Phylloicus Flint, 1991:99 Phylloicus munozi P. blahniki Revision of the Neotropical caddisfly genus Phylloicus (Trichoptera: Calamoceratidae) PRATHER, AYSHA L. Zootaxa 2003 2003-08-29 275 1 1 214 Prather, 2003 Prather 2003 [264,494,964,990] Insecta Calamoceratidae Phylloicus Animalia Trichoptera 74 75 Arthropoda species munozi sp. nov.   Figs. 74, 75     Phylloicus“species 2"  Flint, 1991:99.   Phylloicus munoziis similar to  P. blahniki, but is distinguished by the unmodified tergum III, the absence of posterior processes of tergum IV and a complex lateral sclerite ( Fig. 74F), and the long digitate basodorsal process of tergum X ( Fig. 74A, B).  Adult. Forewing length 9.7­10.5 mm, n = 17. Head golden brown. Maxillary palps dark brown. Antenna twice forewing length; dark brown, with narrow patches of pale sensilla on anteromesal surface of each flagellomere. Dorsal pterothorax golden brown; ventrolateral thorax pale tan. Legs pale tan. Metathoracic leg of male with posterior fringe of long setae, setae dark. Tibial spur formula 2,4,4. Forewing flat; dark brown; with two transverse bands; proximal band pale orange­yellow, reaching posterior wing margin, at least 1/2 width of wing, narrow, less than 10 setae wide; distal band pale orange­yellow, not reaching either wing margin, 1/2 width of wing or less, narrow, less than 10 setae wide; with two basal stripes, ivory; cell Cu and base of cell Cu 2clear. Hind wing basal brush present in male. Male. Preterminalic abdominal terga with anteromesal notch. Corematic structures present. Tergum IV with paired lateral sclerites, mesal coremata and lateral coremata; lateral sclerite spatulate, directed laterally and with acute basomesal process; lateral coremata with basal globose lobes and long tubular posterior lobe; mesal coremata single­lobed, setose, capable of inflation beyond terminalia. Tergum V without sclerotized modifications ( Fig. 74F). Sternum VII with short, acute anteromesal process. Sternum VIII similar to anterior sterna, sternum IX not elongate. Tergum IX without mesal ridge; posterior margin continuous with basodorsal process of tergum X ( Fig. 74B); lateral ridge present; dorsal pleural setae approximately 2, ventral pleural setae absent ( Fig. 74A); sternum IX without mesolateral ridges; sternum IX ( Fig. 74C). Preanal appendage approximately length of tergum X, of uniform diameter throughout length, setae long, but not filamentous or longer than appendage ( Fig. 74A, B). Tergum X without basal lobes; basodorsal process long and digitate; basolateral processes absent; apex, in lateral view, truncate, in dorsal view, entire ( Fig. 74A, B). Harpago slightly tapered; peglike setae many, apical ( Fig. 74A, C). Phallic endotheca with paired apicolateral lobes and paired basolateral lobes, basolateral lobes tapered apically, apicolateral lobes large and rounded; phallotremal sclerites average size, longest dimension less than diameter of phallobase; dorsal sclerite ovoid, in dorsal view horseshoe­shaped ( Fig. 74D, E). Female. Preterminal abdominal terga with anteromesal notch. Sternum VII with short pointed anteromesal process. Tergum VIII without posterolateral brush; sternum VIII posterior margin entire; sternum VIII ( Fig. 75C). Tergum IX without mesal ridge ( Fig. 75B). Sternum IX anterior and posterior lobes darkly sclerotized and striate, with patch of lightly sclerotized cuticle lateral to vaginal opening ( Fig. 75A). Tergum X appendage shorter than mesal lobe, base marked by faint suture line, apex oblique; mesal lobe lightly sclerotized; digitate lateral processes absent ( Fig. 75B). Sternum X with patches of short fine setae posterolaterally to anal opening ( Fig. 75A). Vaginal apparatus anterior and posterior sclerites equal in length; anterior sclerite rounded anteriorly, posterolateral projections absent; posterior sclerite triangular ( Fig. 75A).     