Revision of the Exechia parva group (Diptera: Mycetophilidae) Lindemann, Jon Peder Soli, Geir Kjaerandsen, Jostein Biodiversity Data Journal 2021 2021-09-24 9 67134 67134 urn:lsid:zoobank.org:act:05e60b65-31c0-4583-8580-ff712f5aa0a3 Lindemann Lindemann 2021 Insecta Mycetophilidae Exechia CoL Animalia Exechia spatulata Diptera 0 67134 Arthropoda species spatulata sp. n.  Materials   Type status:  Holotype.  Occurrence:catalogNumber: TSZD-JKJ-111444; recordedBy:  Jarl Birkeland; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned;  Location:country: Norway; county: Rogaland; municipality: Sokndal; locality: Skittmyr; decimalLatitude: 58.350488; decimalLongitude: 6.3054945;  Event:samplingProtocol: Malaise trap; eventDate: 2020-05-21/ 2020-06-11; Record Level:collectionID: TMU-JKJ-COL-001514; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-103590; occurrenceRemarks: TMU-JKJ fieldwork; recordedBy:  Jostein Kjaerandsen& Martin T. Dahl; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Norway; county: Finnmark; municipality:  Sor-Varanger; locality:   Gjokbekken, Pasvik; verbatimElevation:   72 m; locationRemarks: SLAM 1; decimalLatitude: 69.1522; decimalLongitude: 29.1516;  Event:samplingProtocol: SLAM trap; eventDate: 2017-08-25; Record Level:collectionID: TMU-JKJ-COL-000538; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-107937; occurrenceRemarks: ADB - M&M project; recordedBy:  Kai Berggren; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Norway; county: Vest-Agder; municipality: Kristiansand; locality: Nedre Jegersbergvann; verbatimElevation:   21 m; locationRemarks: MT 3, at lake; decimalLatitude: 58.169; decimalLongitude: 8;  Event:samplingProtocol: Malaise trap; eventDate: 2019-06-29; Record Level:collectionID: TMU-JKJ-COL-001099; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-107987; occurrenceRemarks: Giftfrom collector; recordedBy: J. Birkeland; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Norway; county: Rogaland; municipality: Sokndal; locality: Skittmyr; verbatimElevation:   22 m; decimalLatitude: 58.351; decimalLongitude: 6.306;  Event:samplingProtocol: Malaise trap; eventDate: 2019-10-02; Record Level:collectionID: TMU-JKJ-COL-001079; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-107932; occurrenceRemarks: Depth: 7.5 | ADB - M&M project; recordedBy: K. Berggren; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Norway; county: Vest-Agder; municipality: Kristiansand; locality: Nedre Jegersbergvann; verbatimElevation:   21 m; locationRemarks: MT 3, at lake; decimalLatitude: 58.169; decimalLongitude: 8;  Event:samplingProtocol: Malaise trap; eventDate: 2019-08-28; Record Level:collectionID: TMU-JKJ-COL-001101; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-111547; recordedBy: H. Andersson; individualCount: 1; sex: male; lifeStage: adult; preparations: Slide;  Location:country: Sweden; county: Norrbotn; municipality:  Lulea; locality:   Ranea,  Hoegsoen,  Hoegsoefjaerden;  Event:eventDate: 1965-09-06;  Record Level:institutionCode: TMU   Typestatus:  Paratype.  Occurrence:catalogNumber: SPM-060717; recordedBy: M. Soederlund; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Sweden; county:  Vaestra Goetaland; municipality: Ale; locality:  Kollandamosse; locationRemarks: MT 2; decimalLatitude: 57.940833; decimalLongitude: 12.249444;  Event:samplingProtocol: Malaise trap; eventDate: 1995-09-11/ 1995-09-20; Record Level:institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: SPM-060720; recordedBy: M. Soederlund; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Sweden; county:  Vaestra Goetaland; municipality: Ale; locality:  Kollandamosse; locationRemarks: MT 2; decimalLatitude: 57.940833; decimalLongitude: 12.249444;  Event:samplingProtocol: Malaise trap; eventDate: 1995-10-01/ 1995-10-10; Record Level:collectionID: COL-009616; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: SPM-036986; recordedBy: G. Nilsson; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned, with