Myrmeleon pulchellus Rambur, 1842:408
Glenurus pulchellus (Rambur)
Glenoleon pulchellus (Rambur)
Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae)
Machado, Renato Jose Pires
Oswald, John David
Zootaxa
2020
2020-06-16
4796
1
1
322
3G98P
(Rambur, 1842)
Rambur
1842
[151,613,153,180]
Insecta
Myrmeleontidae
Glenoleon
Animalia
Neuroptera
178
179
Arthropoda
species
pulchellus
( Figs. 87–89)
Myrmeleon pulchellus Rambur, 1842:408(OD); Gerstaecker 1885a:25(cit); Gerstaecker 1885b:114(cit). Glenurus pulchellus(Rambur): Hagen 1866:405(ncb); Froggatt 1902:360(rd); Banks 1910:40(cit). Glenoleon pulchellus(Rambur): Banks 1913:224(ncb); Esben-Petersen 1915:72(key) [as “ pulchellum”]; Esben-Petersen 1923:586(cit); Handschin 1935:690(cit); Adams 1936:25(cit); Stange 1976:306(cat); New 1985b:33(key, rd, ill); Lambkin 1987:12(cit); New 1996:83(cat); Whittington 2002:382(cit); Stange 2004:105(cat); Miller & Stange 2012:14(im); Oswald 2018(cat); Machado et al. 2019(tree).
Diagnosis.Legs elongate; tibial spurs reaching T2 apex; forewing costal area uniareolate; hind wing with two broad dark bands; wings broad; male gonarcus/mediuncus complex tubular; male paramere positioned posterior to the gonarcus/mediuncus complex.
Description.Lengths: forewing: 29–34 mm; hind wing: 27–32 mm. Head ( Figs. 88a–b): Labrumbrown, set with a line of elongate setae. Clypeuspale with brown spots in some specimens; set with elongate white setae. Fronspredominantly black, except for ventrolateral corners pale; beset with white setae. Genapale to light brown. Vertexraised; in anterior view pale, with a median black line, usually interrupted centrally; in dorsal view mostly black except for some pale areas around center posteriorly; set with short black setae. Ocularsetae absent. Antennaeclubbed; elongate,> 3x length of pronotum; distance between antennae wider than scape width; all segments dark-brown to black, except by three to five subapical flagellomeres pale in some specimens; torular membrane yellow; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae. Mandiblespale to light brown, but darker at tip. Palpi, maxillary and labial pale to light brown with distal segments generally darker, apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially. Thorax ( Fig. 88b): Pronotumlonger than wide; posterior margin wider than anterior; subapical furrow present; mostly pale with a broad black longitudinal band on midline (flanked by thinner pale longitudinal bands), margins generally black; set with short black setae, and long setae at margins. Mesonotummostly black, with two rounded pale spots on prescutum, scutum with two small pale spots near wing bases and two thin pale longitudinal bands on each side, scutellum black with pale spots laterally or posterolaterally. Metanotumusually black with two pale longitudinal bands. Pterothoracic pleurablack but with few pale areas generally between sclerites; set with elongate white setae; Miller’s organ present. Wings ( Fig. 88c): Fairly broad with tip acute; anterior Banksian line present in both wings, posterior absent; veins set with short black setae, in general brown, except for some specimens with main longitudinal veins are intercalated by numerous short white regions. Male pilula axillarispresent. Forewingmembrane mostly hyaline with brown marks generally concentrated on posterior margin, tip, pterostigma, rhegmal area, and around crossveins; in some specimens the brown areas are darker; CuA fork located between RP origin and first fork; three presectoral crossveins; some subcostal veinlets forked and irregularly linked in some specimens; posterior area narrower than prefork area. Hind wingmembrane mostly hyaline except for two broad brown subapical bands; MP fork located between RP origin and first fork; subcostal veinlets simple; one presectoral crossvein, with a medial spur. Legs ( Figs. 88a–b): All pairs of legs, femur elongate (> 3.5x length of coxa); tibia slightly longer than femur but longer than tarsi; tibial spurs surpassing T2 tip; T2, T3 and T4 about same size, T1 about twice as long as T2, T5 about twice as long as T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with short black setae, and scattered long black setae; T5 ventrally with two rows of thick, long, black setae. Prolegsense hair absent; tibia with antennal cleaning setae ventroapically; colouration varies in intensity among specimens, but in general the femur dorsal surface and tip blackish, remaining areas pale, tibia blackish with two or three pale rings, T1 and base of T2 pale, remaining tarsomeres blackish. Mesolegwith same colour pattern of proleg except for a dorsal subapical yellow spot on femur. Metalegwith femur and tibia slightly longer than in other legs; femur, tibia and T5 pale except for their blackish tip; four basal tarsomeres generally pale but darker in some specimens. Abdomen: Usually entirely black or dark brown, occasionally with irregular pale spots on tergites and sternites. Male Terminalia ( Figs. 89a–d): Ectoproctposterior margin rounded in lateral view, set with elongate black setae. 9 thsternitein lateral view with posterior margin longer than anterior; in ventral view posterior margin with a shallow medial concavity; set with elongate black setae. Gonarcusand mediuncusfused forming a tubular structure. Paramerelocated posteriorly to gonarcus/mediuncus complex; long, curved and with tip acute. Female Terminalia ( Figs. 89e–g): Ectoproctposterior margin rounded set with long black setae. Lateral gonapophysesrounded, shorter than ectoproct, and set with cavisetae. 7 thsternitewith distal margin elongate and covered with long black setae in lateral view. Pregenital platelarge, squared shaped with dorsolateral margins elongate. Posterior gonapophysesbroad, elongate, set with very long black setae and, a group of cavisetae apically. 9 thtergitewith an elongate membranous digitiform process. Anterior gonapophysesa small rounded sclerotized plate, located at base of gonapophyseal plates.
