Myrmeleon pulchellus Rambur, 1842:408 Glenurus pulchellus (Rambur) Glenoleon pulchellus (Rambur) Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae) Machado, Renato Jose Pires Oswald, John David Zootaxa 2020 2020-06-16 4796 1 1 322 3G98P (Rambur, 1842) Rambur 1842 [151,613,153,180] Insecta Myrmeleontidae Glenoleon Animalia Neuroptera 178 179 Arthropoda species pulchellus  ( Figs. 87–89)    Myrmeleon pulchellus Rambur, 1842:408(OD); Gerstaecker 1885a:25(cit); Gerstaecker 1885b:114(cit).   Glenurus pulchellus(Rambur): Hagen 1866:405(ncb); Froggatt 1902:360(rd); Banks 1910:40(cit).   Glenoleon pulchellus(Rambur): Banks 1913:224(ncb); Esben-Petersen 1915:72(key) [as “ pulchellum”]; Esben-Petersen 1923:586(cit); Handschin 1935:690(cit); Adams 1936:25(cit); Stange 1976:306(cat); New 1985b:33(key, rd, ill); Lambkin 1987:12(cit); New 1996:83(cat); Whittington 2002:382(cit); Stange 2004:105(cat); Miller & Stange 2012:14(im); Oswald 2018(cat); Machado et al. 2019(tree).   Diagnosis.Legs elongate; tibial spurs reaching T2 apex; forewing costal area uniareolate; hind wing with two broad dark bands; wings broad; male gonarcus/mediuncus complex tubular; male paramere positioned posterior to the gonarcus/mediuncus complex.   Description.Lengths: forewing: 29–34 mm; hind wing: 27–32 mm. Head ( Figs. 88a–b): Labrumbrown, set with a line of elongate setae. Clypeuspale with brown spots in some specimens; set with elongate white setae. Fronspredominantly black, except for ventrolateral corners pale; beset with white setae. Genapale to light brown. Vertexraised; in anterior view pale, with a median black line, usually interrupted centrally; in dorsal view mostly black except for some pale areas around center posteriorly; set with short black setae. Ocularsetae absent. Antennaeclubbed; elongate,> 3x length of pronotum; distance between antennae wider than scape width; all segments dark-brown to black, except by three to five subapical flagellomeres pale in some specimens; torular membrane yellow; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae. Mandiblespale to light brown, but darker at tip. Palpi, maxillary and labial pale to light brown with distal segments generally darker, apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially. Thorax ( Fig. 88b): Pronotumlonger than wide; posterior margin wider than anterior; subapical furrow present; mostly pale with a broad black longitudinal band on midline (flanked by thinner pale longitudinal bands), margins generally black; set with short black setae, and long setae at margins. Mesonotummostly black, with two rounded pale spots on prescutum, scutum with two small pale spots near wing bases and two thin pale longitudinal bands on each side, scutellum black with pale spots laterally or posterolaterally. Metanotumusually black with two pale longitudinal bands. Pterothoracic pleurablack but with few pale areas generally between sclerites; set with elongate white setae; Miller’s organ present. Wings ( Fig. 88c): Fairly broad with tip acute; anterior Banksian line present in both wings, posterior absent; veins set with short black setae, in general brown, except for some specimens with main longitudinal veins are intercalated by numerous short white regions. Male pilula axillarispresent. Forewingmembrane mostly hyaline with brown marks generally concentrated on posterior margin, tip, pterostigma, rhegmal area, and around crossveins; in some specimens the brown areas are darker; CuA fork located between RP origin and first fork; three presectoral crossveins; some subcostal veinlets forked and irregularly linked in some specimens; posterior area narrower than prefork area. Hind wingmembrane mostly hyaline except for two broad brown subapical bands; MP fork located between RP origin and first fork; subcostal veinlets simple; one presectoral crossvein, with a medial spur. Legs ( Figs. 88a–b): All pairs of legs, femur elongate (> 3.5x length of coxa); tibia slightly longer than femur but longer than tarsi; tibial