Glenoleon rudda New, 1985b:46 Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae) Machado, Renato Jose Pires Oswald, John David Zootaxa 2020 2020-06-16 4796 1 1 322 (New, 1985) New 1985 [151,507,427,454] Insecta Myrmeleontidae Riekoleon Animalia Neuroptera 307 308 Arthropoda species rudda  ( Figs. 162, 165–166) http://zoobank.org/NomenclaturalActs/ 403A3264-34A2-4DC5-9CBA-08FA475EFC29    Glenoleon rudda New, 1985b:46(OD); New 1996:83(cat); Stange 2004:105(cat); Oswald 2018(cat).   Diagnosis.Forewing: first fork of RP in proximal half of wing, dark mediocubital line interrupted (not continuous); hind wing rhegmal area with a small brown infuscation; tibial spurs present; male ectoproct with ventral margin straight; female posterior gonapophysis longer than wide; female anterior gonapophysis a small plate.   Description.Lengths: forewing: 24–32 mm; hind wing: 21–29 mm. Head ( Figs. 165a–b): Labrumpale, with a line of elongate setae. Clypeuspale, with elongate black setae. Fronswith ventral half pale but dorsal half black; beset with short white setae. Genapale. Vertexraised; in anterior view pale, with a dorsal black line; in dorsal view mostly black except for some small pale areas posteriorly; set with short black setae. Ocularsetae absent. Antennaeclubbed; elongate,> 2x length of pronotum; distance between antennae wider than scape width; most flagellomeres black with distal half pale, two or three subapical flagellomeres entirely pale, apical flagellomeres entirely black, torular membrane yellow; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae. Mandiblespale with tip dark brown. Palpi, maxillary and labial with basal segment black but remaining areas pale, apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially. Thorax ( Fig. 165b): Pronotumslightly longer than wide; posterior margin as wide as anterior; subapical furrow present in some specimens; mostly black with two thin pale longitudinal bands around central region and two small- er ones near border; covered with short black setae. Mesonotummostly black, with small longitudinal pale lines around segments central area, some specimens with two small pale spots near wing bases, covered with short black setae. Metanotumentirely black, with some pale marks on anterior margin in some specimens, covered with short black setae. Pterothoracic pleurawith dorsal segments black, but ventral segments mostly pale, with some black marks, mainly on dorsal margin; covered with short white setae and few short black setae. Miller’s organ present. Wings ( Fig. 165c): Rather narrow with tip acute; anterior Banksian line present in both wings, posterior absent; veins set with short black setae, in general black, but the main longitudinal veins are intercalated by numerous short white regions. Male pilula axillarispresent. Forewingmembrane mostly hyaline with small brown marks generally concentrated on posterior margin and tip, and a longitudinal brown line on mediocubital area and costal area base; CuA fork located between RP origin and first fork; four presectoral crossveins, distal crossvein with a medial veinlet connecting with RP; costal area large, with few subcostal veinlets forked; posterior area narrower than prefork area. Hind wingmembrane mostly hyaline except for rhegmal region brown infuscated; MP fork located between RP origin and first fork; subcostal veinlets mostly simple but few distal ones forked; one presectoral crossvein. Legs ( Figs. 165a–b): All pairs of legs, femur elongate (> 2.5x length of coxa); tibia as long as femur but longer than tarsi; tibial spurs surpassing T1 apex; T2, T3 and T4 about same size, T1 slightly longer than T2, T5 about twice as long as T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with short black setae, and scattered long black