Antatelson antennatum Bellan­Santini & Ledoyer 1974 Probolisca nasutigenes Antatelson antennatum Probolisca nasutigenes A. antennatum Metopa nasutigenes Ausatelson Prothaumatelson Schellenberg, 1931 Prothaumatelson nasutum ( Chevreux, 1912 ) Thaumatelson nasutum Thaumatelson inermis Thaumatelson nasutum Prothaumatelson nasutum Ausatelson J.L. Barnard (1972a, 312) Thaumatelsonine Stenothoids (Crustacea, Amphipoda). Part 2 Krapp-Schickel, Traudl Zootaxa 2006 2006-04-03 1165 1 31 66Y3R Bellan-Santini & Ledoyer Bellan-Santini & Ledoyer 1974 [264,877,443,470] Malacostraca Stenothoidae Antatelson GBIF,CoL Animalia Amphipoda 16 17 Arthropoda species antennatum    Antatelson antennatum Bellan­Santini & Ledoyer 1974, 701 fig. 39   Material examinedUL/OC­458 King George Island, Admiralty Bay, 46 m, 17.3.1985, 3 ind., 3 mm.   Diagnosis Antenna 1 article 1 nasiform process reaching half of article 3. Gnathopods 1, 2 both palmar corner rounded, hind margins parallel to front margins; gnathopod 1, 2 propodi twice as long as large. Pereonite 4 widened, without any process. Uropods slightly surpassing length of telson.   Remarks    Probolisca nasutigenesand  Antatelson antennatumare strikingly similar and both described from Kerguelen, thus I argued that some details may have been overlooked or not observed sufficiently, e.g. the telson of  Probolisca nasutigenesis described as ordinary, not "twisted", but Stebbing mentioned "upturned margins" ( Stebbing 1906, 183, not in the original description).­ But  A. antennatum( 2mmlength) is described as living among red algae ( Bellan­Santini & Ledoyer 1974, 641, see K­80, K­81, K­83, 50 m depth), the here studied material from King George Island ( 3 mm) comes from medium to coarse sand and shell, while Stebbing’s  Metopa nasutigenes( 4.5 mm) is found in 222 mdepth among volcanic mud (also Bellan­Santini & Ledoyer 1974found the latter species with 1 specimenat 50 mdepth in the Kerguelen). ­ Thus the synonymy of these two "nosebearing", extremely similar species could not be attested. In the material from King George Island uropod 1 is longer than drawn in Bellan­ Santini, 1974and thus matching with Stebbing, 1888, also uropod 3 article 2 seems longer, but the articulation is not very clear (the same also in  Ausatelson).   Distribution Antarctica, Kerguelen, King George Island.   EcologyRed algae, medium to coarse sand and shell, mud. About 50 mdepth.    Verticotelson gen. nov. Type species.  Verticotelson mantis n. sp.Monotypic.  Characters Antenna 1 peduncle article 1 longer than first body segment, with long nasiform process; accessory flagellum absent. Mandible palp lacking. Maxilla 1 palp with 2 articles. Maxilla 2 plates in tandem position, subequal. Maxilliped outer plates lacking, palp slender and narrow. Coxa 4 trapezium­shaped. Gnathopod 1 propodus long and narrow, subrectangular, subchelate; gnathopod 2 with propodus, carpus and merus extremely long and narrow, dactylus minute. Urosome segments fused. Telson thickened, fleshy, lateral area larger than dorso­ventral one, thus kept in vertical position.   EtymologyFor stressing the rudder­like telson with large lateral area.    Verticotelson mantis n.sp.  Fig. 7–10   Material examined   Holotype. Australia, Victoria, Western Port( 38 o26.48’S, 145 o13.03’E),  23 m, Sand, Smith­McIntyre grab, Coleman, N,  25 November 1973(stn WBES 1748), NMV J39602( 1 male, 2.5 mm).   Allotype. Same locality as holotype, NMV J52748( 1 female, 2.5 mm).   