spurs surpassing T2 tip; T2, T3 and T4 about same size, T1 about twice as long as T2, T5 about twice as long as T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae; femur, tibia and tarsi set with short black setae, and scattered long black setae; T5 ventrally with two rows of thick, long, black setae. Prolegsense hair absent; tibia with antennal cleaning setae ventroapically; colouration varies in intensity among specimens, but in general the femur dorsal surface and tip blackish, remaining areas pale, tibia blackish with two or three pale rings, T1 and base of T2 pale, remaining tarsomeres blackish. Mesolegwith same colour pattern of proleg except for a dorsal subapical yellow spot on femur. Metalegwith femur and tibia slightly longer than in other legs; femur, tibia and T5 pale except for their blackish tip; four basal tarsomeres generally pale but darker in some specimens. Abdomen: Usually entirely black or dark brown, occasionally with irregular pale spots on tergites and sternites. Male Terminalia ( Figs. 89a–d): Ectoproctposterior margin rounded in lateral view, set with elongate black setae. 9 thsternitein lateral view with posterior margin longer than anterior; in ventral view posterior margin with a shallow medial concavity; set with elongate black setae. Gonarcusand mediuncusfused forming a tubular structure. Paramerelocated posteriorly to gonarcus/mediuncus complex; long, curved and with tip acute. Female Terminalia ( Figs. 89e–g): Ectoproctposterior margin rounded set with long black setae. Lateral gonapophysesrounded, shorter than ectoproct, and set with cavisetae. 7 thsternitewith distal margin elongate and covered with long black setae in lateral view. Pregenital platelarge, squared shaped with dorsolateral margins elongate. Posterior gonapophysesbroad, elongate, set with very long black setae and, a group of cavisetae apically. 9 thtergitewith an elongate membranous digitiform process. Anterior gonapophysesa small rounded sclerotized plate, located at base of gonapophyseal plates.   Distribution( Fig. 87). Australia: ACT, NSW, QLD, SA*, VIC*. Widespread and common in eastern Australia; apparently absent from western and central Australia.  Adult activity period.Records for September to April.   Biology. New (1996)stated that the larva is terrestrial and a predator. Miller and Stange (2012)reared a female from a larva collected in coarse sand at the entrance of a cave in Pilliga Nature Reserve, NSW. They did not describe the larva, but mentioned that it had a complex colour pattern and the distinctive Dendroleontinaemedial tuft.   FIGURE 89.  Glenoleon pulchellus: male: a) terminalia, lateral; b) 9 thsternite, ventral; c) genitalia, lateral; d) genitalia, dorsal; female: e) terminalia, lateral; f) ventral margin of 7 thsternite and pregenital plate, posterior; g) terminalia, ventral. Scale bars = 0.5 mm.  Name-bearing type.  Myrmeleon pulchellus: Holotype(by explicit monotypy), male, OUMNH, high resolution images examined. Type locality: Australia(no futher locality data). From original description: “D'après un individu en assez mauvais état, venant de la Nouvelle-Hollande” (According to one individual in poor condition, from New Holland). New Hollandrefers to Australia. From New (1985b): “ Australia(?Paris, not seen)”. From Tauber et al.(2019, fig. 93): “One specimen in the Hope Collection (NEUR0058, Fig. 93), a male, carries Rambur’s labels and is identified as a type. We consider it to be the  M. pulchellus holotype(by explicit monotypy). The abdomen is damaged, and some legs are missing”. Until recently the holotypeof  G. pulchelluswas thought to reside in the MNHN, e.g., New (1985b), Stange (2004), and Oswald (2018), but it had never been confirmed to exist there. For this study we attempted to obtain information about this specimen from the MNHN, but the specimen was not found there (Dr. André Nel, personal communication). More recently, Tauber et al.