Distribution( Fig. 87). Australia: ACT, NSW, QLD, SA*, VIC*. Widespread and common in eastern Australia; apparently absent from western and central Australia. Adult activity period.Records for September to April.
Biology. New (1996)stated that the larva is terrestrial and a predator. Miller and Stange (2012)reared a female from a larva collected in coarse sand at the entrance of a cave in Pilliga Nature Reserve, NSW. They did not describe the larva, but mentioned that it had a complex colour pattern and the distinctive Dendroleontinaemedial tuft. FIGURE 89. Glenoleon pulchellus: male: a) terminalia, lateral; b) 9 thsternite, ventral; c) genitalia, lateral; d) genitalia, dorsal; female: e) terminalia, lateral; f) ventral margin of 7 thsternite and pregenital plate, posterior; g) terminalia, ventral. Scale bars = 0.5 mm. Name-bearing type. Myrmeleon pulchellus: Holotype(by explicit monotypy), male, OUMNH, high resolution images examined. Type locality: Australia(no futher locality data). From original description: “D'après un individu en assez mauvais état, venant de la Nouvelle-Hollande” (According to one individual in poor condition, from New Holland). New Hollandrefers to Australia. From New (1985b): “ Australia(?Paris, not seen)”. From Tauber et al.(2019, fig. 93): “One specimen in the Hope Collection (NEUR0058, Fig. 93), a male, carries Rambur’s labels and is identified as a type. We consider it to be the M. pulchellus holotype(by explicit monotypy). The abdomen is damaged, and some legs are missing”. Until recently the holotypeof G. pulchelluswas thought to reside in the MNHN, e.g., New (1985b), Stange (2004), and Oswald (2018), but it had never been confirmed to exist there. For this study we attempted to obtain information about this specimen from the MNHN, but the specimen was not found there (Dr. André Nel, personal communication). More recently, Tauber et al.(2019)reported on multiple specimens of G. pulchellusfound in the OUMNH. One of these specimens bears labels in Rambur’s handwriting and a post-Rambur label that identifies it as a “Type” that was accessioned into the OUMNH as part of the Marchal Collection. Numerous other Rambur type specimens of neuropterous insects (in the Linnean sense) are known to have entered the OUMNH by way of the Marchal Collection, so the provenance of this specimen is consistent with its identification as the original holotypeof G. pulchellus. Condition: good, antennae broken, right foreleg missing and abdomen damaged. Additional material examined( 142♂, 211♀, 8?). AUSTRALIA: 1917 ( 1♀, 1♂, SAMA); ACT: Black Mountain: 10.ii.1952, L.J. Chinnick( 1♀, ANIC); ii.1967( 1♂, ANIC); 16.i.1950( 1♀, ANIC); Canberra: 11.ii.1967, E. Paramonov( 1♀, ANIC); 23.xi.1929( 1♂, ANIC); 5.ii.1951( 1♂, ANIC); 2.ii.1960( 2♀, 2♂, ANIC); 5.ii.1960( 4♀, 1♂, ANIC, 1♂, FSCA); 8.ii.1960( 2♀, ANIC); 16.ii.1960( 1♂, ANIC); 20.ii.1960, E.F. Riek( 1♀, AMSA); 7.iii.1960( 2♀, 1♂, ANIC); 28.xi.1960, K.G. Sewnson( 1♀, OSU); 12.xii.1960, E.F. Riek( 1♂, TAMU); 15.xii.1960( 1♀, 2♂, ANIC); 27.xii.1960( 2♀, ANIC); 29.xii.1960( 2♀, ANIC); 9.i.1961( 1♀, 1♂, ANIC); 12.i.1961( 1♂, ANIC); 23.iv.1961( 1♀, ANIC); 1.ii.63 ( 1♂, AMSA); 14.ii.1963( 1♀, ANIC); Cotter River Reserve: 26.xi.1977, G. Daniels( 1♀, AMSA); Mount Majura: 27.xi.1977, G. Daniels( 1♂, 1♀, AMSA); New South Wales: no label ( 1♂, ANIC); 22.xii.12 ( 1♂, ANIC); Armidale: 20.xii.1954( 1♀, 1♂, ANIC); 23.xi.1959( 1♂, ANIC); 11.xii.1959( 1♀, ANIC); xii.1960( 2♀, ANIC); Asquith: 21.ii.66, A.B. Rose( 1♂, AMSA); Ballina: x.22, A.N. Burns( 1♀, AMSA); Bank- stown: near Sydney, 