Thiele, 1928: 394
Brown (1980: 111
Pseudocleopatra voltana Mandahl-Barth, 1973: 279–281
Adding the West-African riverine component: Revision of the Recent freshwater snails belonging to Pseudocleopatra Thiele, 1928 (Caenogastropoda, Cerithioidea, Paludomidae)
Neiber, Marco T.
Kahl, Sandra M.
Wiggering, Benedikt
Glaubrecht, Matthias
Zootaxa
2019
2019-09-25
4674
3
301
328
Mandahl-Barth, 1973
Mandahl-Barth
1973
[151,727,1788,1815]
Gastropoda
Thiaridae
Pseudocleopatra
Animalia
11
312
Mollusca
species
voltana
Figs 8B, 11–14
Pseudocleopatra togoensis Thiele, 1928: 394(in part); Brown (1980: 111, fig. 63e). Pseudocleopatra voltana Mandahl-Barth, 1973: 279–281, fig. 3a, h; Brown (1980: 111–112); Brown & Kristensen (1993: 28, 51); Brown (1994: 128–129, 537, 580, fig. 61f); Glaubrecht (1996: 144–145, 147–148, fig. 24b, c); Kristensen & Stensgaard (2010a: e.T165376A6007457).
Typelocality.“ River Voltaat Daboya”; Ghana, Northern Region, West Gonja District, Daboya, White Volta, 9°32’06’’ N, 1°22’48’’ W. Type material. Holotype, DBL 2594-1and two paratypes, DBL 2594- 2,3, leg. F.S. McCullough. On the original label all three specimens are indicated as “Type”. Fromthe original description it is clear, however, that the specimen depicted here in Fig. 11Ais the holotypeoriginally selected by Mandahl-Barth (1973: 281). Additional paratypes: Allfrom Ghana: “Mogeyenga village, NE-Region” (not localised), DBL 2591, 7 specimens; “Apaso village” (= Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta), DBL 2592, 3 specimens; “Northeast region” (= Northeastern Ghana, without exact locality), DBL 2593, 1 specimen; “White Voltaat Daboya” (= Northern Region, West Gonja District, Daboya, White Volta, 9°32’06” N, 1°22’48” W), MRAC 799144, 2 specimens; “Fluss Voltabei Apaso ( Togo)” ( Volta Rivernear Apaso( Togo) = Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta). Other material examined.“Fluss Volta bei Apaso ( Togo)” ( River Voltanear Apaso( Togo) = Ghana, Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta), ZMB 190049, 1 specimen. Note that this specimen is a former syntypeof P. togoensis Thiele, 1928. Shell ( Fig. 11).The shell of P. voltanais reddish to dark brown or yellowish-ochre, sometimes with darker bands or a darker area near the columella, and has an ovate to elongate shape, with up to seven whorls and a pointed apex (if not eroded). The whorls are convex to almost flat, with 1–6 distinct spiral cords; in few specimens these ridges are only weakly developed. The aperture is holostome and ovate, higher than wide and slightly drawn out at the upper angle; outer margin sinuous in lateral view, columella slightly curved to almost straight and the basal margin evenly rounded (in the most mature specimens slightly produced). Umbilicus slit-like or closed.
Measurements ( n= 16).h: 8.52 ± 1.53 mm(range: 6.0–12.0 mm); w: 4.83 ± 0.60 mm(range: 3.7–5.8 mm); bw: 6.83 ± 0.89 mm(range: 4.8–8.3 mm); la: 4.11 ± 0.43 mm(range: 3.2–4.8 mm); wa: 2.72 ± 0.27 mm(range: 2.2–3.2 mm). Ratios ( n= 16).h/w: 1.75 ± 0.16 (range: 1.51–2.17); h/la: 2.06 ± 0.20 (range: 1.86–2.60). Operculum ( Fig. 8B).The operculum is corneous, with a paucispiral, rather large nucleus that can reach up to ⅓ of the maximum diameter and possesses a concentric margin. Radula( Figs 12–13, Table 2).Due to the lack of alcohol material, already prepared radula slides from the collection of the DBL were examined. Two slides of specimens from two different localities were examined (DBL 2592 and DBL 2594). The rachidian tooth is of squarish shape, with a straight to slightly V-shaped upper edge. The number of denticles is variable, with a prominent mesocone and 4–5 flanking denticles on either side, or mesocone not prominent, i.e. about the same size as the flanking denticles. The lateral teeth have relatively short but straight lateral extensions. The cutting edge bears 12–15 small denticles hardly differing in size and lacking a prominent mesocone. Inner and outer marginal teeth subequal in size and slightly curved. Both, the inner and outer marginal teeth bear 10–12 denticles each that are of approximately the same size.
Distribution ( Fig. 14). Pseudocleopatra voltanais known from the southern part of present day Lake Voltaand several localities along the White VoltaRiver.
Remarks. Pseudocleopatra voltanahas a more elongate shell than the remaining three Pseudocleopatraspecies. When the spiral sculpture is well-developed, the species is very distinctive and cannot be confused with any of its congeners. Only in few specimens the spiral sculpture was weak on the last whorl. Nothing is known about the ecology of the species and new records do not exist.
2418059372
[151,617,521,549]
Daboya
River
13
314
1
Volta
holotype
2418059384
Ghana
West Gonja District
9.535001
White
21
-1.38
Daboya
13
314
1
Northern
holotype
2423596352
[380,1281,593,621]
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
holotype
2423596350
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
holotype
2423596349
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
paratype
2423596341
[382,1432,773,801]
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
holotype
2423596345
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
holotype
2423596342
DBL, MRAC
F. S. McCullough. On & From & All
Ghana
Krachi East District
9.535001
Lake
15
0.118
Apase
13
314
DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144
13
Volta Region
holotype
2418059360
ZMB
Togo
River Volta
Krachi East District
0.118
Apaso
13
314
ZMB 190049
1
Volta Region
syntype