Platyleon froggatti Esben-Petersen, 1923:578 Morphological phylogeny and taxonomic revision of the former antlion subtribe Periclystina (Neuroptera: Myrmeleontidae: Dendroleontinae) Machado, Renato Jose Pires Oswald, John David Zootaxa 2020 2020-06-16 4796 1 1 322 (Esben-Petersen, 1923) Machado & Oswald 2020 Esben-Petersen 1923 [151,683,441,468] Insecta Myrmeleontidae Glenoleon Animalia Neuroptera 162 163 Arthropoda species froggatti comb. nov.  ( Figs. 78–80) http://zoobank.org/NomenclaturalActs/ 21B0223B-62EE-44BD-B04A-FB37723131EA    Platyleon froggatti Esben-Petersen, 1923:578(OD); Stange 1976:312(cat); New 1985b:10(key, rd, ill); New 1996:91(cat); Stange 2004:107(cat); Oswald 2018(cat).   Diagnosis.Legs elongate; tibial spurs reaching T2 apex; forewing costal area biareolate; hind wing without broad bands on apical half; male paramere located posterior to gonarcus/mediuncus complex; female pregenital plate conically invaginated.   Description.Lengths: forewing: 24–28 mm; hind wing: 22–26 mm. Head ( Figs. 79a–b): Labrumpale; set with a line of elongate setae. Clypeuspale, with some elongate pale setae. Fronsdark brown except for ventrolateral corners pale; set with short white setae. Genapale. Vertexraised; pale except for two rounded dark brown marks around midline in dorsal view, and some irregular lateral dark marks on posterior border; covered with short black setae. Ocularsetae absent. Antennaeclubbed; elongate,> 2x length of pronotum; distance between antennae wider than scape width; scape and pedicel brown with distal margin pale; flagellum mostly brown, except for five to six subapical segments pale and apex dark brown; torular membrane dark brown; flagellomeres almost as long as wide at base, apical ones much wider than long; all segments set with short black setae, except for scape and pedicel with few short white setae. Mandiblepale to brown, with tip black. Palpi, maxillary and labial mostly pale with irregular dark brown marks mainly on basal segments; apical labial palpomere fusiform, palpimacula opening oval-shaped, located medially Thorax ( Fig. 79b): Pronotumlonger than wide; posterior margin slightly wider than anterior; subapical furrow present; mostly pale except for some dark brown areas: a broad sagittal line (forked on anterior margin in some specimens), two thin and sinuous longitudinal lines near midline (interrupted medially in some specimens), and longitudinal marks on lateral margin; beset with short black setae, and some elongate setae on margins. Mesonotummostly dark brown, with pale areas on each segment: two curved marks on prescutum, two large marks around midline but enclosing a short dark brown line, and a sinuous line laterally at scutum; beset with short black setae. Metanotumdark brown, except for large pale marks at scutum, and some irregular pale areas on scutellum of some specimens; covered with short black setae. Pterothoracic pleuramostly pale, except for dark brown marks, mainly at ventral margin of sclerites; covered with white setae; Miller’s organ present. Wings ( Fig. 79c): Fairly broad; anterior Banksian line present in both wings, but posterior absent; veins mostly dark brown, but some veins intercalated with white spots, beset with short black setae. Male pilula axillarispresent. Forewingmembrane mostly hyaline with some brown areas as follows: surrounding veins at tip of prefork area, beneath pterostigma, around gradate veins, posterior margin and tip with small brown infuscations; CuA fork located between RP origin and first fork; three presectoral crossveins; costal area large, biareolate, with most of subcostal veinlets forked; posterior area narrower than prefork area. Hind wingmembrane mostly hyaline except for a small brown mark hypostigmatic cell base, a large mark on rhegmal area, and small brown infuscations at tip; MP fork located between RP origin and first fork; subcostal veinlets mostly simple but few forked; one presectoral crossvein with a medial spur in some specimens. Legs ( Figs. 79a–b): All pairs of legs, femur elongate (> 3.5x length of coxa); tibia slightly longer than femur and about 1.5x longer than tarsi; tibial spurs reaching T2 apex; T2, T3 and T4 about same size, T1 about twice longer than T2, T5> 2x length of T1; claws about half of T5 length; coxa, trochanter, and femur set with short white setae and few scattered long white setae at femur; femur, tibia, and tarsi set with long and short black setae; T5 ventrally with two rows of thick, long, black setae. Prolegsense hair slightly longer than femur width; tibia with antennal cleaning setae ventroapically; coxa pale with irregular dark marks; trochanter pale, femur and tibia pale to brown with apex dark brown; tarsi pale with T3, T4, and apex of T5 dark brown. Mesolegcolour pattern similar to proleg, except for coxa entirely pale. Metalegsimilar to mesoleg except with femur and tibia slightly longer than in other legs. Abdomen: Mostly dark brown with irregular pale marks, mainly at lateral margins on tergites and medial areas of sternites. Beset with short black setae, and some white ones on basal segments, mainly sternites. Male Terminalia ( Figs. 80a–d): Ectoproctposterior margin rounded in lateral view, in posterior view with inner ventral margin enlarged; covered with elongate black setae. 