16.xi.1980, B.J. Day( 1♀, AMSA); Banyabba Natural Reserve: 20.i.1971( 1♀, ANIC); Bay- view: 29.x.1970, L.C. Haines( 1♀, AMSA); Belmont: 4.i.1947( 2♂, 1♀, MVM); 30.xii.1946( 1♂, MVM); Bend- along: 24.i.1970, G. Daniels( 1♂, AMSA); Berowra: Galston Gorgenear Hawkesburry R., ii.1927, T.G. Campbell( 1♀, AMSA); 6 kmNE of Bilpinnear Nurrajong, 2.ii.1980 N.W. Rodd( 1♂, AMSA); Blue Mountains: Clarence, 8.i.1983, N.W. Rodd( 1♂, AMSA); Brindabella Range: iii.1984( 1♂, ANIC); Loqnet ValleySchool, 6.ii.1984, at light, L.C. Haines( 1♀, AMSA); Mount Tomah, 9.i.1984, N.W. Rodd( 1♀, AMSA); Braidwood: 14.i.1934, Fuller( 1♀, 1♂, ANIC); Bundeena: 12.xi.1960( 1♀, FSCA); Cabramatta: 10.i.1961( 1♂, FSCA); 1.5miN of Colo River, 23.x.65, G.L. Bush( 1♀, TAMU); Como: 7.i.1911, Waterhousecoll. ( 1♀, ANIC); 18.i.1933( 1♀, MVM); 5.ii.1922( 1♀, MVM); Conjola: 26.xii.1950( 1♂, MVM); Coonabarabran: 2.xii.1974( 1♂, ANIC); Cuttler’s Pass: William’s river, 10.i.1943, A. Musgrave( 1♀, AMSA); Deriah Aboriginal Area: 23 kmE of Narrabri, - 30.345 S– 150.014 E, 10.xi.2009, Ooline, light, D.R. Britton, J. Recsei( 1♀, AMSA); Ebenezer: xii.1996( 1♀, ANIC); Hazelbrook, 1984, M. Dingley( 2♂, 1♀, AMSA); i.1985, light ( 1♂, AMSA); Elanora Heights: xi.1986, T. Robinson( 1♀, AMSA); Elizabeth Bay: 29.i.1940, K.C. McKeown( 1♀, AMSA); Engadine: near Sydney, 23.xii.1976, R. Eastwood( 1♀, AMSA); Fletcher: 29.xii.1960( 1♂, ANIC); Fraser Park: 2.i.1947( 1♀, MVM); Glenbrook: 15.1.1993, M.S. & B.J. Moulds( 1♂, 1♀, AMSA); 27.xii.1964( 1♀, ANIC); Gosford: ( 1♂, ANIC); Greta: 1951 ( 1♂, FSCA); Groose Vale: 29.x.1988, N.W. Rodd( 1♂, AMSA); Gundamain National Park: 14.ii.1926, Mackerras( 1♂, FSCA); Helensburgh: 23.i.1979, R. Eastwood( 1♀, AMSA); Hornsby: 3.i.1916, Tillyard( 1♂, ANIC); 9.3 kmS of Ilford, 29.xii.1977, G. Daniels( 1♀, AMSA); Killara: 22.xii.1934, M.F. Day( 1♀, ANIC); Killarney Gap: Kaputar National Park, - 30.141 S– 150.066 E, 14.xi.2009, Callitris& sclerophyll scrub / rocky ridge, MV Lamp, D.R. Britton, J. Recsei( 1♀, AMSA); 6 kmNE of Kyogle, Murwillumbah Road, 9.i, B.J. Day( 1♂, AMSA); Liverpool: Heathcote Road., 33 o56’ S– 150 o56’ E, 13.xii.1998, light, B.J. & C.J.R. Day( 1♂, AMSA); Loftus: 27.xii.1978, J. Olive( 1♂, AMSA); Mann River: 5 kmupstream from Old Glen Innes, Grafton Roadcrossing, 27.xi.1977, M.S. & B.J. Moulds( 1♀, AMSA); Mittagong( 1♂, SAMA); Moonbi: 7.ii.1968, D.H. Collens( 1♀, ANIC); Mooney Mooney Creek: near Gosforo, 13.i.1977, D.K. McAlpine( 1♀, AMSA); Mount Kaputar National Park: 15.i.1978, G. Daniels( 1♀, AMSA); Mount Victoria: 4.xii.1946( 1♀, MVM); Nambucca: 26.x.1949( 1♀, MVM); Narrabeen: 18.ix.1934( 1♀, ANIC); Narrabri: 17.xii.1934, M.F. Day( 1♀, ANIC); xi.1997( 1♀, ANIC); Nepean River: x.1931, A. Musgrave( 1♂, AMSA); New- castle: Sugarloaf Mount, 2.i.1947( 1♂, MVM); Newport: 2.xii.57, G. Dolezal( 1♀, AMSA); Pilliga Nature Reserve: 34 kmN of Coonabarabran, 13.x.2005, L.A. Stange( 1♀, FSCA); Royal National Park: Goondera Ridge, 3.xii.1977, G. & A. Daniels( 1♀, AMSA); Singleton: 23.iii.1957, E.F. Riek( 1♂, ANIC); Sutton: 19.i.1979( 1♂, QMBA); Syd- ney: C. Gibbons( 1♀, AMSA); 28.xii.1961, D.H. Collens( 1♂, ANIC); xii.1946( 1♀, MVM); ( 1♀, SAMA); Waita- ra, 23.i.1983, M.S. & B.J. Moulds( 1♂, AMSA); Tomerong: Nowra district, 3miN, 7.ii.1970, D.K. McAlpine( 1♀, AMSA); Tuggerah: xii.1904( 1♀, SAMA); Ulan: xi.1982( 1♂, ANIC); Wahroonga: 12.xii.1970, A.B. Rose( 1♀, AMSA); H.J. Carter( 1♀, AMSA); Wapengo: 12.i.1978( 1♀, MVM); Warialda: Adam’s Scrub 14.6 kmS, - 29.667 S– 150.567 E, 18.xi.2009, Callitris/sclerophyll forest, MV lamp, D.R. Britton, J. Recsei( 1♂, AMSA); Wastepoint: 