(2019)reported on multiple specimens of  G. pulchellusfound in the OUMNH. One of these specimens bears labels in Rambur’s handwriting and a post-Rambur label that identifies it as a “Type” that was accessioned into the OUMNH as part of the Marchal Collection. Numerous other Rambur type specimens of neuropterous insects (in the Linnean sense) are known to have entered the OUMNH by way of the Marchal Collection, so the provenance of this specimen is consistent with its identification as the original holotypeof  G. pulchellus. Condition: good, antennae broken, right foreleg missing and abdomen damaged.  Additional material examined( 142♂, 211♀, 8?).   AUSTRALIA: 1917 ( 1♀, 1♂, SAMA); ACT: Black Mountain:  10.ii.1952, L.J. Chinnick( 1♀, ANIC);  ii.1967( 1♂, ANIC);   16.i.1950( 1♀, ANIC); Canberra: 11.ii.1967, E. Paramonov( 1♀, ANIC);  23.xi.1929( 1♂, ANIC);   5.ii.1951( 1♂, ANIC);   2.ii.1960( 2♀, 2♂, ANIC);   5.ii.1960( 4♀, 1♂, ANIC, 1♂, FSCA);   8.ii.1960( 2♀, ANIC);   16.ii.1960( 1♂, ANIC); 20.ii.1960, E.F. Riek( 1♀, AMSA);  7.iii.1960( 2♀, 1♂, ANIC); 28.xi.1960, K.G. Sewnson( 1♀, OSU);  12.xii.1960, E.F. Riek( 1♂, TAMU);  15.xii.1960( 1♀, 2♂, ANIC);   27.xii.1960( 2♀, ANIC);   29.xii.1960( 2♀, ANIC);   9.i.1961( 1♀, 1♂, ANIC);   12.i.1961( 1♂, ANIC);   23.iv.1961( 1♀, ANIC); 1.ii.63 ( 1♂, AMSA);  14.ii.1963( 1♀, ANIC);  Cotter River Reserve:  26.xi.1977, G. Daniels( 1♀, AMSA); Mount Majura:  27.xi.1977, G. Daniels( 1♂, 1♀, AMSA);  New South Wales: no label ( 1♂, ANIC); 22.xii.12 ( 1♂, ANIC); Armidale:  20.xii.1954( 1♀, 1♂, ANIC);   23.xi.1959( 1♂, ANIC);   11.xii.1959( 1♀, ANIC);   xii.1960( 2♀, ANIC);  Asquith: 21.ii.66, A.B. Rose( 1♂, AMSA); Ballina: x.22, A.N. Burns( 1♀, AMSA); Bank- stown: near Sydney,  16.xi.1980, B.J. Day( 1♀, AMSA); Banyabba Natural Reserve:  20.i.1971( 1♀, ANIC); Bay- view: 29.x.1970, L.C. Haines( 1♀, AMSA); Belmont:  4.i.1947( 2♂, 1♀, MVM);   30.xii.1946( 1♂, MVM); Bend- along: 24.i.1970,  G. Daniels( 1♂, AMSA); Berowra: Galston Gorgenear Hawkesburry R.,  ii.1927,  T.G. Campbell( 1♀, AMSA);  6 kmNE of Bilpinnear Nurrajong,  2.ii.1980  N.W. Rodd( 1♂, AMSA); Blue Mountains: Clarence,  8.i.1983, N.W. Rodd( 1♂, AMSA); Brindabella Range:  iii.1984( 1♂, ANIC);  Loqnet ValleySchool,  6.ii.1984, at light, L.C. Haines( 1♀, AMSA); Mount Tomah,  9.i.1984,  N.W. Rodd( 1♀, AMSA); Braidwood:  14.i.1934,  Fuller( 1♀, 1♂, ANIC); Bundeena:  12.xi.1960( 1♀, FSCA);  Cabramatta:  10.i.1961( 1♂, FSCA);   1.5miN of Colo River, 23.x.65, G.L. Bush( 1♀, TAMU); Como:  7.i.1911, Waterhousecoll. ( 1♀, ANIC);   18.i.1933( 1♀, MVM);   5.ii.1922( 1♀, MVM);  Conjola:  26.xii.1950( 1♂, MVM);  Coonabarabran:  2.xii.1974( 1♂, ANIC);  Cuttler’s Pass: William’s river,  10.i.1943, A. Musgrave( 1♀, AMSA); Deriah Aboriginal Area:  23 kmE of Narrabri, - 30.345 S– 150.014 E,  10.xi.2009,  Ooline, light, D.R. Britton, J. Recsei( 1♀, AMSA); Ebenezer:  xii.1996( 1♀, ANIC);  Hazelbrook, 1984, M. Dingley( 2♂, 1♀, AMSA);  i.1985, light ( 1♂, AMSA); Elanora Heights:  xi.1986, T. Robinson( 1♀, AMSA); Elizabeth Bay:  29.i.1940,  K.C. McKeown( 1♀, AMSA); Engadine: near Sydney,  23.xii.1976, R. Eastwood( 1♀, AMSA); Fletcher:  29.xii.1960( 1♂, ANIC);  Fraser Park:  2.i.1947( 1♀, MVM); Glenbrook: 15.1.1993, M.S. & B.J. Moulds( 1♂, 1♀, AMSA);  27.xii.1964( 1♀, ANIC);  Gosford: ( 1♂, ANIC); Greta: 1951 ( 1♂, FSCA); Groose Vale:  29.x.1988, N.W. Rodd( 1♂, AMSA); Gundamain National Park:  14.ii.1926,  Mackerras( 1♂, FSCA); Helensburgh:  23.i.1979,  R. Eastwood( 1♀, AMSA); Hornsby:  3.i.1916,  Tillyard( 1♂, ANIC);  9.3 kmS of Ilford,  29.xii.1977,  G. Daniels( 1♀, AMSA); Killara:  22.xii.1934,  M.F. Day( 1♀, ANIC); Killarney Gap: Kaputar National Park, - 30.141 S– 150.066 E,  14.xi.2009,   Callitris& sclerophyll scrub / rocky ridge, MV Lamp, D.R. Britton, J. Recsei( 1♀, AMSA);  6 kmNE of Kyogle, Murwillumbah Road, 9.i, B.J. Day( 1♂, AMSA); Liverpool: Heathcote Road., 33 o56’ S– 150 o56’ E,  13.xii.1998, light, B.J. & C.J.R. Day( 1♂, AMSA); Loftus:  27.xii.1978, J. Olive( 1♂, AMSA); Mann River:  5 kmupstream from Old Glen Innes, Grafton Roadcrossing,  