Glenoleon radialis Banks, 1913:224 Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae) Machado, Renato Jose Pires Oswald, John David Zootaxa 2020 2020-06-16 4796 1 1 322 (Banks, 1913) Machado & Oswald 2020 Banks 1913 [151,610,1773,1800] Insecta Myrmeleontidae Megagonoleon Animalia Neuroptera 239 240 Arthropoda species radialis comb. nov.  ( Figs. 119, 124–125) http://zoobank.org/NomenclaturalActs/ 94ECC4EC-4A32-4A54-B4C1-A03A18C9B0A4    Glenoleon radialis Banks, 1913:224(OD); Stange 1976:306(cat); New 1985b:33(key, rd, ill); New 1996:84(cat); Stange 2004:106(cat); Oswald 2018(cat).   Diagnosis.Body mostly black; clypeus mostly dark; hind wing with some radial crossveins brown margined; tibial spurs present; hind wing rhegmal and stigmal marks about same size; posterior gonapophysis elongate and narrow.   Description.Lengths: forewing: 33–36 mm; hind wing: 30–33 mm. Head ( Figs. 124a–b): Labrumblack centrally and pale marginally, set with a line of elongate setae. Clypeuspale with central brown marks; set with few elongate black setae. Fronspredominantly black, except for ventrolateral margin pale; some specimens with area between antennae pale; beset with short white setae. Genapale. Vertexraised; in anterior view pale, with a median black line interrupted centrally; in dorsal view mostly black except by some pale areas around central region posteriorly; set with short black or white setae. Ocularsetae very short, black or white. Antennaeclubbed; elongate,> 2x length of pronotum; distance between width; entirely dark-brown to black; torular membrane yellow; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae. Mandiblesdark brown, with some pale areas externally. Palpi, maxillary and labial dark brown with distal margin of segments generally pale, apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially.   FIGURE 124.  Megagonoleon radialis: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal. Thorax ( Fig. 124b): Pronotumabout as long as wide; posterior margin as wide as anterior; subapical furrow present in some specimens; generally pale with a broad black longitudinal band on midline (flanked on each side by a thinner pale longitudinal band), margins generally black; set with many short or long black setae, and some long white setae mostly located at margins. Mesonotumgenerally black, with some small longitudinal yellow bands on each side, scutellum mostly pale with black dorsal marks, but entirely black in some specimens; covered with short black or white setae. Metanotumusually black with some pale areas, mostly at sclerites margins; covered with short white setae. Pterothoracic pleurablack with some pale areas generally between sclerites; set with white setae; Miller’s organ present.   FIGURE 125.  Megagonoleon radialis: male: a) terminalia, lateral; b) genitalia, lateral; c) genitalia, posterior; female: d) terminalia, lateral; e) pregenital plate, posterior. Scale bars = 0.5 mm. Wings ( Fig. 124c): Fairly broad; anterior and posterior Banksian lines present in both wings; veins mostly brown but intercalated with white spots, set with short black setae. Male pilula axillarispresent. Forewingmembrane mostly hyaline with brown marks generally concentrated at tip, pterostigma, and around crossveins (mainly radial crossveins); CuA fork located near origin of RP; three presectoral crossveins; subcostal veinlets simple; prefork area slightly wider than posterior area. Hind wingmembrane mostly hyaline except by brown marks around crossveins, tip and rhegmal and stigmal areas; MP fork located between RP origin and first fork; subcostal veinlets simple; one presectoral crossvein. Legs ( Figs. 124a–b): All pairs of legs, femur elongate (near three times longer than coxa); tibia and femur about same size but longer than tarsi; tibial spurs reaching apex of T1; T2, T3 and T4 about same size, T1 about twice longer T2, T5 about twice longer T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with short black setae, and scattered long black setae; T5 ventrally with two rows of thick, long, black setae. Prolegsense hair as long as femur width; tibia with antennal cleaning setae ventroapically; coxa, trochanter, and femur dark brown, some specimens with a pale area on femur anterior surface; tibia dark brown with three pale rings; tarsi dark brown, except for base of T1, pale. Mesolegwith same colour pattern of proleg. Metalegwith femur, tibia and T1 slightly longer than in other legs; colour pattern similar to anterior legs, except for tibia pale but tip and ventral surface, dark brown. Abdomen: Usually entirely black or dark brown, but occasionally with irregular pale spots on sclerites posterior margins. Beset with short white setae. Male Terminalia ( Figs. 125a–c): Ectoproctposterior margin rounded in lateral view, set with elongate black setae. 