setae, femur and tibia with few long white setae; T5 ventrally with two rows of thick, long, black setae. Prolegsense hair absent; femur tip and tibia base slightly enlarged; tibia with antennal cleaning setae ventroapically; coxa, tibia and tarsi entirely black; trochanter pale; femur mostly black with basal half of ventral and internal surfaces pale. Mesolegcolour pattern very similar to proleg, except coxa pale, and femur pale areas (larger in some specimens). Metalegfemur, tibia and T1 slightly longer than in other legs; coxa, trochanter, T1, most of femur and tibia (except apex) pale, remaining areas black. Abdomen: Mostly black, with scattered yellow marks generally on sternites; ectoproct pale with tip black; beset with short white setae.   FIGURE 165.  Riekoleon rudda: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal. Male Terminalia ( Figs. 166a–c): Ectoproctventral margin straight and slightly longer than dorsal margin in lateral view; set with elongate black setae. 9 thsternitewith posterior border straight in ventral view; with some black setae. Gonarcusbroad and arched, with medial region less sclerotized in posterior view; anterior margin elongated and rounded. Mediuncusabsent. Paramerelarge, with tip elongate, rounded, and curving inwards in posterior view. Female Terminalia ( Figs. 166d–e): Ectoproctposterior margin rounded, set with long black setae. Lateral gonapophysesrounded, shorter than ectoproct, beset with long black setae. 7 thsterniteelongate, with distal border rounded in ventral view; covered with short setae. Pregenital platerelatively small, shaped as upside down “U” in ventral view. Posterior gonapophysesbroad, elongate, beset with very long black setae and a large group of cavisetae at apex, and some small thickened black setae on internal margin. 9 thtergitewith a membranous digitiform process. Anterior gonapophysesvery small, a minute sclerotized plate, covered with short setae. Ventral membranegonapophyseal plates present and connected basally by a short transverse bar.   Distribution( Fig. 162). Australia: ACT, NSW*, QLD*, WA*. Known from scattered records in southeastern Queenslandand eastern New South Wales, with outliers in Western Australia.   FIGURE 166.  Riekoleon rudda: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female: d) terminalia, ventral (note pregenital plate hidden beneath sternite); e) terminalia, lateral. Scale bars = 0.5 mm.  Adult activity period.Records for October to May.   Biology.Unknown, larva unknown.  Name-bearing type.  Glenoleon rudda:  Holotype(by original designation), female, ANIC, examined. Type locality: Australia, Australian Capital Territory, Canberra(city). Fromoriginal description: “Types / Holotype, ♀, Australian Capital Territory, Canberra,  19.xii.1957, E. F. Riek( ANIC)”. Condition: good, terminalia dissected.  Additional material examined( 48♂, 62♀, 8?).   AUSTRALIA: ACT: Black Mountain: ii–iii.1967, light trap( 1♂, ANIC);  Canberra:   i.1952( 1♀, PT ANIC);   ii.1952( 1♀, PT ANIC);   i.1955( 1♀, SAMA);   ii.1957( 1♀, 1♂, PT ANIC);   ii.1957( 1♂, PT ANIC);   i.1960( 1♀, PT ANIC);   ii.1960( 11♀, 11♂, PT ANIC, 1♂, FSCA);   i.1961( 2♀, 1♂, PT ANIC);   ii.1961( 3♀, PT ANIC);   ii.1963( 9♀, 4♂, PT ANIC, 1♀, FSCA);   iii.1963( 4♀, 4♂, PT ANIC);   i.1966( 4♀, 5♂, PT ANIC);   New South Wales:  8 kmS of Mendooran, 31 o53’S– 149 o03’E,   12.ii.1992, G. Daniels& C.J. Burwell( 1♂, QMBA);  Nombinnie Nature Reserve: 18 air km SSE Mount Hope,  160m, 32.99613 oS–145.94006 oE±  70m,   3.i.2013, M.V., Oswald, Diehl& Machado, Oswald#626 ( 10♂, 7♀, TAMU);  Pillaga Scrub: 30:56S–149:23E,   10.ii.1997, M.S. & B.J. Moulds( 1♀, AMSA);   Queensland: Brisbane: (8?, QMBA);   2.ii.1925, H Hacker( 1♀, QMBA);   6.xi.1926( 2♀, QMBA);   21.i.1927( 1♂, QMBA);   3.ii.1927( 1♀, ZMUC);   