27.xi.1977, M.S. & B.J. Moulds( 1♀, AMSA); Mittagong( 1♂, SAMA); Moonbi:  7.ii.1968,  D.H. Collens( 1♀, ANIC); Mooney Mooney Creek: near Gosforo,  13.i.1977, D.K. McAlpine( 1♀, AMSA); Mount Kaputar National Park:  15.i.1978,  G. Daniels( 1♀, AMSA); Mount Victoria:  4.xii.1946( 1♀, MVM);  Nambucca:  26.x.1949( 1♀, MVM);  Narrabeen:  18.ix.1934( 1♀, ANIC);  Narrabri:  17.xii.1934, M.F. Day( 1♀, ANIC);  xi.1997( 1♀, ANIC);  Nepean River:  x.1931, A. Musgrave( 1♂, AMSA); New- castle: Sugarloaf Mount,  2.i.1947( 1♂, MVM);  Newport: 2.xii.57, G. Dolezal( 1♀, AMSA); Pilliga Nature Reserve:  34 kmN of Coonabarabran,  13.x.2005,  L.A. Stange( 1♀, FSCA); Royal National Park: Goondera Ridge,  3.xii.1977,  G. & A. Daniels( 1♀, AMSA); Singleton:  23.iii.1957, E.F. Riek( 1♂, ANIC); Sutton:  19.i.1979( 1♂, QMBA); Syd- ney: C. Gibbons( 1♀, AMSA);  28.xii.1961, D.H. Collens( 1♂, ANIC);  xii.1946( 1♀, MVM); ( 1♀, SAMA); Waita- ra, 23.i.1983,  M.S. & B.J. Moulds( 1♂, AMSA); Tomerong: Nowra district, 3miN,  7.ii.1970, D.K. McAlpine( 1♀, AMSA); Tuggerah:  xii.1904( 1♀, SAMA);  Ulan:  xi.1982( 1♂, ANIC); Wahroonga: 12.xii.1970,  A.B. Rose( 1♀, AMSA); H.J. Carter( 1♀, AMSA); Wapengo:  12.i.1978( 1♀, MVM); Warialda: Adam’s Scrub 14.6 kmS, - 29.667 S– 150.567 E, 18.xi.2009,  Callitris/sclerophyll forest, MV lamp, D.R. Britton, J. Recsei( 1♂, AMSA); Wastepoint:  8.3 kmN of Jindabyne, KosciuszkoN.P., - 36.347 S– 148.606 E,  8.xii.2007, MV lamp, D.R. Britton, D.J. Smith, P. Hinton( 1♂, AMSA); Watalgan Forrest:  24.xi.1973( 1♀, ANIC); Wedderburr: 3 kmE, 34 o08’ S– 150 o49’ E, 13.xii.2005, MV light, D. Britton( 1♂, AMSA); Wee Jasper: 25.xii.56, E. shipp ( 1♂, AMSA); Wellington:  28.ix.1957,  E.F. Riek( 2♂, ANIC); West Head: PiHwater,  2.i.1985, L.C. Haines( 1♂, AMSA); Whiskers:  xii.1992( 1♂, ANIC);   Queensland: ( 1♀, ZMHB); ( 1♀, SAMA); Archer Rivercrossing: 13 o25’S– 142 o56’E,  iv.1989, mv lamp, G. & A. Daniels(6?, QMBA); Barankula State Forrest: 26 o26’07’’ S– 150 o49’31’’ E,  3–7.iv.2000, malaise, M. Mathieson( 1♀, TAMU); Bardon:  x.1943( 1♀, 1♂, SAMA);  Barrine:  10.x.1967( 2♂, ANIC);  Blackdown Tableland: Expedi- tion Range,  17.i.1987, M.S. & B.J. Moulds( 1♂, AMSA); Bluff Range:  16.xii.1970( 1♀, 1♂, ANIC);   9.i.1971( 2♀, 1♂, ANIC);  Biggenden:  22–23.xi.1977( 12♀, 9♂, ANIC);   1.i.1972( 2♀, ANIC);   2.xii.1974( 1♂, ANIC);  Biloela:  17.i.1947( 1♀, ANIC);  Bin Bin Range:  4.xii.1974( 1♀, ANIC);   12.i.1972( 1♀, 1♂, ANIC);  Bingera:  22.xi.1971( 1♀, ANIC);  Bon Accord Falls: Montville, Blackall Ranges, 22.ix.55, R. Dobson( 1♀, AMSA); Boolaboonda Range:  10.xi.1973( 1♀, 1♂, ANIC); Boompa: 4.xii.1980,  H. Franca( 1♂, ANIC); Bowen:  1.x.1950, E.F. Riek( 1♀, ANIC); Brisbane: ( 6♂, 24♀, QMBA);  i.1888( 2♀, SAMA);   xi.1965( 2♀, ANIC);   20.x.1900( 1♀, ANIC); 1903 ( 1♀, ZMUC);  x.1911( 1♀, SAMA); 20.ii.27, L. Franzen( 2♂, ZMUC);  19.ix.1927( 1♀, QMBA);   30.xii.1927( 1♀, MVM);   19.x.1938( 1♀, ANIC);   9.i.1960( 1♂, 1♀, MVM);   10.i.1960( 1♀, MVM);   24.xii.1955( 1♀, MVM);   27.xii.1959( 1♀, MVM);   2.xii.1922( 2♀, MVM);   29.x.1936( 1♀, MVM);   4.xi.1936( 1♂, MVM);   26.xi.1956( 1♂, MVM);   x.1910( 1♂, 2♀, MVM);  Moggill Farm,  23.i.1961( 1♂, FSCA);  Bucasia:  i.2003( 1♂, ANIC);  Bundaberg:  19.xi.1978( 2♀, 3♂, ANIC);   22.xii.1979( 1♀, 2♂, ANIC);  Bunya Mountains:  17.i.1955( 1♂, MVM);  Burleigh Heads:  3.i.1956( 1♀, MVM);   xi.1942( 1♂, MVM);   xii.1942( 1♂, MVM);   10.x.1956( 1♀, MVM); Byfield State Forrest: x.1924, A. Musgrave( 1♂, AMSA);  1.i.1976, G. Daniels( 1♀, AMSA); Carnarvon National Park:  7.xii.1938, N. Geary( 2♀, AMSA);  8.xii.1938( 1♂, AMSA);   12.xii.1938( 1♂, 1♀, AMSA);   5.i.1939( 1♀, AMSA);   8.i.1939( 1♂, AMSA);   ii.1944( 1♀, AMSA); 92 air km NNW of Injune, 25.06695°S 148.24611°E±  30m, R Machado, MV light,  25.xii.2015, AustRM#10 ( 2♀, TAMU); Charleville:  27.xii.1925( 1♀, QMBA); Cordalba National Forrest: 30 kmS of Bunda- berg, 29.xi.1979, H. frauca ( 1♀, ANIC); Degilbo:  9.xii.1978( 2♂, ANIC);  Electra:  8.ii.1977( 1♂, ANIC); Fletcher: 