Thiele, 1928: 394 Brown (1980: 111 Pseudocleopatra voltana Mandahl-Barth, 1973: 279–281 Adding the West-African riverine component: Revision of the Recent freshwater snails belonging to Pseudocleopatra Thiele, 1928 (Caenogastropoda, Cerithioidea, Paludomidae) Neiber, Marco T. Kahl, Sandra M. Wiggering, Benedikt Glaubrecht, Matthias Zootaxa 2019 2019-09-25 4674 3 301 328 Mandahl-Barth, 1973 Mandahl-Barth 1973 [151,727,1788,1815] Gastropoda Thiaridae Pseudocleopatra Animalia 11 312 Mollusca species voltana   Figs 8B, 11–14     Pseudocleopatra togoensis  Thiele, 1928: 394(in part);  Brown (1980: 111, fig. 63e).   Pseudocleopatra voltana Mandahl-Barth, 1973: 279–281, fig. 3a, h; Brown (1980: 111–112); Brown & Kristensen (1993: 28, 51); Brown (1994: 128–129, 537, 580, fig. 61f); Glaubrecht (1996: 144–145, 147–148, fig. 24b, c); Kristensen & Stensgaard (2010a: e.T165376A6007457).     Typelocality.“ River Voltaat Daboya”;  Ghana, Northern Region, West Gonja District, Daboya, White Volta, 9°32’06’’ N, 1°22’48’’ W.  Type material.  Holotype, DBL 2594-1and two paratypes, DBL 2594- 2,3, leg. F.S. McCullough. On the original label all three specimens are indicated as “Type”. Fromthe original description it is clear, however, that the specimen depicted here in Fig. 11Ais the holotypeoriginally selected by Mandahl-Barth (1973: 281). Additional paratypes: Allfrom Ghana: “Mogeyenga village, NE-Region” (not localised), DBL 2591, 7 specimens; “Apaso village” (= Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta), DBL 2592, 3 specimens; “Northeast region” (= Northeastern Ghana, without exact locality), DBL 2593, 1 specimen; “White Voltaat Daboya” (= Northern Region, West Gonja District, Daboya, White Volta, 9°32’06” N, 1°22’48” W), MRAC 799144, 2 specimens; “Fluss Voltabei Apaso ( Togo)” ( Volta Rivernear Apaso( Togo) = Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta).   Other material examined.“Fluss Volta bei Apaso ( Togo)” ( River Voltanear Apaso( Togo) = Ghana, Volta Region, Krachi East District, Apase, 7°31’33.6”, 0°07’04.8” E, today inundated by Lake Volta), ZMB 190049, 1 specimen. Note that this specimen is a former syntypeof  P. togoensis Thiele, 1928.  Shell ( Fig. 11).The shell of  P. voltanais reddish to dark brown or yellowish-ochre, sometimes with darker bands or a darker area near the columella, and has an ovate to elongate shape, with up to seven whorls and a pointed apex (if not eroded). The whorls are convex to almost flat, with 1–6 distinct spiral cords; in few specimens these ridges are only weakly developed. The aperture is holostome and ovate, higher than wide and slightly drawn out at the upper angle; outer margin sinuous in lateral view, columella slightly curved to almost straight and the basal margin evenly rounded (in the most mature specimens slightly produced). Umbilicus slit-like or closed.   Measurements ( n= 16).h: 8.52 ± 1.53 mm(range: 6.0–12.0 mm); w: 4.83 ± 0.60 mm(range: 3.7–5.8 mm); bw: 6.83 ± 0.89 mm(range: 4.8–8.3 mm); la: 4.11 ± 0.43 mm(range: 3.2–4.8 mm); wa: 2.72 ± 0.27 mm(range: 2.2–3.2 mm).  Ratios ( n= 16).h/w: 1.75 ± 0.16 (range: 1.51–2.17); h/la: 2.06 ± 0.20 (range: 1.86–2.60).  Operculum ( Fig. 8B).The operculum is corneous, with a paucispiral, rather large nucleus that can reach up to ⅓ of the maximum diameter and possesses a concentric margin.   Radula( Figs 12–13, Table 2).Due to the lack of alcohol material, already prepared radula slides from the collection of the DBL were examined. Two slides of specimens from two different localities were examined (DBL 2592 and DBL 2594). The rachidian tooth is of squarish shape, with a straight to slightly V-shaped upper edge. The number of denticles is variable, with a prominent mesocone and 4–5 flanking denticles on either side, or mesocone not prominent, i.e. about the same size as the flanking denticles. The lateral teeth have relatively short but straight lateral extensions. The cutting edge bears 12–15 small denticles hardly differing in size and lacking a prominent mesocone. Inner and outer marginal teeth subequal in size and slightly curved. Both, the inner and outer marginal teeth bear 10–12 denticles each that are of approximately the same size.   Distribution ( Fig. 14).  Pseudocleopatra voltanais known from the southern part of present day Lake Voltaand several localities along the White VoltaRiver.   Remarks.  Pseudocleopatra voltanahas a more elongate shell than the remaining three  Pseudocleopatraspecies. When the spiral sculpture is well-developed, the species is very distinctive and cannot be confused with any of its congeners. Only in few specimens the spiral sculpture was weak on the last whorl. Nothing is known about the ecology of the species and new records do not exist. 2418059372 [151,617,521,549] Daboya River 13 314 1 Volta holotype 2418059384 Ghana West Gonja District 9.535001 White 21 -1.38 Daboya 13 314 1 Northern holotype 2423596352 [380,1281,593,621] DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region holotype 2423596350 DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region holotype 2423596349 DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region paratype 2423596341 [382,1432,773,801] DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region holotype 2423596345 DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region holotype 2423596342 DBL, MRAC F. S. McCullough. On & From & All Ghana Krachi East District 9.535001 Lake 15 0.118 Apase 13 314 DBL 2594-1, DBL 2594, 2,3, DBL 2591, DBL 2592, DBL 2593, MRAC 799144 13 Volta Region holotype 2418059360 ZMB Togo River Volta Krachi East District 0.118 Apaso 13 314 ZMB 190049 1 Volta Region syntype