8.3 kmN of Jindabyne, KosciuszkoN.P., - 36.347 S– 148.606 E, 8.xii.2007, MV lamp, D.R. Britton, D.J. Smith, P. Hinton( 1♂, AMSA); Watalgan Forrest: 24.xi.1973( 1♀, ANIC); Wedderburr: 3 kmE, 34 o08’ S– 150 o49’ E, 13.xii.2005, MV light, D. Britton( 1♂, AMSA); Wee Jasper: 25.xii.56, E. shipp ( 1♂, AMSA); Wellington: 28.ix.1957, E.F. Riek( 2♂, ANIC); West Head: PiHwater, 2.i.1985, L.C. Haines( 1♂, AMSA); Whiskers: xii.1992( 1♂, ANIC); Queensland: ( 1♀, ZMHB); ( 1♀, SAMA); Archer Rivercrossing: 13 o25’S– 142 o56’E, iv.1989, mv lamp, G. & A. Daniels(6?, QMBA); Barankula State Forrest: 26 o26’07’’ S– 150 o49’31’’ E, 3–7.iv.2000, malaise, M. Mathieson( 1♀, TAMU); Bardon: x.1943( 1♀, 1♂, SAMA); Barrine: 10.x.1967( 2♂, ANIC); Blackdown Tableland: Expedi- tion Range, 17.i.1987, M.S. & B.J. Moulds( 1♂, AMSA); Bluff Range: 16.xii.1970( 1♀, 1♂, ANIC); 9.i.1971( 2♀, 1♂, ANIC); Biggenden: 22–23.xi.1977( 12♀, 9♂, ANIC); 1.i.1972( 2♀, ANIC); 2.xii.1974( 1♂, ANIC); Biloela: 17.i.1947( 1♀, ANIC); Bin Bin Range: 4.xii.1974( 1♀, ANIC); 12.i.1972( 1♀, 1♂, ANIC); Bingera: 22.xi.1971( 1♀, ANIC); Bon Accord Falls: Montville, Blackall Ranges, 22.ix.55, R. Dobson( 1♀, AMSA); Boolaboonda Range: 10.xi.1973( 1♀, 1♂, ANIC); Boompa: 4.xii.1980, H. Franca( 1♂, ANIC); Bowen: 1.x.1950, E.F. Riek( 1♀, ANIC); Brisbane: ( 6♂, 24♀, QMBA); i.1888( 2♀, SAMA); xi.1965( 2♀, ANIC); 20.x.1900( 1♀, ANIC); 1903 ( 1♀, ZMUC); x.1911( 1♀, SAMA); 20.ii.27, L. Franzen( 2♂, ZMUC); 19.ix.1927( 1♀, QMBA); 30.xii.1927( 1♀, MVM); 19.x.1938( 1♀, ANIC); 9.i.1960( 1♂, 1♀, MVM); 10.i.1960( 1♀, MVM); 24.xii.1955( 1♀, MVM); 27.xii.1959( 1♀, MVM); 2.xii.1922( 2♀, MVM); 29.x.1936( 1♀, MVM); 4.xi.1936( 1♂, MVM); 26.xi.1956( 1♂, MVM); x.1910( 1♂, 2♀, MVM); Moggill Farm, 23.i.1961( 1♂, FSCA); Bucasia: i.2003( 1♂, ANIC); Bundaberg: 19.xi.1978( 2♀, 3♂, ANIC); 22.xii.1979( 1♀, 2♂, ANIC); Bunya Mountains: 17.i.1955( 1♂, MVM); Burleigh Heads: 3.i.1956( 1♀, MVM); xi.1942( 1♂, MVM); xii.1942( 1♂, MVM); 10.x.1956( 1♀, MVM); Byfield State Forrest: x.1924, A. Musgrave( 1♂, AMSA); 1.i.1976, G. Daniels( 1♀, AMSA); Carnarvon National Park: 7.xii.1938, N. Geary( 2♀, AMSA); 8.xii.1938( 1♂, AMSA); 12.xii.1938( 1♂, 1♀, AMSA); 5.i.1939( 1♀, AMSA); 8.i.1939( 1♂, AMSA); ii.1944( 1♀, AMSA); 92 air km NNW of Injune, 25.06695°S 148.24611°E± 30m, R Machado, MV light, 25.xii.2015, AustRM#10 ( 2♀, TAMU); Charleville: 27.xii.1925( 1♀, QMBA); Cordalba National Forrest: 30 kmS of Bunda- berg, 29.xi.1979, H. frauca ( 1♀, ANIC); Degilbo: 9.xii.1978( 2♂, ANIC); Electra: 8.ii.1977( 1♂, ANIC); Fletcher: 12.i.1967, E. Sutton( 1♂, ANIC); Forty Mile Scrub: 4.x.1989( 1♂, ANIC); Hayman Island: 24.x.1950, R. Dobson( 3♀, 1♂, ANIC); Herberton: 6.xii.1968, E. Britton& S. Misko( 1♂, ANIC); x.1990( 1♀, ANIC); 55 kmNNE of Injune, 23.xi.1986, M.S. & B.J. Moulds( 1♂, AMSA); Kinbombi falls: Goomeri, 19.xii.1976, M.S. & B.J. Moulds( 1♀, AMSA); Kingston: xii.1981( 1♂, QMBA); Kuranda: ( 1♀, SAMA); 19.xi.1951( 1♀, MVM); 5.x.1949( 1♀, MVM); Mackay: 1905 ( 1♀, 1♂, ANIC); Milmerran: 14.xi.1945( 1♂, MVM); Montville: Bon Accord Falls, 20.ix.1955( 1♂, 1♀, MVM); Mount Cootha: x.1942( 1♀, SAMA); 16.xii.1973( 2♀, QMBA); Mount Garmet: i.1961( 1♂, ANIC); Mount Hay: 24.i.2009( 1♀, QMBA); Mount Moffat National Park: Mahogany Forest, 11–12.xii.1987, M.T. Yeates( 1♂, 2♀, QMBA); Mount Tamborine: 30.iii.1950( 1♂, MVM); 26.xii.1950( 1♀, MVM); MountTin- beerwah: 1.i.1979( 1♀, QMBA); Murphy’s Creek: 29.xi.1992( 1♀, QMBA); Noosa Heads: 1.x.1956( 1♀, MVM); Ravenshoe: xi.1998( 1♀, ANIC); Stanthorpe: 30.i.1927( 1♀, QMBA); 29.xii.1977( 2♂, QMBA; 1♂, ANIC); 9.ii.1981(1?, QMBA); Stradbroke