12.iii.1927( 1♂, ZMUC; 1♀, QMBA);   18.x.1927( 1♀, ZMUC);   1.xi.1930( 1♀, QMBA);   29.x.1936( 1♀, MVM);   40 kmWNW of Goondiwindi,   17.xii.1983, M.S. & B.J. Moulds( 1♀, AMSA);   17 kmW of Miles,   11.xii.1990, T.A. Lambkin( 1♂, QMBA);  Millmerran:   12.xi.1987, at light, T.A. Lambkin( 1♂, QMBA);  Mount Coot-tha: Hilltoping, 27 o29’S– 152 o57’E,   10.v.1997, J.A. Skevington( 1♀, QMBA);  Nine Mile Creek:  14 kmNNW of Miles,   24.i.1990, M.S. & B.J. Moulds( 3♀, AMSA);  Rocky Barvia Eldsvold:   18–19.i.1996, at light, K.J. Lambkin( 2♀, QMBA);   Western Australia:  1 kmNE of Millstream, 21.35S– 117.04E,   6.xi.1970, Upton& Feehan( 1♂, ANIC);   23.x.1970, J.C. Car- dale, at light ( 1♂, ANIC);   iv.1971( 1♂, ANIC);  Norseman:   i.1966( 1♀, ANIC).   Comments.  Riekoleon ruddacan easily be distinguished from other  Riekoleonspecies by the colour pattern of its wings, the broad costal area of its forewing, and the general shape of its male and female terminalia. For phylogenetic relationships see Fig. 4. 1957-12-19 ANIC E. F. Riek Australia From Canberra 310 311 1 1 Australian Capital Territory holotype ANIC Australia Black Mountain 310 311 1 1 [302,409,369,395] Australia Canberra 310 311 1 [420,677,368,395] 1952-01 PT, ANIC Australia 310 311 1 1 [689,954,368,395] 1952-02 PT, ANIC Australia 310 311 1 1 [966,1197,368,395] 1955-01 SAMA Australia 310 311 1 1 1957-02 PT, ANIC Australia 310 311 2 1 1 [246,516,404,431] 1957-02 PT, ANIC Australia 310 311 1 1 [530,792,405,431] 1960-01 PT, ANIC Australia 310 311 1 1 [806,1291,405,431] 1960-02 ANIC, FSCA Australia 310 311 23 11 12 1961-01 PT, ANIC Australia 310 311 3 2 1 [339,611,441,467] 1961-02 PT, ANIC Australia 310 311 3 3 [625,1087,441,467] 1963-02 ANIC, FSCA Australia 310 311 14 10 4 [1101,1432,441,467] 1963-03 PT, ANIC Australia 310 311 8 4 4 [151,467,476,503] 1966-01 PT, ANIC Australia 310 311 9 4 5 [481,1197,476,503] Australia -31.883333 8 km S of Mendooran 1213 149.05 310 311 1 New South Wales 1992-02-12 QMBA Daniels & C. J. Burwell Australia 310 311 1 1 Australia 160 Mount Hope Nombinnie Nature Reserve 310 311 1 [378,1260,549,575] 2013-01-03 TAMU Oswald & Diehl & Machado Australia 310 311 17 7 10 Australia Pillaga Scrub 310 311 1 [353,864,585,611] 1997-02-10 AMSA Moulds Australia 310 311 1 1 [875,1292,584,611] QMBA Australia Brisbane 310 311 1 Queensland 1925-02-02 QMBA Hacker Australia 310 311 1 1 [411,682,621,647] 1926-11-06 QMBA Australia 310 311 2 2 [695,967,621,647] 1927-01-21 QMBA Australia 310 311 1 1 [980,1242,621,647] 1927-02-03 ZMUC Australia 310 311 1 1 1927-03-12 ZMUC, QMBA Australia 310 311 2 1 1 [415,693,657,683] 1927-10-18 ZMUC Australia 310 311 1 1 [707,983,657,683] 1930-11-01 QMBA Australia 310 311 1 1 [997,1265,657,683] 1936-10-29 MVM Australia 310 311 1 1 Australia 40 km WNW of Goondiwindi 310 311 1 [348,889,693,719] 1983-12-17 AMSA Moulds Australia 310 311 1 1 [903,1120,693,719] Australia 17 km W of Miles 310 311 1 1990-12-11 QMBA Lambkin Australia 310 311 1 1 [317,449,729,755] Australia Millmerran 310 311 1 [458,1001,729,755] 1987-11-12 QMBA Lambkin Australia 310 311 1 1 Australia -27.483334 Hilltoping 1237 152.95 Mount Coot-tha 310 311 1 [271,743,765,791] 1997-05-10 QMBA Skevington Australia 310 311 1 1 [756,1218,765,791] Australia Nine Mile Creek 310 311 1 1990-01-24 AMSA Moulds Australia 310 311 3 3 [460,736,801,827] Eldsvold Australia Rocky Bar 310 311 1 [748,1324,801,827] 1996-01-18 1996-01-19 1996-01-18 QMBA Lambkin Australia 310 311 2 2 Australia -21.35 1 km NE of Millstream 758 117.04 310 311 1 Western Australia [751,1200,837,863] 1970-11-06 ANIC Upton & Feehan Australia 310 311 1 1 1970-10-23 ANIC Australia 310 311 1 1 [447,675,873,899] 1971-04 ANIC Australia 310 311 1 1 [686,806,873,899] Australia Norseman 310 311 1 [816,1032,873,899] 1966-01 ANIC Australia 310 311 1 1