12.i.1967, E. Sutton( 1♂, ANIC); Forty Mile Scrub:  4.x.1989( 1♂, ANIC);  Hayman Island:  24.x.1950,  R. Dobson( 3♀, 1♂, ANIC); Herberton:  6.xii.1968, E. Britton& S. Misko( 1♂, ANIC);  x.1990( 1♀, ANIC);  55 kmNNE of Injune,  23.xi.1986,  M.S. & B.J. Moulds( 1♂, AMSA); Kinbombi falls: Goomeri,  19.xii.1976, M.S. & B.J. Moulds( 1♀, AMSA); Kingston:  xii.1981( 1♂, QMBA);  Kuranda: ( 1♀, SAMA);  19.xi.1951( 1♀, MVM);   5.x.1949( 1♀, MVM);  Mackay: 1905 ( 1♀, 1♂, ANIC); Milmerran:  14.xi.1945( 1♂, MVM);  Montville: Bon Accord Falls,  20.ix.1955( 1♂, 1♀, MVM);  Mount Cootha:  x.1942( 1♀, SAMA);   16.xii.1973( 2♀, QMBA);  Mount Garmet:  i.1961( 1♂, ANIC);  Mount Hay:  24.i.2009( 1♀, QMBA);  Mount Moffat National Park: Mahogany Forest,  11–12.xii.1987, M.T. Yeates( 1♂, 2♀, QMBA); Mount Tamborine:  30.iii.1950( 1♂, MVM);   26.xii.1950( 1♀, MVM);  MountTin- beerwah:  1.i.1979( 1♀, QMBA);  Murphy’s Creek:  29.xi.1992( 1♀, QMBA);  Noosa Heads:  1.x.1956( 1♀, MVM);  Ravenshoe:  xi.1998( 1♀, ANIC);  Stanthorpe:  30.i.1927( 1♀, QMBA);   29.xii.1977( 2♂, QMBA; 1♂, ANIC);   9.ii.1981(1?, QMBA);  Stradbroke Island:  21.ix.1954( 1♀, MVM);  Tamborine: ( 1♂, QMBA); Toowoomba:  11.ii.1960( 1♂, ANIC);  Townsville:  12.xii.1908( 1♀, MVM);  Wacol:  16.x.1952( 1♂, QMBA);  West Burleigh: 1960, A.T. Bandsoma( 1♂, AMSA); Woodridge:  2.i.1968( 1♂, MVM);   South Australia: Aldgate: 19.iii.19309( 1♀, MVM); Barossa: ( 1♀, SAMA); Blackwood:  ii.1965( 2♀, SAMA);   20.i.1969( 1♀, MVM; 2♀, SAMA);   xii.1975( 1♀, SAMA);  Bridgewater:  xii.1967( 1♀, SAMA);  Cara Pook:  i.1973( 1♂, SAMA);  Heywood Park:  xii.1958( 1♂, SAMA);  Highbury:  i.1969( 1♂, SAMA);  Kangaroo Creek Reserve:  i.1971( 1♂, SAMA); “Kurlge” Blackwood:  xi.1957( 1♀, SAMA);   xii.1957( 1♂, SAMA);   i.1958( 2♀, SAMA);   ii.1958( 1♀, SAMA);  Mount Lofty Range:  ii.1965( 1♀, SAMA);   xii.1973( 1♂, SAMA);  Nairne:  i.1988( 1♂, SAMA);  Locationnot recognized: Saunder’s Coll., iii.68 ( 1♀, BMNH); Nolabel ( 2♀, 3♂, ANIC);  Victoria: Bendigo:  5.i.1930( 1♀, MVM);  Hurstbridge:  18.ii.1947( 1♂, MVM);  Monbulk:  27.i.1929( 1♀, MVM);  Nowa Nowa:  21.ii.1947( 1♂, MVM); Moe: 24.xii.1944 ( 1♂, MVM); Oakleigh: ( 1♂, MVM); Research:  ii.1983- ( 1♀, ANIC); Warrandyte:  i.1988( 1♀, ANIC); Wartook:  xii.1935( 1♀, SAMA).   Comments.  Glenoleon pulchellusis a common species in eastern Australia. Its color intensity varies considerably among specimens; some are almost completely dark, while others have many pale areas. Despite these coloration differences the species is easily recognized by the two dark bands in the apical half of the hind wing; no other species in the  Periclystusgenus group has this character. The long legs and pronotum, and broad wings, of this species support its close relationship with the several other species retained here in  Glenoleon, and is further demonstrated by both molecular ( Machado et al. 2019) and other morphological data ( Fig. 4). 1952-02-10 SAMA Australia Black Mountain 181 182 2 1 1 [339,883,765,791] 1967-02 ANIC Chinnick Australia 181 182 2 1 1 [896,1151,764,791] 1950-01-16 ANIC Chinnick Australia true 181 182 1 1 1929-11-23 ANIC Paramonov Australia 181 182 2 1 1 [714,960,800,827] 1951-02-05 ANIC Paramonov Australia true 181 182 1 1 [971,1266,801,827] 1960-02-02 ANIC Australia 181 182 4 2 2 1960-02-05 ANIC, FSCA Australia 181 182 6 4 2 [418,660,837,863] 1960-02-08 ANIC Australia 181 182 2 2 [670,926,837,863] 1960-02-16 ANIC Australia 181 182 1 1 1960-03-07 AMSA, ANIC Riek Australia 181 182 4 3 1 [482,918,873,899] 1960-12-12 OSU Sewnson Australia 181 182 1 1 1960-12-15 TAMU, ANIC Riek Australia 181 182 4 1 3 [249,533,909,935] 1960-12-27 ANIC Riek Australia true 181 182 2 2 [550,834,909,935] 1960-12-29 ANIC Australia 181 182 2 2 [851,1153,909,935] 1961-01-09 ANIC Australia 181 182 2 1 1 [1170,1432,909,935] 1961-01-12 ANIC Australia 181 182 1 1 [151,414,945,971] 1961-04-23 ANIC Australia 181 182 1 1 [425,922,945,971] 1963-02-14 AMSA, ANIC Australia 181 182 2 1 1 [933,1305,945,972] 1977-11-26 Australia Cotter