Island: 21.ix.1954( 1♀, MVM); Tamborine: ( 1♂, QMBA); Toowoomba: 11.ii.1960( 1♂, ANIC); Townsville: 12.xii.1908( 1♀, MVM); Wacol: 16.x.1952( 1♂, QMBA); West Burleigh: 1960, A.T. Bandsoma( 1♂, AMSA); Woodridge: 2.i.1968( 1♂, MVM); South Australia: Aldgate: 19.iii.19309( 1♀, MVM); Barossa: ( 1♀, SAMA); Blackwood: ii.1965( 2♀, SAMA); 20.i.1969( 1♀, MVM; 2♀, SAMA); xii.1975( 1♀, SAMA); Bridgewater: xii.1967( 1♀, SAMA); Cara Pook: i.1973( 1♂, SAMA); Heywood Park: xii.1958( 1♂, SAMA); Highbury: i.1969( 1♂, SAMA); Kangaroo Creek Reserve: i.1971( 1♂, SAMA); “Kurlge” Blackwood: xi.1957( 1♀, SAMA); xii.1957( 1♂, SAMA); i.1958( 2♀, SAMA); ii.1958( 1♀, SAMA); Mount Lofty Range: ii.1965( 1♀, SAMA); xii.1973( 1♂, SAMA); Nairne: i.1988( 1♂, SAMA); Locationnot recognized: Saunder’s Coll., iii.68 ( 1♀, BMNH); Nolabel ( 2♀, 3♂, ANIC); Victoria: Bendigo: 5.i.1930( 1♀, MVM); Hurstbridge: 18.ii.1947( 1♂, MVM); Monbulk: 27.i.1929( 1♀, MVM); Nowa Nowa: 21.ii.1947( 1♂, MVM); Moe: 24.xii.1944 ( 1♂, MVM); Oakleigh: ( 1♂, MVM); Research: ii.1983- ( 1♀, ANIC); Warrandyte: i.1988( 1♀, ANIC); Wartook: xii.1935( 1♀, SAMA).
Comments. Glenoleon pulchellusis a common species in eastern Australia. Its color intensity varies considerably among specimens; some are almost completely dark, while others have many pale areas. Despite these coloration differences the species is easily recognized by the two dark bands in the apical half of the hind wing; no other species in the Periclystusgenus group has this character. The long legs and pronotum, and broad wings, of this species support its close relationship with the several other species retained here in Glenoleon, and is further demonstrated by both molecular ( Machado et al. 2019) and other morphological data ( Fig. 4).
1952-02-10
SAMA
Australia
Black Mountain
181
182
2
1
1
[339,883,765,791]
1967-02
ANIC
Chinnick
Australia
181
182
2
1
1
[896,1151,764,791]
1950-01-16
ANIC
Chinnick
Australia
true
181
182
1
1
1929-11-23
ANIC
Paramonov
Australia
181
182
2
1
1
[714,960,800,827]
1951-02-05
ANIC
Paramonov
Australia
true
181
182
1
1
[971,1266,801,827]
1960-02-02
ANIC
Australia
181
182
4
2
2
1960-02-05
ANIC, FSCA
Australia
181
182
6
4
2
[418,660,837,863]
1960-02-08
ANIC
Australia
181
182
2
2
[670,926,837,863]
1960-02-16
ANIC
Australia
181
182
1
1
1960-03-07
AMSA, ANIC
Riek
Australia
181
182
4
3
1
[482,918,873,899]
1960-12-12
OSU
Sewnson
Australia
181
182
1
1
1960-12-15
TAMU, ANIC
Riek
Australia
181
182
4
1
3
[249,533,909,935]
1960-12-27
ANIC
Riek
Australia
true
181
182
2
2
[550,834,909,935]
1960-12-29
ANIC
Australia
181
182
2
2
[851,1153,909,935]
1961-01-09
ANIC
Australia
181
182
2
1
1
[1170,1432,909,935]
1961-01-12
ANIC
Australia
181
182
1
1
[151,414,945,971]
1961-04-23
ANIC
Australia
181
182
1
1
[425,922,945,971]
1963-02-14
AMSA, ANIC
Australia
181
182
2
1
1
[933,1305,945,972]
1977-11-26
Australia
Cotter River Reserve
181
182
1
1977-11-27
AMSA
Daniels
Australia
Mount Majura
181
182
1
1
1954-12-20
AMSA, ANIC
Daniels
Australia
Armidale
181
182
6
2
4
New South Wales
[875,1143,1016,1043]
1959-11-23
ANIC
Daniels
Australia
true
Armidale
181
182
1
1
New South Wales
[1156,1432,1016,1043]
1959-12-11
ANIC
Australia
181
182
1
1
[151,393,1053,1079]
1960-12
ANIC
Australia
181
182
2
2
1980-11-16
AMSA
Asquith & Rose & Burns
Australia
Sydney
Ballina
181
182
2
1
1
[524,1369,1089,1115]
1971-01-20
AMSA, ANIC
Day
Australia
Banyabba Natural Reserve
181
182
2
2
[349,1064,1125,1151]
1947-01-04
AMSA, MVM
Haines
Australia
Belmont
181
182
4
2
2
[1077,1355,1125,1151]
1946-12-30
MVM
Haines
Australia
true
Belmont
181
182
1
1
[349,1259,1161,1187]