9 thsterniteposterior margin with medial region acute in ventral view; set with elongate black setae. Genitaliasemi triangular in dorsal view. Gonarcusbroad and connected to Mediuncus, which is acute apically in lateral view. Paramereweakly sclerotized, thin, and elongate in lateral view. Female Terminalia ( Figs. 125d–e): Ectoproctposterior margin rounded, set with thin elongate setae and some long cavisetae ventrally. Lateral gonapophysesrounded, about same size as ectoproct, beset with elongate cavisetae. 8 thand 9 thtergitewith few thickened setae on ventral area in lateral view. 7 thsternitedistal margin with a small invagination in ventral view; covered with short black setae. Pregenital platelarge, with ventral margin curved in lateral view; curved and concave in posterior view. Posterior gonapophysesthin, elongate, covered with short black setae. 9 thtergitewith a short membranous digitiform process. Anterior gonapophysesabsent. Ventral membranegonapophyseal plates absent.   Distribution( Fig. 119). Australia: NT, QLD, WA. Widespread across northern Australia.  Adult activity period.Records for March to June; apparently a northern fall emerging species.   Biology.Unknown, larva unknown.  Name-bearing type.  Megagonoleon radialis:  Holotype(by original designation), female, ANIC, examined. Type locality: Australia, Northern Territory, Darwin(city). Fromoriginal description: “ Type— ♀. From Port Dar- win, Australia, 12th May (Dodd)”. From New (1985b; 1986): “ Holotype, ♀, Northern Territory, Darwin, 12 May (Dodd) (formerly MCZ, now ANIC) (seen)”. Condition: good; terminalia dissected.  Additional material examined( 17♂, 37♀).   AUSTRALIA:  Northern Territory: ½ mi SE of Batchelor, Camp 9,  12.iv.1966, U.V. light, N. McFarland( 5♀, 8♂, SAMA);  Creswell Downs:  9.iv.1986, J. Howard( 3♀, QMBA);  Daly Watters:  iv.1966( 1♀, SAMA);  Highway Inn:  3.iv.2008, ex woodland, at light, G. Williamsand W. Pulawski( 1♂, 4♀, AMSA);  98 kmSW of Katherine, 15.07S– 131.42E,  1.iv.1995, E.D. Edwards& M. Mat- thews ( 1♀, ANIC);  Nabarlek Dam:  14 kmSSW of Nimbuwah Rock, 12.19S– 133.19E,  2.vi.1973, M.S. Upton( 2♀, ANIC);   Queensland: Georgetown: Newcastle Range, 18.18S- 143.44E,  21.iii.1995, E.D. Edwards& M. Matthews( 2♂, ANIC);  Mica Creek:  16.iii.2001( 1♂, 6♀, QMBA);  Mornington IslandMission:  11–12.v.1963, at light, P. Aitken& N.B. Tindale( 1♂, 6♀, SAMA);  Normanton:  14.iv.1961, at light, E.N. Marks( 1♂, 1♀, ANIC, QMBA);  Riversleigh Homestead:  iv.1986( 1♀, SAMA);   Western Australia:  70 miESE of Kununurra(NT):  17.iv.1966, U.V. light, N. McFarland( 2♀, 1♂, SAMA);  Fitzroy Crossing area:  24.iii.1984, at light, K. & E. Carnaby( 2♀, AMSA);  24 kmNNW of Lissadell, 16.33S– 128.12E,  7.iv.1995, E.D. Edwards& M. Matthews( 1♀, 2♂, ANIC).   Comments.  Megagonoleon radialisis closely related to  M. stigmatus, with which it forms a species-pair clade within the larger  stigmatusspecies group ( Fig. 4). Both species appear to be restricted to the northern parts of Australia. They have dark bodies and many similarities in both the female and male terminalia, but can be easily separated based on wing and clypeal markings. ANIC Australia Darwin From From 242 243 2 2 Northern Territory holotype 1966-04-12 SAMA N. McFarland Australia Northern Territory 242 243 13 5 8 Northern Territory 1986-04-09 QMBA J. Howard Australia Creswell Downs 242 243 3 3 Northern Territory [263,679,1521,1547] 1966-04 SAMA Australia Daly Watters 242 243 1 1 Northern Territory 2008-04-03 AMSA G. Williams & W. Pulawski Australia Highway Inn 242 243 5 4 1 Northern Territory 1995-04-01 ANIC E. D. Edwards Australia -15.07 Katherine 772 131.42 242 243 1 1 Northern Territory 1973-06-02 ANIC M. S. Upton Australia -12.19 Nabarlek Dam 776 133.19 242 243 2 2 Northern Territory 1995-03-21 ANIC E. D. Edwards & M. Matthews Australia -18.18 Newcastle Range 765 143.44 Georgetown 242 243 2 2 Queensland [307,803,1665,1691] 2001-03-16 QMBA Australia Mica Creek 242 243 7 6 1 Queensland 1963-05-11 1963-05-12 1963-05-11 SAMA P. Aitken & N. B. Tindale Australia Mornington Island 242 243 7 6 1 Queensland [634,1432,1701,1727] 1961-04-14 ANIC, QMBA E. N. Marks Australia Normanton 242 243 2 1 1 Queensland [151,665,1737,1763] 1986-04 SAMA Australia Riversleigh Homestead 242 243 1 1 Queensland 1966-04-17 SAMA N. McFarland Australia 70 mi ESE of Kununurra 242 243 3 2 1 Western Australia [596,1432,1773,1799] 1984-03-24 AMSA K. & E. Carnaby Australia Fitzroy Crossing area 242 243 2 2 Western Australia [151,1302,1808,1835] 1995-04-07 ANIC E. D. Edwards & M. Matthews Australia -16.33 Lissadell 769 128.12 242 243 3 1 2 Western Australia