River Reserve 181 182 1 1977-11-27 AMSA Daniels Australia Mount Majura 181 182 1 1 1954-12-20 AMSA, ANIC Daniels Australia Armidale 181 182 6 2 4 New South Wales [875,1143,1016,1043] 1959-11-23 ANIC Daniels Australia true Armidale 181 182 1 1 New South Wales [1156,1432,1016,1043] 1959-12-11 ANIC Australia 181 182 1 1 [151,393,1053,1079] 1960-12 ANIC Australia 181 182 2 2 1980-11-16 AMSA Asquith & Rose & Burns Australia Sydney Ballina 181 182 2 1 1 [524,1369,1089,1115] 1971-01-20 AMSA, ANIC Day Australia Banyabba Natural Reserve 181 182 2 2 [349,1064,1125,1151] 1947-01-04 AMSA, MVM Haines Australia Belmont 181 182 4 2 2 [1077,1355,1125,1151] 1946-12-30 MVM Haines Australia true Belmont 181 182 1 1 [349,1259,1161,1187] 1927-02 AMSA G. Daniels & Hawkesburry R. Australia Galston Gorge Berowra 181 182 1 1 1980-02-02 AMSA T. G. Campbell Australia Nurrajong 181 182 1 1 1983-01-08 AMSA N. W. Rodd Australia Clarence Blue Mountains 181 182 1 1 [259,1025,1233,1259] 1984-03 AMSA, ANIC Rodd Australia Brindabella Range 181 182 2 2 [1038,1403,1233,1259] 1984-02-06 Australia Loqnet Valley 181 182 1 1984-01-09 AMSA Haines Australia Mount Tomah 181 182 1 1 [808,1356,1269,1295] 1934-01-14 AMSA N. W. Rodd Australia Braidwood 181 182 1 1 1960-11-12 ANIC, FSCA Fuller Australia Bundeena 181 182 3 2 1 [762,1162,1305,1331] 1961-01-10 FSCA Cabramatta Australia 181 182 1 1 1911-01-07 TAMU, ANIC G. L. Bush & Waterhouse Australia Como 181 182 2 2 [1070,1325,1341,1368] 1933-01-18 MVM Australia 181 182 1 1 1922-02-05 MVM Australia 181 182 1 1 [305,687,1376,1404] 1950-12-26 MVM Conjola Australia 181 182 1 1 [699,1145,1377,1403] 1974-12-02 ANIC Australia Coonabarabran 181 182 1 1 1943-01-10 Cuttler's Pass Australia William's river 181 182 1 2009-11-10 AMSA Musgrave Australia -30.345 Deriah Aboriginal Area 72 150.014 181 182 1 1 [285,1218,1449,1476] 1996-12 AMSA, ANIC Ooline & D. R. Britton & Recsei Australia Ebenezer 181 182 2 2 1985-01 AMSA Hazelbrook & Dingley Australia 181 182 3 1 2 [599,1116,1485,1511] 1986-11 AMSA Australia Elanora Heights 181 182 1 1 1940-01-29 AMSA Robinson Australia Elizabeth Bay 181 182 1 1 [451,1217,1521,1548] 1976-12-23 AMSA K. C. McKeown Australia Sydney Engadine 181 182 1 1 1960-12-29 AMSA, ANIC Eastwood Australia Fletcher 181 182 2 1 1 [647,1031,1557,1584] 1947-01-02 MVM Fraser Park Australia 181 182 1 1 1964-12-27 AMSA, ANIC Moulds Australia 181 182 3 2 1 1988-10-29 ANIC, FSCA Gosford Australia Groose Vale Greta 181 182 2 2 [277,996,1629,1656] 1926-02-14 AMSA Rodd Australia Gundamain National Park 181 182 1 1 1979-01-23 FSCA Mackerras Australia Helensburgh 181 182 1 1 [272,790,1665,1692] 1916-01-03 AMSA R. Eastwood Australia Hornsby 181 182 1 1 [803,1397,1665,1691] 1977-12-29 ANIC Tillyard Australia 9.3 km S of Ilford 181 182 1 1 1934-12-22 AMSA G. Daniels Australia Killara 181 182 1 1 2009-11-14 ANIC M. F. Day Australia -30.141 Kaputar National Park 73 150.066 Killarney Gap 181 182 1 1 1998-12-13 AMSA Lamp & D. R. Britton & J. Recsei & B. J. Day Australia Murwillumbah Road -33.933334 Heathcote Road. 1202 150.93333 Liverpool 181 182 2 1 1 [446,1102,1809,1835] 1978-12-27 AMSA B. J. & C. J. R. Day Australia Loftus 181 182 1 1 1977-11-27 AMSA Olive Australia Grafton Road Mann River 181 182 1 1 1968-02-07 AMSA, SAMA Moulds Australia Moonbi Mittagong 181 182 2 1 1 1977-01-13 ANIC D. H. Collens Australia Gosforo Mooney Mooney Creek 181 182 1 1 [270,1064,1916,1943] 1978-01-15 AMSA McAlpine Australia Mount Kaputar National Park 181 182 1 1 1946-12-04 AMSA, MVM G. Daniels Australia Mount 181 182 2 2 Victoria [526,919,1953,1979] 1949-10-26 MVM Nambucca Australia 181 182 1 1 [929,1323,1953,1979] 1934-09-18 ANIC Narrabeen Australia 181 182 1 1 1934-12-17 Australia Narrabri 181 182 1 [293,787,1989,2016] 1997-11 ANIC Day Australia 181 182 2 2 [798,1047,1989,2016] 1931-10 Australia Nepean River 181 182 1 1947-01-02 AMSA, MVM Musgrave Australia 182 183 Sugarloaf Mount 181 182 2 2 2005-10-13 AMSA Newport & Dolezal Australia Pilliga Nature Reserve 182 183 1 1 [590,1432,189,215] 1977-12-03 FSCA L. A. Stange Australia Goondera Ridge Royal National Park 182 183 1 1 [151,746,224,251] 1957-03-23 AMSA G. & A. Daniels Australia Singleton 182 183 1 1 [757,1372,225,251] 1979-01-19 ANIC, QMBA Riek Australia Sutton 182 183 2 2 1961-12-28 AMSA Gibbons Australia 182 183 1 1 [644,1352,261,287] 1946-12 ANIC, MVM, SAMA Collens Australia 182 183 3 2 1 [307,1192,297,323] 1970-02-07 AMSA M. S. & B. J. Moulds Australia Nowra district Tomerong 182 183 1 1 1904-12 AMSA, SAMA McAlpine Australia Tuggerah 182 183 2 2 [651,957,333,359] 1982-11 ANIC Ulan Australia 182 183 1 1 1978-01-12 AMSA, MVM A. B. Rose & H. J. Carter Australia Wapengo 182 183 3 3 2007-12-08 AMSA D. R. Britton & J. Recsei Australia -36.347 Kosciuszko 70 148.606 Waste 182 183 1 1 1973-11-24 AMSA, ANIC D. R. Britton & Smith & Hinton Australia Watalgan Forrest 182 183 2 1 1 [290,1432,512,539] 1957-09-28 AMSA Britton Australia Wellington Wee Jasper 182 183 2 2 [151,760,548,575] 1985-01-02 ANIC E. F. Riek Australia PiHwater West Head 182 183 2 2 [771,1432,549,575] 1992-12 AMSA, ANIC Haines Australia Whiskers 182 183 2 2 [151,1216,584,611] 1989-04 ZMHB, SAMA Australia -13.416667 Archer River 1291 142.93333 182 183 2 2 Queensland 2000-04-03 2000-04-07 2000-04-03 QMBA G. & A. Daniels Australia -26.435276 Barankula State Forrest 20 150.82527 182 183 1 1943-10 TAMU, SAMA Mathieson Australia Bardon 182 183 3 2 1 [715,1074,657,683] 1967-10-10 ANIC Barrine Australia 182 183 2 2 1987-01-17 Blackdown Tableland Australia Range 182 183 1 [407,1275,693,719] 1970-12-16 AMSA, ANIC Moulds Australia Bluff Range 182 183 3 1 2 1971-01-09 ANIC Moulds Australia true Bluff Range 182 183 3 2 1 [294,807,729,755] 1977-11-22 1977-11-23 1977-11-22 ANIC Biggenden Australia 182 183 21 12 9 [820,1060,729,755] 1972-01-01 1977-11-23 1977-11-22 ANIC Biggenden Australia true 182 183 2 2 [1073,1335,729,755] 1974-12-02 ANIC Australia 182 183 1 1 1947-01-17 ANIC Biloela Australia 182 183 1 1 [409,842,765,791] 1974-12-04 ANIC Bin Bin Range Australia 182 183 1 1 [852,1148,765,791] 1972-01-12 ANIC Bin Bin Range Australia true 182 183 2 1 1 1971-11-22 ANIC Bingera Australia 182 183 1 1 1973-11-10 AMSA, ANIC Montville & Blackall Ranges & Dobson Australia Boolaboonda Range Bon Accord Falls 182 183 3 2 1 [712,1172,836,863] 1950-10-01 ANIC H. Franca Australia Bowen 182 183 1 1 1888-01 ANIC, QMBA, SAMA Riek & Brisbane Australia 182 183 33 27 6 [723,949,872,899] 1965-11 ANIC Riek & Brisbane Australia true 182 183 2 2 [958,1211,873,899] 1900-10-20 ANIC Australia 182 183 1 1 1911-10 ZMUC, SAMA Australia 182 183 2 2 [415,1128,909,935] 1927-09-19 ZMUC, QMBA Franzen Australia 182 183 3 1 2 [1145,1432,909,935] 1927-12-30 MVM Franzen Australia true 182 183 1 1 [151,414,945,971] 1938-10-19 ANIC Australia 182 183 1 1 [428,725,945,971] 1960-01-09 MVM Australia 182 183 2 1 1 [739,999,945,971] 1960-01-10 MVM Australia 182 183 1 1 [1013,1295,944,971] 1955-12-24 MVM Australia 182 183 1 1 1959-12-27 MVM Australia 182 183 1 1 [311,581,981,1008] 1922-12-02 MVM Australia 182 183 2 2 [596,864,981,1008] 1936-10-29 MVM Australia 182 183 1 1 [879,1141,981,1008] 1936-11-04 MVM Australia 182 183 1 1 [1156,1432,980,1008] 1956-11-26 MVM Australia 182 183 1 1 [151,420,1017,1044] 1910-10 MVM Australia 182 183 3 2 1 [430,848,1017,1044] 1961-01-23 FSCA Moggill Farm Australia 182 183 1 1 [857,1171,1017,1043] 2003-01 ANIC Bucasia Australia 182 183 1 1 1978-11-19 ANIC Bundaberg Australia 182 183 5 2 3 [348,668,1053,1079] 1979-12-22 ANIC Bundaberg Australia true 182 183 3 1 2 [679,1145,1052,1080] 1955-01-17 MVM Australia Bunya Mountains 182 183 1 1 1956-01-03 MVM Burleigh Heads Australia 182 183 1 1 [304,537,1089,1116] 1942-11 MVM Burleigh Heads Australia true 182 183 1 1 [548,789,1089,1116] 1942-12 MVM Australia 182 183 1 1 [800,1060,1088,1116] 1956-10-10 MVM Australia 182 183 1 1 1976-01-01 AMSA Musgrave Australia 182 183 1 1 [545,1258,1125,1151] 1938-12-07 AMSA Daniels Australia Carnarvon National Park 182 183 1 1 1938-12-08 AMSA Geary