genitalia in glycerine in separate microvial;  Location:country: Sweden; county:  Vaestmanland; municipality:  Vaesteras; locality:  Solbacken; decimalLatitude: 59.517222; decimalLongitude: 16.740000;  Event:samplingProtocol: Malaise trap; eventDate: 1990-08-25/ 1990-09-09; Record Level:collectionID: COL-008186; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-111490; recordedBy: J. Birkeland; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned;  Location:country: Norway; county: Rogaland; municipality: Sokndal; locality:  Skittmyr2; decimalLatitude: 58.3501773; decimalLongitude: 6.3055563;  Event:samplingProtocol: Malaise trap; eventDate: 2020-08-08/ 2020-09-05; Record Level:collectionID: TMU-JKJ-COL-001525; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-111221; recordedBy: J. Kjaerandsen; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned;  Location:country: Norway; county:  Trrndelag; municipality: Oppdal; locality:  Dovre,  Gronbakken; decimalLatitude: 62.278890; decimalLongitude: 9.59833;  Event:samplingProtocol: Sweep net; eventDate: 2020-07-07; Record Level:collectionID: TMU-JKJ-COL-001481; institutionCode: TMU  Type status:  Paratype.  Occurrence:catalogNumber: TSZD-JKJ-111439; recordedBy: J. Birkeland; individualCount: 1; sex: male; lifeStage: adult; preparations: Pinned;  Location:country: Norway; county: Rogaland; municipality: Sokndal; locality:  Skittmyr2; decimalLatitude: 58.3501773; decimalLongitude: 6.3055563;  Event:samplingProtocol: Malaise trap; eventDate: 2020-07-10/ 2020-08-08; Record Level:collectionID: TMU-JKJ-COL-001524; institutionCode: TMU  Description Male (n = 12): Body length 2.9-3.2 mm. Wing length 2.4-2-5 mm. Colouration(Dry specimen). Head, face and clypeus dark brown; labellum dark brown; palpus yellow. Antenna with scape and pedicel yellow; flagellum dark brown, basal part of first flagellomere pale. Thorax with scutum, lateral sclerites and propleura dark brown; halteres whitish-yellow. Legs yellow. Abdomen dark brown. Terminalia brown. Head. Vertex, frons and clypeus covered with pale setae. Antenna 1.78 times as long as length from vertex to ventral margin of clypeus; flagellomeres quadrate, with sixth flagellomere as long as broad. Thorax. Scutum covered with pale setae. Legs(n = 3). Fore leg with tibia as long as first tarsomere. Mid-tibia with 20 anterior, 3 posterodorsal, 4 posterior and 2 posteroventral bristles. Hind tibia with 6 anterodorsal, 5 posterodorsal and 3 posterior bristles. Wings. Vein r-m 2.8 times longer than stem of M-fork. Abdomen.Tergites covered with pale setae. Terminalia(n = 4, Fig. 36 a, b). Each part of divided tergite IX with about 3-4 setae, most apical seta stout. GL with length 0.6-0.75 of gonocoxite width (Fig. 36 a, b); apico-internal margin slightly angled exteriorly; basal third covered with setae; apex with 3 setae. Aedaegal guides short and acute. Hypandrium covered with 10-12 setae, with apical pair reaching two thirds to three-fourths of GL (Fig. 36 a, b). Hypandrial lobe with each part slender, apical half slightly widened. Gonostylus (Fig. 36 c) with DB elongate, about 3.52 times longer than broad, virtually straight or very slightly curved interiorly, apical lobe distinctly spathulate, apex rounded; dorsal side evenly covered with setae, except on most apical part; external margin almost straight, not forming any angle, with row of 4 elongate setae. VB with 2 small setae. IB apically with 1 seta close to apex and pair of setae one-third from apex. MB apex acute, with 1 seta close to the base. Female: Unknown.  Diagnosis Distinguished from  E. longilobatain having the dorsal gonostylus branch apically spathulate and about 3.5 times longer than broad (Fig. 36 c), the hypandrium less setose with 10-12 setae (Fig. 36 a, b), each part of the hypandrial lobe with apical half only slightly widened and the gonocoxal lobe length 0.6-0.75 the gonocoxite width (Fig. 36 c); from other species in the  E. parvagroup in having the dorsal branch spathulate and more elongated (Fig. 36 c), in combination with the gonocoxal lobes with apico-internal margin slightly angled exteriorly (Fig. 36 a).  