9 thsterniteshort with posterior margin rounded, in ventral view; set with very long black setae. Gonarcus/ mediuncuscomplex long and tubular. Paramerestrongly sclerotized, fused at apex of gonarcus/mediuncus complex; curved with tip acute.   FIGURE 79.  Glenoleon froggatti: a) head, anterior; b) head and thorax, dorsal; c) wings, dorsal. Female Terminalia ( Figs. 80e–f): Ectoproctposterior margin rounded, covered with thin elongate setae. Lateral gonapophysesrounded, smaller than ectoproct, covered with cavisetae. 7 thsternitelong with distal margin straight in ventral view, covered with long black setae. Pregenital platelarge and conically invaginated. Posterior gonapophysesbroad, elongate, curved and with tip rounded; covered with long black setae, and many elongate cavisetae apically. 9 thtergitewith a short membranous digitiform process. Anterior gonapophysesas two internal small lobes located at base of posterior gonapophyses, bearing short thickened setae; only seem in ventral view. Ventral membranewith gonapophyseal plates present, and with a sclerotized rectangular bar ventral to the anterior gonapophyses, also bearing short thickened setae.   Distribution ( Fig. 78). Australia: NSW, NT*, QLD, VIC*. Known from scattered records in eastern Australia, with an outlier in southern Northern Territory.  Adult activity period.Records for November to February.   Biology.Nothing published exists for this species, but one male deposited in the FSCA was reared by Robert B. Miller in 1998 from a larva collected in a tree hole near a river in Queensland. The larval collection site suggests that  G. froggattilarvae may develop in tree holes like many other Dendroleontinispecies ( Stange 1976; Stange & Miller 1990). The exuviae also shows that the larva possesses three large mandibular teeth, and the typical Dendroleontinaesetal tuft located medially on mesonotum ( Stange & Miller 1990).  Name-bearing type.  Platyleon froggatti:  Holotype(by explicit monotypy), female, ANIC, examined. Type locality: Australia, New South Wales, Hay(town). Fromoriginal description: “One ♀, N.S. Wales,  20.12.1915( W. W. Froggattleg.). Coll. Froggatt”. From New 1985b: “ Holotype, ♀, New South Wales, Hay (Eucalyptus),  20.xii.1915, W. W. Froggattleg. ( ANIC) (seen) (antennae missing)”. Condition: relatively good; antennae missing, abdomen broken but stored with the specimen, body with some white mold.  Additional material examined( 9♂, 18♀).   AUSTRALIA:  New South Wales: Deniliquin:  1.i.1944( 1♀, MVM);  Hay: 1917, W.W. Froggatt( 1♀, ANIC);  Murray River  50 miW of Wentworth,  22.xi.1967( 1♀, MVM);  Niemur River: Near Wakool,  28.xii.1954( 1♂, MVM);   Northern Territory: Alice Springs: Stuart Highwayroadhouse rest area,  19.xi.1989, R.B. Miller( 1♂, FSCA);   Queensland: Brisbane: ( 3♀, QMBA);  Dalby:  3.ii.1932( 4♂, 5♀, QMBA);  Roma:  30.i.1932( 2♂, 3♀, QMBA);   30.ii.1930, Franzenleg. ( 1♀, QMBA);  Coll. Esben-Petersen( 1♀, ZMUC);  Stradbroke Island:  xii.1912( 1♀, ANIC);  White MountainNatural Park: Julia Creek,  xii.1998, R.B. Miller, in tree along river (in tree hole) ( 1♂, FSCA);   Victoria: Yarrawonga:  4.i.1948( 1♀, MVM).   Comments.See Comments for  Glenoleon. 1915-12-20 ANIC W. W. Froggatt & Coll. Froggatt Australia From Hay 165 166 2 2 New South Wales holotype 1944-01-01 MVM Australia Deniliquin 165 166 1 1 New South Wales [248,699,657,683] ANIC W. W. Froggatt Australia Hay 165 166 1 1 New South Wales [712,1432,656,683] 1967-11-22 MVM Australia Murray River 165 166 1 1 New South Wales [151,764,692,719] 1954-12-28 MVM Australia Near Wakool Niemur River 165 166 1 1 New South Wales 1989-11-19 FSCA R. B. Miller Australia Stuart Highway Alice Springs 165 166 1 1 Northern Territory [756,1183,728,755] QMBA Australia Brisbane 165 166 3 3 Queensland 1932-02-03 QMBA Australia Dalby 165 166 9 5 4 Queensland [306,701,765,791] 1932-01-30 QMBA Australia Roma 165 166 5 3 2 Queensland [711,1135,765,791] 1930-02-30 QMBA Franzen Australia 165 166 1 1 Queensland 1930-02-30 ZMUC Coll. Esben-Petersen Australia true 165 166 1 1 Queensland [255,703,801,827] 1912-12 ANIC Australia Stradbroke Island 165 166 1 1 Queensland 1998-12 FSCA R. B. Miller Australia Julia Creek White Mountain 165 166 1 1 Queensland [662,1163,836,863] 1948-01-04 MVM Australia Yarrawonga 165 166 1 1 Victoria