Holotypemale: COSTA RICA: Cartago: Reserva Tapantí, Quebrada Palmitos& falls, ca. 9 km(road) NW tunnel, 09°43'12”N, 83°46'48"W,  1400 m,  1­2.viii.1990, Holzenthal, Blahnik, & Muñoz ( UMSP).    Paratypes: COLOMBIA: Antioquia: Mun. Envigado, Quebrada La Ayura,  1750 m,  2.xii.1983, Matthias— 1 female( NMNH); site B,  1750 m,  9.xii.1983, Matthias — 2 females( NMNH); trap B,  1750 m,  20.iv.1983, Matthias — 1 male( NMNH);   23.viii.1983, Matthias — 2 males( NMNH);   COSTA RICA: Límon:Limon,  16 kmW Guapiles,  400 m, Ii.­  Mar 1989, Hanson — 1 male( UMSP);   Puntarenas: Río Bellavistatrib., Las Alturas, road to quarry, 08°57'07”N, 82°50'53"W,  1480 m,  19.iii.1991, Holzenthal, Muñoz, & Huisman — 1 male( INBIO); —  1 male( UMSP);  Río Jaba, rock quarry, 1.4 km (air) W Las Cruces, 08°47'24”N, 82°58'12"W,  1150 m,  9.viii.1990, Holzenthal, Blahnik, & Muñoz — 1 male( UMSP);   15.iii.1991, Holzenthal, Muñoz, & Huisman — 2 males( UMSP);  Jardín Botanico R & C Wilson, unnamed trib., Sendro del Agua, 08°48'00”N, 82°57'36"W,  1180 m,  8.viii.1990, Holzenthal, Blahnik, & Muñoz — 2 males( UMSP);   San José: Reserva Biológica Carara, Río del Sur, 1.5 km (rd) S Carara, 09°46'08”N, 84°31'52"W,  160 m,  13.iii.1991, Holzenthal, Muñoz, & Huisman — 1 female( UMSP);   PANAMA: Chiriqui: Fortuna Dam Sitenr. Hornitos, 08°55'00”N, 82°16'00"W,  1050 m,  4.i.­ 7.iii.1978, Wolda — 1 male( NMNH). Distribution. Colombia, Costa Rica, Panama. Etymology. This species is named for Fernando Muñoz­Quesada, who collected the typespecimen. 3319300443 1990-08-01 1990-08-02 1990-08-01 UMSP Costa Rica 1400 9.719999 Quebrada Palmitos 21 -83.78001 Reserva Tapanti 75 76 1 Cartago holotype 3319300822 1983-12-02 NMNH Colombia Mun. Envigado 1750 Matthias Quebrada La Ayura 75 76 1 1 Antioquia paratype 3319301023 1983-12-09 NMNH Colombia 1750 75 76 2 2 Antioquia paratype 3319300919 [488,1161,1364,1390] 1983-04-20 NMNH Colombia 1750 75 76 1 1 Antioquia paratype 3319300672 1983-08-23 NMNH Colombia 75 76 2 2 Antioquia paratype 3319300977 1989-03 UMSP Costa Rica 400 16 km W Guapiles 75 76 1 1 Limon paratype 3319300826 1991-03-19 INBIO Las Alturas Costa Rica 1480 8.951944 Rio Bellavista 21 -82.84806 75 76 1 1 Puntarenas paratype 3319300403 [659,852,1524,1550] 1991-03-19 UMSP Las Alturas Costa Rica 1480 true 8.951944 Rio Bellavista 21 -82.84806 75 76 1 1 Puntarenas paratype 3319300355 1990-08-09 UMSP W Las Cruces Costa Rica 1150 8.790001 Rio Jaba 21 -82.97 75 76 1 1 Puntarenas paratype 3319300602 [519,1317,1604,1630] 1991-03-15 UMSP Costa Rica 75 76 2 2 Puntarenas paratype 3319300998 1990-08-08 UMSP Holzenthal, Blahnik, & Muñoz Costa Rica 1180 8.8 Sendro del Agua 21 -82.96 75 76 2 2 Puntarenas paratype 3319301030 1991-03-13 UMSP S Carara Costa Rica 160 9.768888 Rio del Sur 21 -84.53111 Reserva Biologica Carara 75 76 1 1 San Jose paratype 3319300536 1978-01-04 1978-03-07 1978-01-04 NMNH Fortuna Dam Site & Hornitos Panama 1050 8.916667 Chiriqui 21 -82.26667 75 76 1 1 Chiriqui paratype