Etymology From Latin spatula, flat piece, relating to the the apical shape of the dorsal branch of the gonostylus.  Distribution West Palaearctic, Norway, Sweden (Fig. 15)  Biology Unknown. 2020-05-21 2020-11-06 2020-05-21 Malaise trap Jarl Birkeland Norway Occurrence 58.350487 Pinned 6.3054943 Jarl Birkeland 0 67134 TSZD-JKJ- 111444 1 1 Rogaland Holotype 2020-05-21 2020-11-06 2020-05-21 Malaise trap Jarl Birkeland Norway Occurrence 58.350487 Pinned 6.3054943 Jarl Birkeland TSZD-JKJ- 111444 1 1 Rogaland Holotype 2017-08-25 SLAM trap Occurrence, Jostein Kjaerandsen & Martin T. Dahl, Jostein Kjaerandsen, Martin T. Dahl Norway Pinned 0 69.1522 Sor-Varanger 29.1516 Location 0 67134 TSZD-JKJ- 103590 1 1 Finnmark Paratype 2017-08-25 SLAM trap Occurrence, Jostein Kjaerandsen & Martin T. Dahl, Jostein Kjaerandsen, Martin T. Dahl Norway Pinned 0 69.1522 Sor-Varanger 29.1516 Location TSZD-JKJ- 103590 1 1 Finnmark Paratype 2019-06-29 Malaise trap Kai Berggren Norway Occurrence 0 58.169 Pinned 8.0 Kai Berggren 0 67134 TSZD-JKJ- 107937 1 1 Vest-Agder Paratype 2019-06-29 Malaise trap Kai Berggren Norway Occurrence 0 58.169 Pinned 8.0 Kai Berggren TSZD-JKJ- 107937 1 1 Vest-Agder Paratype 2019-02-10 Malaise trap Occurrence, Gift, J. Birkeland Norway Pinned 0 58.351 Sokndal 6.306 Location 0 67134 TSZD-JKJ- 107987 1 1 Rogaland Paratype 2019-02-10 Malaise trap Occurrence, Gift, J. Birkeland Norway Pinned 0 58.351 Sokndal 6.306 Location TSZD-JKJ- 107987 1 1 Rogaland Paratype 2019-08-28 Malaise trap Occurrence, Depth, K. Berggren Norway Pinned 0 58.169 Kristiansand 8.0 Location 0 67134 TSZD-JKJ- 107932 1 1 Vest-Agder Paratype 2019-08-28 Malaise trap Occurrence, Depth, K. Berggren Norway Pinned 0 58.169 Kristiansand 8.0 Location TSZD-JKJ- 107932 1 1 Vest-Agder Paratype 1965-06-09 Occurrence, H. Andersson Sweden Slide Record Level 0 67134 TSZD-JKJ- 111547 1 1 Paratype 1965-06-09 Occurrence, H. Andersson Sweden Type Slide Record Level TSZD-JKJ- 111547 1 1 Paratype 1995-09-20 1995-11-09 1995-09-20 Malaise trap SPM Occurrence, M. Soederlund Sweden Type 57.940834 Pinned 12.249444 Kollanda 0 67134 SPM- 060717 1 1 Paratype 1995-09-20 1995-11-09 1995-09-20 Malaise trap SPM Occurrence, M. Soederlund Sweden Event 57.940834 Pinned 12.249444 Kollanda SPM- 060717 1 1 Paratype 1995-01-10 1995-10-10 1995-01-10 Malaise trap SPM Occurrence, M. Soederlund Sweden Event 57.940834 Pinned 12.249444 Kollanda 0 67134 SPM- 060720 1 1 Paratype 1995-01-10 1995-10-10 1995-01-10 Malaise trap SPM Occurrence, M. Soederlund Sweden Event 57.940834 Pinned 12.249444 Kollanda SPM- 060720 1 1 Paratype 1990-08-25 1990-09-09 1990-08-25 Malaise trap SPM Occurrence, G. Nilsson Sweden Event 59.517223 Pinned 16.74 Solbacken 0 67134 SPM- 036986 1 1 Paratype 1990-08-25 1990-09-09 1990-08-25 Malaise trap SPM Occurrence, G. Nilsson Sweden Event 59.517223 Pinned 16.74 Solbacken SPM- 036986 1 1 Paratype 2020-05-09 2020-08-08 2020-05-09 Malaise trap Occurrence, J. Birkeland Norway Pinned 58.350178 Sokndal 6.3055563 Location 0 67134 TSZD-JKJ- 111490 1 1 Rogaland Paratype 2020-05-09 2020-08-08 2020-05-09 Malaise trap Occurrence, J. Birkeland Norway Pinned 58.350178 Sokndal 6.3055563 Location TSZD-JKJ- 111490 1 1 Rogaland Paratype 2020-07-07 Sweep net Occurrence, J. Kjaerandsen Norway Event 62.27889 Pinned 9.59833 Gronbakken 0 67134 TSZD-JKJ- 111221 1 1 Paratype 2020-07-07 Sweep net Occurrence, J. Kjaerandsen Norway Event 62.27889 Pinned 9.59833 Gronbakken TSZD-JKJ- 111221 1 1 Paratype 2020-08-08 2020-10-07 2020-08-08 Malaise trap Occurrence, J. Birkeland Norway Pinned 58.350178 Sokndal 6.3055563 Location 0 67134 TSZD-JKJ- 111439 1 1 Rogaland Paratype 2020-08-08 2020-10-07 2020-08-08 Malaise trap Occurrence, J. Birkeland Norway Pinned 58.350178 Sokndal 6.3055563 Location TSZD-JKJ- 111439 1 1 Rogaland Paratype