Phylloicus
Flint, 1991:99
Phylloicus munozi
P. blahniki
Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae)
PRATHER, AYSHA L.
Zootaxa
2003
2003-08-29
275
1
1
214
Prather, 2003
Prather
2003
[264,494,964,990]
Insecta
Calamoceratidae
Phylloicus
Animalia
Trichoptera
74
75
Arthropoda
species
munozi
sp. nov.
Figs. 74, 75
Phylloicus“species 2" Flint, 1991:99. Phylloicus munoziis similar to P. blahniki, but is distinguished by the unmodified tergum III, the absence of posterior processes of tergum IV and a complex lateral sclerite ( Fig. 74F), and the long digitate basodorsal process of tergum X ( Fig. 74A, B).
Adult. Forewing length 9.710.5 mm, n = 17. Head golden brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax golden brown; ventrolateral thorax pale tan. Legs pale tan. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band pale orangeyellow, reaching posterior wing margin, at least 1/2 width of wing, narrow, less than 10 setae wide; distal band pale orangeyellow, not reaching either wing margin, 1/2 width of wing or less, narrow, less than 10 setae wide; with two basal stripes, ivory; cell Cu and base of cell Cu 2clear. Hind wing basal brush present in male. Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired lateral sclerites, mesal coremata and lateral coremata; lateral sclerite spatulate, directed laterally and with acute basomesal process; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata singlelobed, setose, capable of inflation beyond terminalia. Tergum V without sclerotized modifications ( Fig. 74F). Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX without mesal ridge; posterior margin continuous with basodorsal process of tergum X ( Fig. 74B); lateral ridge present; dorsal pleural setae approximately 2, ventral pleural setae absent ( Fig. 74A); sternum IX without mesolateral ridges; sternum IX ( Fig. 74C). Preanal appendage approximately length of tergum X, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage ( Fig. 74A, B). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, entire ( Fig. 74A, B). Harpago slightly tapered; peglike setae many, apical ( Fig. 74A, C). Phallic endotheca with paired apicolateral lobes and paired basolateral lobes, basolateral lobes tapered apically, apicolateral lobes large and rounded; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoeshaped ( Fig. 74D, E). Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII posterior margin entire; sternum VIII ( Fig. 75C). Tergum IX without mesal ridge ( Fig. 75B). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 75A). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 75B). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 75A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular ( Fig. 75A).
Holotypemale: COSTA RICA: Cartago: Reserva Tapantí, Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12”N, 83°46'48"W, 1400 m, 12.viii.1990, Holzenthal, Blahnik, & Muñoz ( UMSP). Paratypes: COLOMBIA: Antioquia: Mun. Envigado, Quebrada La Ayura, 1750 m, 2.xii.1983, Matthias— 1 female( NMNH); site B, 1750 m, 9.xii.1983, Matthias — 2 females( NMNH); trap B, 1750 m, 20.iv.1983, Matthias — 1 male( NMNH); 23.viii.1983, Matthias — 2 males( NMNH); COSTA RICA: Límon:Limon, 16 kmW Guapiles, 400 m, Ii. Mar 1989, Hanson — 1 male( UMSP); Puntarenas: Río Bellavistatrib., Las Alturas, road to quarry, 08°57'07”N, 82°50'53"W, 1480 m, 19.iii.1991, Holzenthal, Muñoz, & Huisman — 1 male( INBIO); — 1 male( UMSP); Río Jaba, rock quarry, 1.4 km (air) W Las Cruces, 08°47'24”N, 82°58'12"W, 1150 m, 9.viii.1990, Holzenthal, Blahnik, & Muñoz — 1 male( UMSP); 15.iii.1991, Holzenthal, Muñoz, & Huisman — 2 males( UMSP); Jardín Botanico R & C Wilson, unnamed trib., Sendro del Agua, 08°48'00”N, 82°57'36"W, 1180 m, 8.viii.1990, Holzenthal, Blahnik, & Muñoz — 2 males( UMSP); San José: Reserva Biológica Carara, Río del Sur, 1.5 km (rd) S Carara, 09°46'08”N, 84°31'52"W, 160 m, 13.iii.1991, Holzenthal, Muñoz, & Huisman — 1 female( UMSP); PANAMA: Chiriqui: Fortuna Dam Sitenr. Hornitos, 08°55'00”N, 82°16'00"W, 1050 m, 4.i. 7.iii.1978, Wolda — 1 male( NMNH). Distribution. Colombia, Costa Rica, Panama. Etymology. This species is named for Fernando MuñozQuesada, who collected the typespecimen.
3319300443
1990-08-01
1990-08-02
1990-08-01
UMSP
Costa Rica
1400
9.719999
Quebrada Palmitos
21
-83.78001
Reserva Tapanti
75
76
1
Cartago
holotype
3319300822
1983-12-02
NMNH
Colombia
Mun. Envigado
1750
Matthias
Quebrada La Ayura
75
76
1
1
Antioquia
paratype
3319301023
1983-12-09
NMNH
Colombia
1750
75
76
2
2
Antioquia
paratype
3319300919
[488,1161,1364,1390]
1983-04-20
NMNH
Colombia
1750
75
76
1
1
Antioquia
paratype
3319300672
1983-08-23
NMNH
Colombia
75
76
2
2
Antioquia
paratype
3319300977
1989-03
UMSP
Costa Rica
400
16 km W Guapiles
75
76
1
1
Limon
paratype
3319300826
1991-03-19
INBIO
Las Alturas
Costa Rica
1480
8.951944
Rio Bellavista
21
-82.84806
75
76
1
1
Puntarenas
paratype
3319300403
[659,852,1524,1550]
1991-03-19
UMSP
Las Alturas
Costa Rica
1480
true
8.951944
Rio Bellavista
21
-82.84806
75
76
1
1
Puntarenas
paratype
3319300355
1990-08-09
UMSP
W Las Cruces
Costa Rica
1150
8.790001
Rio Jaba
21
-82.97
75
76
1
1
Puntarenas
paratype
3319300602
[519,1317,1604,1630]
1991-03-15
UMSP
Costa Rica
75
76
2
2
Puntarenas
paratype
3319300998
1990-08-08
UMSP
Holzenthal, Blahnik, & Muñoz
Costa Rica
1180
8.8
Sendro del Agua
21
-82.96
75
76
2
2
Puntarenas
paratype
3319301030
1991-03-13
UMSP
S Carara
Costa Rica
160
9.768888
Rio del Sur
21
-84.53111
Reserva Biologica Carara
75
76
1
1
San Jose
paratype
3319300536
1978-01-04
1978-03-07
1978-01-04
NMNH
Fortuna Dam Site & Hornitos
Panama
1050
8.916667
Chiriqui
21
-82.26667
75
76
1
1
Chiriqui
paratype