1927-02
AMSA
G. Daniels & Hawkesburry R.
Australia
Galston Gorge
Berowra
181
182
1
1
1980-02-02
AMSA
T. G. Campbell
Australia
Nurrajong
181
182
1
1
1983-01-08
AMSA
N. W. Rodd
Australia
Clarence
Blue Mountains
181
182
1
1
[259,1025,1233,1259]
1984-03
AMSA, ANIC
Rodd
Australia
Brindabella Range
181
182
2
2
[1038,1403,1233,1259]
1984-02-06
Australia
Loqnet Valley
181
182
1
1984-01-09
AMSA
Haines
Australia
Mount Tomah
181
182
1
1
[808,1356,1269,1295]
1934-01-14
AMSA
N. W. Rodd
Australia
Braidwood
181
182
1
1
1960-11-12
ANIC, FSCA
Fuller
Australia
Bundeena
181
182
3
2
1
[762,1162,1305,1331]
1961-01-10
FSCA
Cabramatta
Australia
181
182
1
1
1911-01-07
TAMU, ANIC
G. L. Bush & Waterhouse
Australia
Como
181
182
2
2
[1070,1325,1341,1368]
1933-01-18
MVM
Australia
181
182
1
1
1922-02-05
MVM
Australia
181
182
1
1
[305,687,1376,1404]
1950-12-26
MVM
Conjola
Australia
181
182
1
1
[699,1145,1377,1403]
1974-12-02
ANIC
Australia
Coonabarabran
181
182
1
1
1943-01-10
Cuttler's Pass
Australia
William's river
181
182
1
2009-11-10
AMSA
Musgrave
Australia
-30.345
Deriah Aboriginal Area
72
150.014
181
182
1
1
[285,1218,1449,1476]
1996-12
AMSA, ANIC
Ooline & D. R. Britton & Recsei
Australia
Ebenezer
181
182
2
2
1985-01
AMSA
Hazelbrook & Dingley
Australia
181
182
3
1
2
[599,1116,1485,1511]
1986-11
AMSA
Australia
Elanora Heights
181
182
1
1
1940-01-29
AMSA
Robinson
Australia
Elizabeth Bay
181
182
1
1
[451,1217,1521,1548]
1976-12-23
AMSA
K. C. McKeown
Australia
Sydney
Engadine
181
182
1
1
1960-12-29
AMSA, ANIC
Eastwood
Australia
Fletcher
181
182
2
1
1
[647,1031,1557,1584]
1947-01-02
MVM
Fraser Park
Australia
181
182
1
1
1964-12-27
AMSA, ANIC
Moulds
Australia
181
182
3
2
1
1988-10-29
ANIC, FSCA
Gosford
Australia
Groose Vale
Greta
181
182
2
2
[277,996,1629,1656]
1926-02-14
AMSA
Rodd
Australia
Gundamain National Park
181
182
1
1
1979-01-23
FSCA
Mackerras
Australia
Helensburgh
181
182
1
1
[272,790,1665,1692]
1916-01-03
AMSA
R. Eastwood
Australia
Hornsby
181
182
1
1
[803,1397,1665,1691]
1977-12-29
ANIC
Tillyard
Australia
9.3 km S of Ilford
181
182
1
1
1934-12-22
AMSA
G. Daniels
Australia
Killara
181
182
1
1
2009-11-14
ANIC
M. F. Day
Australia
-30.141
Kaputar National Park
73
150.066
Killarney Gap
181
182
1
1
1998-12-13
AMSA
Lamp & D. R. Britton & J. Recsei & B. J. Day
Australia
Murwillumbah Road
-33.933334
Heathcote Road.
1202
150.93333
Liverpool
181
182
2
1
1
[446,1102,1809,1835]
1978-12-27
AMSA
B. J. & C. J. R. Day
Australia
Loftus
181
182
1
1
1977-11-27
AMSA
Olive
Australia
Grafton Road
Mann River
181
182
1
1
1968-02-07
AMSA, SAMA
Moulds
Australia
Moonbi
Mittagong
181
182
2
1
1
1977-01-13
ANIC
D. H. Collens
Australia
Gosforo
Mooney Mooney Creek
181
182
1
1
[270,1064,1916,1943]
1978-01-15
AMSA
McAlpine
Australia
Mount Kaputar National Park
181
182
1
1
1946-12-04
AMSA, MVM
G. Daniels
Australia
Mount
181
182
2
2
Victoria
[526,919,1953,1979]
1949-10-26
MVM
Nambucca
Australia
181
182
1
1
[929,1323,1953,1979]
1934-09-18
ANIC
Narrabeen
Australia
181
182
1
1
1934-12-17
Australia
Narrabri
181
182
1
[293,787,1989,2016]
1997-11
ANIC
Day
Australia
181
182
2
2
[798,1047,1989,2016]
1931-10
Australia
Nepean River
181
182
1
1947-01-02
AMSA, MVM
Musgrave
Australia
182
183
Sugarloaf Mount
181
182
2
2
2005-10-13
AMSA
Newport & Dolezal
Australia
Pilliga Nature Reserve
182
183
1
1
[590,1432,189,215]
1977-12-03
FSCA
L. A. Stange
Australia
Goondera Ridge
Royal National Park
182
183
1
1
[151,746,224,251]
1957-03-23
AMSA
G. & A. Daniels
Australia
Singleton
182
183
1
1
[757,1372,225,251]
1979-01-19
ANIC, QMBA
Riek
Australia
Sutton
182
183
2
2
1961-12-28
AMSA
Gibbons
Australia
182
183
1
1
[644,1352,261,287]
1946-12
ANIC, MVM, SAMA
Collens
Australia
182
183
3
2
1
[307,1192,297,323]
1970-02-07
AMSA
M. S. & B. J. Moulds
Australia
Nowra district
Tomerong
182
183
1
1