Paratypes. Samelocality as holotype NMV J39600(1 ind. 2.5 mm); ( 38 o24.36’S, 145 o14.66’E),   20 m, Sand, Smith­McIntyre grab, Colemancoll, N,  21 November 1973(stn WBES 1729), NMV J39601(2). Other material. Australia, Victoria, Western Port ( 38 o29.78’S, 145 o6.28’E), 24 m, Sand, Smith­McIntyre grab, Coleman, N, 25 November 1973(stn WBES 1746), NMV J39603(juvenile 1.2 mm); Off Crib Point ( 38 o22.06’S, 145 o14.10’E),  13 m, reef songe, Smith­McIntyre grab, Gilmour, AJ, Marine Studies Group, Melita, FV,  5 March 1965(stn CPBS­S 33), NMV J39604( 1 juvenile).   Diagnosis Antenna 1 peduncle article 1 huge, nasiform process as long as article, rest of antenna continuing in a right angle. Eyes rounded. Gnathopod 2 characteristically lengthened with very long carpus and reduced dactylus. Pereopods feeble, scarcely beset with setae. Uropods slim and naked. Telson elevated.   Description Length, 1.5–2.5mm Antenna 1 <antenna 2; article 1 with nasiform process, longer than first body segment, front end somewhat serrated and beset with single setae. Insertion of article 2 medioventrally on article 1, creating a right angle. Accessory flagellum lacking. Article 3 subequal article 2, flagellum with 3 segments. Antenna 2 article 5> article 4, flagellum with 8 articles. Mandibular incisor with sharp teeth, lacinia mobilis like a scraper with many short teeth; all other parts smooth and reduced, palp lacking. Maxilla 1 inner plate lacking seta, round, outer plate with 6 robust setae and some subapical setae. Maxilla 2 inner plate somewhat shorter and broader than outer, both rounded and in tandem position, short setae distally. Maxilliped inner plates small, separated, outer plates lacking, palp narrow, with 4 articles, the second one on outer margin densely beset with setae. Coxa 1 subquadrate, minute. Coxa 2, 3 rectangular, length more than twice of width; coxa 4 trapezoidal to rectangular, width> length. Coxa 5–7 subequal, minute, even shorter than coxa 1. Gnathopod 1 basis ratio length to width = 5; ischium with few long setae on outer margin, merus and carpus with many fine setae on outer margin; carpus triangular, length subequal to propodus; propodus rectangular, length> 3 times width, on outer margin 3 combed robust setae, hind margin finely serrated; dactylus curved, palm margin well defined by corner and robust seta. Gnathopod 2 strongly aberrant having a strong long basis with length 4.5 times the width, and propodus + carpus extremely thin, carpus length: width = 12, propodus length: width = 7. The dactylus is reduced, distal part of propodus with a transversally "striped" area which seems to have a special, but hitherto unknown function. Pereopods like in most of thaumatelsonids feeble and with only few setae. Urosomites fused. Uropods extending less far posteriorly in succession, even in uropod 1 lower ramus> the upper one, which is longer than uropod 2 lower ramus> upper one> uropod 3> tip of telson. Telson insertion hidden under end of urosomite 1; there was no fusion with urosomite 3 visible; distally rounded, very short medial setae.   Etymology Mantis, ­eos is the Greek expression for prophet, soothsayer; the situation of the folded long gnathopod 2 reminds of praying arms like in Mantis religiosa.Used as noun in apposition.   Distribution Australia, Victoria, off Melbourne (Phillip Island, Western Port).   EcologyFine sand, sponges, 13–24mdepth. FIGUREgnathopod 2; d) Gn2 distally enlarged; Gn1, Gn 2 inscale x of Fig. 2= 0.2mm; Gn1 enlarged in scale x = 0.1mm; Gn2 enlarged in scale x = 0.04 mm; e) peraeopod 3; f) peraeopod 4; g) peraeopod 5; h) peraeopod 6; i) peraeopod 7; P3,4 in scale x = 0.4 mm; P5,6,7 in scale y of Fig. 2= 0.1mm; k) pleon; l) urosomite 1; m) telson; n) uropod 1; o) uropod 2; p) uropod 3; k,l,m in scale y = 0.1mm; below l­p in scale y = 0.4mm.     Prothaumatelson Schellenberg, 1931  Schellenberg, 1931, p. 113  Type species.  Prothaumatelson nasutumChevreux.  