brown infuscate; scutal chaetotaxy: notopleural, 4; supra alar, 1; post alar, 1; dorsocentral, 4–5; scutellar, 2. Abdomen dark brown, slight yellowish suffusion medially, intersegmental membrane pale, narrow male without silver velutum on tergites, male vestiture as short dark setae dorsally, longer pale setae laterally, female setae uniformly distributed, short, dark, erect; terminalia brown. Male and female genitalia as per genus description except: epandrium brownish, darker medially, distiphallus narrow distally.
Comments. Webb et al.(2013)noted that the syntypesdeposited in the National Museum in Santiago, Chilecould not be located yet could not be confirmed as destroyed. A ‘cotype’ determined by Kröber (1911: p. 489) as “ Dialineura pictipennisPhil.” is deposited in the USNM but cannot be part of the syntypeseries as it was collected decades after the original description was published. Pachyrrhiza pictipennisis distinctive compared with other species in the genus based on the dark body colouration, extensive wing infuscation with small hyaline fenestrations distally and lack of silver velutum on the male abdomen; these are all evident in the original description and the figure plate provided by Philippi (1865: fig. 25); the identity of the species is not in question and a neotypeis not necessary at this time. This species appears to be locally abundant in central Chile. Pachyrrhiza pictipennisappears closely related to P. subpictipennis sp. n.and P. elegantissima sp. n.based on wing pattern and presence of a series of anteroventral macrosetae along the hind femur.
Type material. three syntypefemales, CHILE: Santiago Province: Region Metropolitana de Santiago, Cordillere(location unknown, presumed lost). FIGURE 25. Pachyrrhiza pictipennis Philippi, 1865, adult female, lateral view. Other material examined. CHILE: Arauco Province: 9 males, 1 female, Naluelbuta National Park, 2 kmE Cerro Anai[ -37.801, -73.0153], Malaisein Araucariaforest, 1–8.XII.2003, M.E. Irwin, 1253 m( MEI166325, 166319–20, 166321, 166323, 166328–9, 166336, 166342, 167019; CSCA); 1 male, Parque Nacional Nahuelbuta, Centro de Informaciones[-37.8166, -73.00], 8.XII.1994, S.L. Heydon, E. Arias( MEI116832; UCDC). Biobío Province: 4 males, 2 females, El Albanico, Ross& Michelbacher, 30.XII.1950[ -37.3322, -71.5225] ( MEI167452, 167463–5, 167467, 167476; CAS). Choapa Province: 1 male, Quilimari[ -32.1136, -71.5035], 13.XII.1950, Ross& Michelbacher( MEI167475; CAS). Concepción Province: 1 female, Concepción, Parque Bot. Hualpen[ -36.7845, -73.1021], 10–15.II.1970 Malaise( MEI112369; UCCC). Curicó Province: 3 females, Caminoal Relvo, 5 km E. Potrero Grande[ -35.2061, -70.9627], Malaisein Nothofagusforest, 14–24.X.2004, J.E. Barriga( CSCA); 10 males, 15 km E. Curicó, CerroHueca-Huecan [ -35.0665, -71.1226], Malaise, 400 m, 10–22.I.1998, J.E. Barriga( CSCA); 6 females, Fundo