1904-12
AMSA, SAMA
McAlpine
Australia
Tuggerah
182
183
2
2
[651,957,333,359]
1982-11
ANIC
Ulan
Australia
182
183
1
1
1978-01-12
AMSA, MVM
A. B. Rose & H. J. Carter
Australia
Wapengo
182
183
3
3
2007-12-08
AMSA
D. R. Britton & J. Recsei
Australia
-36.347
Kosciuszko
70
148.606
Waste
182
183
1
1
1973-11-24
AMSA, ANIC
D. R. Britton & Smith & Hinton
Australia
Watalgan Forrest
182
183
2
1
1
[290,1432,512,539]
1957-09-28
AMSA
Britton
Australia
Wellington
Wee Jasper
182
183
2
2
[151,760,548,575]
1985-01-02
ANIC
E. F. Riek
Australia
PiHwater
West Head
182
183
2
2
[771,1432,549,575]
1992-12
AMSA, ANIC
Haines
Australia
Whiskers
182
183
2
2
[151,1216,584,611]
1989-04
ZMHB, SAMA
Australia
-13.416667
Archer River
1291
142.93333
182
183
2
2
Queensland
2000-04-03
2000-04-07
2000-04-03
QMBA
G. & A. Daniels
Australia
-26.435276
Barankula State Forrest
20
150.82527
182
183
1
1943-10
TAMU, SAMA
Mathieson
Australia
Bardon
182
183
3
2
1
[715,1074,657,683]
1967-10-10
ANIC
Barrine
Australia
182
183
2
2
1987-01-17
Blackdown Tableland
Australia
Range
182
183
1
[407,1275,693,719]
1970-12-16
AMSA, ANIC
Moulds
Australia
Bluff Range
182
183
3
1
2
1971-01-09
ANIC
Moulds
Australia
true
Bluff Range
182
183
3
2
1
[294,807,729,755]
1977-11-22
1977-11-23
1977-11-22
ANIC
Biggenden
Australia
182
183
21
12
9
[820,1060,729,755]
1972-01-01
1977-11-23
1977-11-22
ANIC
Biggenden
Australia
true
182
183
2
2
[1073,1335,729,755]
1974-12-02
ANIC
Australia
182
183
1
1
1947-01-17
ANIC
Biloela
Australia
182
183
1
1
[409,842,765,791]
1974-12-04
ANIC
Bin Bin Range
Australia
182
183
1
1
[852,1148,765,791]
1972-01-12
ANIC
Bin Bin Range
Australia
true
182
183
2
1
1
1971-11-22
ANIC
Bingera
Australia
182
183
1
1
1973-11-10
AMSA, ANIC
Montville & Blackall Ranges & Dobson
Australia
Boolaboonda Range
Bon Accord Falls
182
183
3
2
1
[712,1172,836,863]
1950-10-01
ANIC
H. Franca
Australia
Bowen
182
183
1
1
1888-01
ANIC, QMBA, SAMA
Riek & Brisbane
Australia
182
183
33
27
6
[723,949,872,899]
1965-11
ANIC
Riek & Brisbane
Australia
true
182
183
2
2
[958,1211,873,899]
1900-10-20
ANIC
Australia
182
183
1
1
1911-10
ZMUC, SAMA
Australia
182
183
2
2
[415,1128,909,935]
1927-09-19
ZMUC, QMBA
Franzen
Australia
182
183
3
1
2
[1145,1432,909,935]
1927-12-30
MVM
Franzen
Australia
true
182
183
1
1
[151,414,945,971]
1938-10-19
ANIC
Australia
182
183
1
1
[428,725,945,971]
1960-01-09
MVM
Australia
182
183
2
1
1
[739,999,945,971]
1960-01-10
MVM
Australia
182
183
1
1
[1013,1295,944,971]
1955-12-24
MVM
Australia
182
183
1
1
1959-12-27
MVM
Australia
182
183
1
1
[311,581,981,1008]
1922-12-02
MVM
Australia
182
183
2
2
[596,864,981,1008]
1936-10-29
MVM
Australia
182
183
1
1
[879,1141,981,1008]
1936-11-04
MVM
Australia
182
183
1
1
[1156,1432,980,1008]
1956-11-26
MVM
Australia
182
183
1
1
[151,420,1017,1044]
1910-10
MVM
Australia
182
183
3
2
1
[430,848,1017,1044]
1961-01-23
FSCA
Moggill Farm
Australia
182
183
1
1
[857,1171,1017,1043]
2003-01
ANIC
Bucasia
Australia
182
183
1
1
1978-11-19
ANIC
Bundaberg
Australia
182
183
5
2
3
[348,668,1053,1079]
1979-12-22
ANIC
Bundaberg
Australia
true
182
183
3
1
2
[679,1145,1052,1080]
1955-01-17
MVM
Australia
Bunya Mountains
182
183
1
1
1956-01-03
MVM
Burleigh Heads
Australia
182
183
1
1
[304,537,1089,1116]
1942-11
MVM
Burleigh Heads
Australia
true
182
183
1
1
[548,789,1089,1116]
1942-12
MVM
Australia
182
183
1
1
[800,1060,1088,1116]
1956-10-10
MVM
Australia
182
183
1
1
1976-01-01
AMSA
Musgrave
Australia
182
183
1
1
[545,1258,1125,1151]
1938-12-07
AMSA
Daniels
Australia
Carnarvon National Park
182
183
1
1
1938-12-08
AMSA
Geary
Australia
182
183
3
2
1
[550,893,1161,1187]
1938-12-12
AMSA
Geary
Australia
true
182
183
2
1
1
[907,1163,1160,1187]
1939-01-05
AMSA
Australia
182
183
1
1
[1177,1432,1161,1187]
1939-01-08
AMSA
Australia
182
183
1
1
[151,384,1197,1223]
1944-02