Australia 182 183 3 2 1 [550,893,1161,1187] 1938-12-12 AMSA Geary Australia true 182 183 2 1 1 [907,1163,1160,1187] 1939-01-05 AMSA Australia 182 183 1 1 [1177,1432,1161,1187] 1939-01-08 AMSA Australia 182 183 1 1 [151,384,1197,1223] 1944-02 AMSA Australia 182 183 1 1 [393,1431,1196,1224] 2015-12-25 Australia 30 -25.06695 Machado 1 148.24611 182 183 1 [151,895,1232,1259] 1925-12-27 TAMU, QMBA Australia Charleville 182 183 3 3 [349,975,1269,1295] 1978-12-09 ANIC Australia Degilbo 182 183 3 1 2 [986,1325,1269,1295] 1977-02-08 ANIC Electra Australia 182 183 1 1 [272,990,1305,1331] 1989-10-04 ANIC Sutton Australia Forty Mile Scrub 182 183 2 2 [1002,1303,1304,1331] 1950-10-24 Australia Hayman Island 182 183 1 1968-12-06 ANIC R. Dobson Australia Herberton 182 183 4 3 1 [613,1253,1341,1367] 1990-10 ANIC Britton & Misko Australia 182 183 2 1 1 1986-11-23 Australia Injune 182 183 1 [371,1196,1377,1403] 1976-12-19 AMSA M. S. & B. J. Moulds Australia Goomeri Kinbombi falls 182 183 1 1 1981-12 AMSA, QMBA Moulds Australia Kingston 182 183 2 1 1 [710,1264,1412,1439] 1951-11-19 SAMA, MVM Kuranda Australia 182 183 2 2 1949-10-05 MVM Kuranda Australia true 182 183 1 1 [252,1071,1448,1475] 1945-11-14 ANIC, MVM Mackay Australia Milmerran 182 183 3 1 2 1955-09-20 MVM Montville Australia Bon Accord Falls 182 183 2 1 1 [474,882,1485,1511] 1942-10 SAMA Australia Mount Cootha 182 183 1 1 [891,1178,1485,1511] 1973-12-16 QMBA Australia true Mount Cootha 182 183 2 2 1961-01 ANIC Australia Mount Garmet 182 183 1 1 [299,708,1521,1547] 2009-01-24 QMBA Australia Mount Hay 182 183 1 1 [718,1431,1521,1547] 1987-12-11 1987-12-12 1987-12-11 Australia Mahogany Forest Mount Moffat National Park 182 183 1 [151,1004,1556,1584] 1950-03-30 QMBA, MVM Yeates Australia Mount Tamborine 182 183 4 2 2 [1016,1294,1556,1584] 1950-12-26 MVM Yeates Australia true Mount Tamborine 182 183 1 1 1979-01-01 QMBA Mount Australia 182 183 1 1 [533,1012,1593,1619] 1992-11-29 QMBA Australia Murphy's Creek 182 183 1 1 [1023,1432,1592,1619] 1956-10-01 MVM Noosa Heads Australia 182 183 1 1 [151,535,1629,1655] 1998-11 ANIC Ravenshoe Australia 182 183 1 1 [552,973,1629,1655] 1927-01-30 QMBA Stanthorpe Australia 182 183 1 1 [990,1432,1629,1655] 1977-12-29 QMBA, ANIC Australia 182 183 3 3 [151,419,1665,1691] 1981-02-09 QMBA Australia 182 183 1 [437,948,1664,1691] 1954-09-21 MVM Australia Stradbroke Island 182 183 1 1 1960-02-11 QMBA, ANIC Tamborine Australia Toowoomba 182 183 2 2 [416,826,1701,1727] 1908-12-12 MVM Townsville Australia 182 183 1 1 [836,1192,1700,1727] 1952-10-16 QMBA Wacol Australia 182 183 1 1 1968-01-02 AMSA, MVM Bandsoma Australia Woodridge West Burleigh 182 183 2 2 1965-02 MVM, SAMA Aldgate & Barossa Australia Blackwood 182 183 4 4 South Australia [922,1330,1773,1799] 1969-01-20 MVM, SAMA Australia 182 183 3 3 1975-12 SAMA Australia 182 183 1 1 [314,719,1809,1835] 1967-12 SAMA Bridgewater Australia 182 183 1 1 [730,1092,1809,1835] 1973-01 SAMA Cara Pook Australia 182 183 1 1 1958-12 SAMA Heywood Park Australia 182 183 1 1 [259,620,1845,1871] 1969-01 SAMA Highbury Australia 182 183 1 1 [634,1174,1845,1871] 1971-01 SAMA Australia Kangaroo Creek Reserve 182 183 1 1 1957-11 SAMA Blackwood Australia 182 183 1 1 [416,675,1880,1907] 1957-12 SAMA Blackwood Australia true 182 183 1 1 [689,926,1880,1907] 1958-01 SAMA Australia 182 183 2 2 [940,1185,1880,1907] 1958-02 SAMA Australia 182 183 1 1 1965-02 SAMA Australia Mount Lofty Range 182 183 1 1 [410,668,1917,1943] 1973-12 SAMA Australia true Mount Lofty Range 182 183 1 1 [683,1014,1917,1943] 1988-01 SAMA Nairne Australia 182 183 1 1 1930-01-05 BMNH, ANIC, MVM Location & Saunder's Coll. & No Australia Bendigo 182 183 7 4 3 Victoria 1947-02-18 MVM Hurstbridge Australia 182 183 1 1 [426,799,1989,2015] 1929-01-27 MVM Monbulk Australia 182 183 1 1 [811,1230,1989,2015] 1947-02-21 MVM Nowa Nowa Australia 182 183 1 1 [308,571,153,179] MVM 183 184 1 1 [584,941,153,179] 1983-02 ANIC 183 184 1 1 [953,1320,153,179] 1988-01 ANIC 183 184 1 1 1935-12 SAMA 183 184 1 1