la Montana Esterola Palma at Rio Teno, 6 kmE Los Quenes[ -34.9669, -70.9606], 14.I.1967, M.E. Irwin( MEI167015, 167020–1, 167023, 167030, 167032; CSCA); 9 females, Estero la Jaula, Curicó[ -35.0145, -70.7465], I.1964. L. Peña, Nothofagus( MEI167180–88; CNC). Elqui Province: 1 male, 1 female, Quebrada El Arrayán, 33 km S. La Villa[ -30.1951, -70.9338], Malaisein vegetated ravine, 13–21.XI.2003, M.E. Irwin, 653 m( CSCA); 1 male, 1 female, Quebrada El Arrayán, 15 km S. La Villa[ -30.1045, -70.9901], pan trapin damp wash, 1.XI.2003, F.D. Parker, M.E. Irwin, 472 m( CSCA). Limari Province: 2 males, 8 kmN San Pedro de Quile[ -30.7506, -71.2531], 16.XI.1961, L. Peña( MEI167417, 167480; CNC). Milipilla Province: 2 males, El Bollenar, 2 km N. Pichi Alhue[ -33.5729, -71.0215], Malaisein dry wash, 100 m, 1–10.I.2000, ME Irwin, E.I. Schlinger( CSCA). Ñuble Province: 12 females, 7.5 km E. Recinto[ -36.8773, -71.5785], Malaisein dry riverbed, 760 m, 4–6.I.2000, M.E. Irwin, E.I. Schlinger( MEI118861, 118867, 121224; CSCA); 4 males, 2 kmN. Las Trancas[ -36.903, -71.481], Malaisein Nothofagusforest, 6–9.I.2000, M.E. Irwin, E.I. Schlinger( MEI12195–8; CSCA); 3 males, 2 kmN. Las Trancas( -36.903, -71.481), hand netted, 4.I.2000, M.E. Irwin( MEI121264–6; CSCA); 1 male, 1 female, Termas de Chíllan, 8 km E. Las Trancas( -36.906, -71.410), hand netted, 9.I.2000, M.E. Irwin( MEI121215, 121267; CSCA). Quillota Province: 9 males, Palma de Ocoa, Parque Nacional La Campana[ -32.9324, -71.0781], Malaisein hillside draw, 6–13.XI.1997, M.E. Irwin, E.I. Schlinger( MEI107556–64; CSCA). Santiago Province: 10 males, 10 females, Queb. de la Plata, Rinconada, Maipú[ -33.561, -70.783], 510m, [various dates] 26.X.1966– 1.I.1967, M.E. Irwin( CAS); 2 males, 26 females, El Canelo[ -33.5788, -70.4499], 26.XI.1954, L.E. Peña( CNC); 5 males, 2 km E. Corral Quemado[-33.35,-70.3], 2250 m, 11.XII.1994, S.L. Heydon, E. Arias( MEI116834–8; UCDC); 2 females, Pilay, Rio Peuco, ca. 45 kmS. Santiago[ -33.9481, -70.6413], 23–24.XI.1981, D.R. Davis, 800 m( MEI167275, 167277; USNM). Talca Province: female, Alto de Vilches, 26–28.I.1964, L. Peña, Nothofagus( MEI167161; CNC); 1 male, El Radal[ -35.459, -71.024], 900 m, 26.XI.1957, L. Peña, ( MEI079761; CNC).
3128592341
Chile
Cordillere
20
293
1
1
Region Metropolitana
syntype
3128592320
2003-12-01
2003-12-08
2003-12-01
CSCA
Malaise & M. E. Irwin
Chile
1253
-37.801
Naluelbuta National Park
55
-73.0153
20
293
MEI166325, 166319-20, 166321, 166323, 166328-9, 166336, 166342, 167019
10
1
9
Araucania
3128592347
1994-12-08
UCDC
S. L. Heydon & E. Arias
Chile
Centro de Informaciones
Parque Nacional Nahuelbuta
20
293
MEI116832
1
1
Araucania
3128592319
1950-12-30
CAS
Chile
El Albanico
-37.3322
Ross
6
-71.5225
El Albanico
20
293
MEI167452, 167463-5, 167467, 167476
6
2
4
Biobio
3128592349
1950-12-13
CAS
Ross & Michelbacher
Chile
-32.1136
Quilimari
6