AMSA
Australia
182
183
1
1
[393,1431,1196,1224]
2015-12-25
Australia
30
-25.06695
Machado
1
148.24611
182
183
1
[151,895,1232,1259]
1925-12-27
TAMU, QMBA
Australia
Charleville
182
183
3
3
[349,975,1269,1295]
1978-12-09
ANIC
Australia
Degilbo
182
183
3
1
2
[986,1325,1269,1295]
1977-02-08
ANIC
Electra
Australia
182
183
1
1
[272,990,1305,1331]
1989-10-04
ANIC
Sutton
Australia
Forty Mile Scrub
182
183
2
2
[1002,1303,1304,1331]
1950-10-24
Australia
Hayman Island
182
183
1
1968-12-06
ANIC
R. Dobson
Australia
Herberton
182
183
4
3
1
[613,1253,1341,1367]
1990-10
ANIC
Britton & Misko
Australia
182
183
2
1
1
1986-11-23
Australia
Injune
182
183
1
[371,1196,1377,1403]
1976-12-19
AMSA
M. S. & B. J. Moulds
Australia
Goomeri
Kinbombi falls
182
183
1
1
1981-12
AMSA, QMBA
Moulds
Australia
Kingston
182
183
2
1
1
[710,1264,1412,1439]
1951-11-19
SAMA, MVM
Kuranda
Australia
182
183
2
2
1949-10-05
MVM
Kuranda
Australia
true
182
183
1
1
[252,1071,1448,1475]
1945-11-14
ANIC, MVM
Mackay
Australia
Milmerran
182
183
3
1
2
1955-09-20
MVM
Montville
Australia
Bon Accord Falls
182
183
2
1
1
[474,882,1485,1511]
1942-10
SAMA
Australia
Mount Cootha
182
183
1
1
[891,1178,1485,1511]
1973-12-16
QMBA
Australia
true
Mount Cootha
182
183
2
2
1961-01
ANIC
Australia
Mount Garmet
182
183
1
1
[299,708,1521,1547]
2009-01-24
QMBA
Australia
Mount Hay
182
183
1
1
[718,1431,1521,1547]
1987-12-11
1987-12-12
1987-12-11
Australia
Mahogany Forest
Mount Moffat National Park
182
183
1
[151,1004,1556,1584]
1950-03-30
QMBA, MVM
Yeates
Australia
Mount Tamborine
182
183
4
2
2
[1016,1294,1556,1584]
1950-12-26
MVM
Yeates
Australia
true
Mount Tamborine
182
183
1
1
1979-01-01
QMBA
Mount
Australia
182
183
1
1
[533,1012,1593,1619]
1992-11-29
QMBA
Australia
Murphy's Creek
182
183
1
1
[1023,1432,1592,1619]
1956-10-01
MVM
Noosa Heads
Australia
182
183
1
1
[151,535,1629,1655]
1998-11
ANIC
Ravenshoe
Australia
182
183
1
1
[552,973,1629,1655]
1927-01-30
QMBA
Stanthorpe
Australia
182
183
1
1
[990,1432,1629,1655]
1977-12-29
QMBA, ANIC
Australia
182
183
3
3
[151,419,1665,1691]
1981-02-09
QMBA
Australia
182
183
1
[437,948,1664,1691]
1954-09-21
MVM
Australia
Stradbroke Island
182
183
1
1
1960-02-11
QMBA, ANIC
Tamborine
Australia
Toowoomba
182
183
2
2
[416,826,1701,1727]
1908-12-12
MVM
Townsville
Australia
182
183
1
1
[836,1192,1700,1727]
1952-10-16
QMBA
Wacol
Australia
182
183
1
1
1968-01-02
AMSA, MVM
Bandsoma
Australia
Woodridge
West Burleigh
182
183
2
2
1965-02
MVM, SAMA
Aldgate & Barossa
Australia
Blackwood
182
183
4
4
South Australia
[922,1330,1773,1799]
1969-01-20
MVM, SAMA
Australia
182
183
3
3
1975-12
SAMA
Australia
182
183
1
1
[314,719,1809,1835]
1967-12
SAMA
Bridgewater
Australia
182
183
1
1
[730,1092,1809,1835]
1973-01
SAMA
Cara Pook
Australia
182
183
1
1
1958-12
SAMA
Heywood Park
Australia
182
183
1
1
[259,620,1845,1871]
1969-01
SAMA
Highbury
Australia
182
183
1
1
[634,1174,1845,1871]
1971-01
SAMA
Australia
Kangaroo Creek Reserve
182
183
1
1
1957-11
SAMA
Blackwood
Australia
182
183
1
1
[416,675,1880,1907]
1957-12
SAMA
Blackwood
Australia
true
182
183
1
1
[689,926,1880,1907]
1958-01
SAMA
Australia
182
183
2
2
[940,1185,1880,1907]
1958-02
SAMA
Australia
182
183
1
1
1965-02
SAMA
Australia
Mount Lofty Range
182
183
1
1
[410,668,1917,1943]
1973-12
SAMA
Australia
true
Mount Lofty Range
182
183
1
1
[683,1014,1917,1943]
1988-01
SAMA
Nairne
Australia
182
183
1
1
1930-01-05
BMNH, ANIC, MVM
Location & Saunder's Coll. & No
Australia
Bendigo
182
183
7
4
3
Victoria
1947-02-18
MVM
Hurstbridge
Australia
182
183
1
1
[426,799,1989,2015]
1929-01-27
MVM
Monbulk
Australia
182
183
1
1
[811,1230,1989,2015]
1947-02-21
MVM
Nowa Nowa
Australia
182
183
1
1
[308,571,153,179]
MVM
183
184
1
1
[584,941,153,179]
1983-02
ANIC
183
184
1
1
[953,1320,153,179]
1988-01
ANIC
183
184
1
1
1935-12
SAMA
183
184
1
1