Characters Antenna 1 article 1 nasiform; accessory flagellum absent. Mandible palp with 1 long article; maxilla 1 palp with 2 articles; maxilla 2 ordinary. Gnathopod 1 and 2 very different in size and shape, gnathopod 1 small, subchelate­rectipalmate, merus incipiently chelate; gnathopod 2 enlarged, propodochelate. Pereopod 3–7 basis rectilinear. Pereonite 4 elongate, tumid. Pleonite 3 lacking process. Urosomite 1 folds weakly vaulting over urosomite 2. Urosomites 2+3 coalesced. Telson huge, vertically elevated, laterally compress, lateral surface area equal to lateral area of urosome.     Prothaumatelson nasutum( Chevreux, 1912)  Chevreux, 1912, 5, 212 (  Thaumatelson nasutum). Chilton, 1912, 201, 483, t. i, fig. 16, 17 (  Thaumatelson inermis). Chevreux, 1912, 109– 113, fig. 16–18 (  Thaumatelson nasutum). Schellenberg, 1931, 113 (  Prothaumatelson nasutum). Barnard, K.H. 1932, 112–113. Thurston, 1974, 25–26.   Diagnosis A1 article 1 strongly nasiform; Md palp well developed, no articulation visible; gna­ thopod 1 subchelate­rectipalmate, gnathopod 2 propodochelate, dactylus ca 40% propo­ dus. Urosomite 1 has a short, but clearly visible fold over the following segment folding dorsally, reaching, but not overlapping insertion of telson. Uropod 3 shorter than telson, scarcely surpassing peduncle of uropod 2. Telson vertically inserted and huge, lance­ or blade­shaped, with a sclerotified dorsal crest.   Material examined  Type materialSIW, 1 slide, 1 ind. in alcohol, Inv. Nr. 195166, 4 ind. in alcohol, Inv. Nr. 195167. Other material. SIW, 1 ind. alcohol, Inv. Nr. 149531. MNHUB Grytviken, S­ Georgia, Nr. 22805 18 ind. in alcohol (leg. + det. Schellenberg). ­ BML, many ind. in alcohol, Inv. Nr. 1969.143–147 Signy Isl., S­Orkneys (leg. + det. Thurston), 143, 1 ind., 144, 1 female( 2 eggs) 2mm; 145, 1 ind. 2.2mm; 146, 2 ind. 1.2 mm; 147 1 fem. ( 1 egg) 2mm. UL/OC­ 590 King George Isl., Admiralty Bay, 25.1.1988 2 ind. leg. Jazdzewski. UL/28 King George Isl., Admiralty Bay, from rhizoids of  Phyllogigas grandifolius  30–60m,  24.1. 1980.   DistributionSubantarctic and near islands.   EcologyIn large algae, probably in the epizoic growth on the holdfasts, 5–15 m( Thurston 1974, 25); 1– 60m.    AusatelsonJ.L. Barnard, 1972    Ausatelson J.L. Barnard (1972a, 312)  Type species.  Ausatelson ule L.J. Barnard, 1972a.  Characters Antenna 1 nasiform process on article 1. Accessory flagellum minute to absent. Palp of mandible 1­articulate; palp of first maxilla 2­articulate. Inner plate of maxilla 2 obsolescent. Inner plates of maxillipeds partially fused. Gnathopod 1 small, almost simple, palm oblique and longer than posterior margin of propodus; merus not lengthened. Gnathopod 2 enlarged, palm weakly oblique, merus short, lobed. Pereonite 4 highly elongate and tumid. Pleonites 5–6 coalesced; pleonite 3 dorsally tumid but lacking process; pleonite 4 weakly extended posterodorsally. Telson huge, laterally compressed, lateral surface area equal to lateral area of urosome.   Remarks This genus has closest relationship to  Antatelson(see humps on pleonite 3), but gnathopods are here much more different in size and shape from each other, mandible palp has one article (vs. 2?– 3 in  Antatelson) and pereonite 4 is tumid (vs. smooth in  Antatelson). 2236572474 1973-11-25 NMV Coleman, N Australia 23 -38.441334 Sand 11 145.21716 Western Port 17 18 WBES 1748 1 1 Victoria holotype 2236572425 [312,1146,1228,1254] NMV 17 18 1 1 allotype 2236572443 NMV Same -38.406 11 145.24434 17 18 2 paratype 2236572449 1973-11-21 NMV Sand & Coleman 20 17 18 WBES 1729 1 paratype 2236572463 AJ 13 -38.36767 11 145.235 17 18 1 2236572446 1965-03-05 NMV Melita, FV 17 18 CPBS-S 33 1 1 2236572465 1980-01-24 UL Jazdzewski. 45 King George Isl. Admiralty Bay 23 24 1