-71.5035
20
293
MEI167475
1
1
Choapa Province
3128592301
1970-02-10
1970-02-15
1970-02-10
UCCC
Malaise
Chile
-36.7845
Parque Bot. Hualpen
6
-73.1021
Concepcion
20
293
MEI112369
1
1
Concepcion Province
3128592336
2004-10-14
2004-10-24
2004-10-14
CSCA
J. E. Barriga
Chile
-35.2061
Relvo
6
-70.9627
Camino
20
293
3
3
Curico Province
3128592305
1998-01-10
1998-01-22
1998-01-10
CSCA
J. E. Barriga
Chile
400
-35.0665
Cerro
6
-71.1226
20
293
10
10
Curico Province
3128592318
1967-01-14
CSCA
M. E. Irwin
Chile
21
294
-34.9669
Rio Teno
6
-70.9606
Fundo la Montana Estero
20
293
MEI167015, 167020-1, 167023, 167030, 167032
6
6
Curico Province
3128592344
1964-01
CNC
L. Pena
Chile
-35.0145
Curico
6
-70.7465
Estero la Jaula
21
294
MEI167180-88
9
9
Curico Province
3128592310
2003-11-13
2003-11-21
2003-11-13
CSCA
M. E. Irwin
Chile
653
-30.1951
Quebrada El Arrayan
6
-70.9338
21
294
2
1
1
Elqui Province
3128592307
2003-11-01
CSCA
F. D. Parker & M. E. Irwin
Chile
472
-30.1045
Quebrada El Arrayan
6
-70.9901
21
294
2
1
1
Elqui Province
3128592321
1961-11-16
CNC
L. Pena
Chile
-30.7506
8 km N San Pedro de Quile
6
-71.2531
21
294
MEI167417, 167480
2
2
Limari Province
3128592322
2000-01-01
2000-01-10
2000-01-01
CSCA
ME Irwin & E. I Schlinger
Chile
100
-33.5729
El Bollenar
6
-71.0215
21
294
2
2
Milipilla Province
3128592311
2000-01-04
2000-01-06
2000-01-04
CSCA
M. E. Irwin & E. I. Schlinger
Chile
760
-36.8773
Nuble Province
6
-71.5785
21
294
MEI118861, 118867, 121224
12
12
Nuble Province
3128592342
MEI12195
2000-01-06
2000-01-09
2000-01-06
CSCA
M. E. Irwin & E. I. Schlinger
Chile
-36.903
Las Trancas
70
-71.481
21
294
MEI12195-8
4
4
Nuble Province
3128592329
2000-01-04
CSCA
M. E. Irwin
Chile
-36.903
Las Trancas
70
-71.481
21
294
MEI121264-6
3
3
Nuble Province
3128592348
2000-01-09
CSCA
E. Las Trancas & M. E. Irwin
Chile
-36.906
Termas de Chillan
70
-71.41
21
294
MEI121215, 121267
2
1
1
Nuble Province
3128592350
1997-11-06
1997-11-13
1997-11-06
CSCA
M. E. Irwin & E. I. Schlinger
Chile
-32.9324
Parque Nacional La Campana
6
-71.0781
Palma de Ocoa
21
294
MEI107556-64
9
9
Quillota Province
3128592338
1966-10-26
1967-01-01
1966-10-26
CAS
M. E. Irwin
Chile
Queb. de la Plata
510
-33.561
Maipu
72
-70.783
Rinconada
21
294
20
10
10
Region Metropolitana
3128592315
[481,1432,690,717]
1954-11-26
CNC
L. E. Pena
Chile
-33.5788
El Canelo
6
-70.4499
21
294
28
26
2
Region Metropolitana
3128592339
1994-12-11
UCDC
S. L. Heydon & E. Arias
Chile
2250
21
294
MEI116834-8
5
5
Region Metropolitana
3128592324
1981-11-23
1981-11-24
1981-11-23
USNM
D. R. Davis
Chile
Pilay
800
-33.9481
Santiago
6
-70.6413
Rio Peuco
21
294
MEI167275, 167277
2
2
Region Metropolitana
3128592323
1964-01-26
1964-01-28
1964-01-26
CNC
L. Pena
Chile
Alto de Vilches
21
294
MEI167161
1
Talca Province
3128592314
[392,1361,834,861]
1957-11-26
CNC
L. Pena
Chile
900
-35.459
El Radal
71
-71.024
21
294
MEI079761
1
1
Talca Province