Tomosvaryella Aczél (Diptera: Pipunculidae) of Colombia, with description of two new species Ramos-Pastrana, Yardany Rafael, José Albertino Zootaxa 2021 2021-06-11 4985 1 37 68 Aczél, 1939 [287,462,151,178] Insecta Pipunculidae Tomosvaryella Animalia Diptera 28 65 Arthropoda species undetermined    1 Hind trochanter with two bare spines ( Fig. 42); phallus with central branch having three stout, reclined, apical spines and one of the lateral branches with one short reclined spine ( Fig. 50)....................................  T. martae  sp. nov.  - Hind trochanter with one distinct protuberance projected or slightly elevated ( Figs 2, 12, 26, 54, 64, 74, 84, 94, 104–108, 122); phallus branches with variable reclined spines, if three spines, then spines medially placed on the branches ( Figs 8, 18, 34, 60, 70, 80, 90, 100, 114, 129).............................................................................. 2   2 Hind trochanter slightly elevated, truncated, without acute protuberance ( Fig. 94); syntergosternite 8 sub-hemispherical ( Fig. 96); right surstylus wider than left, with distinct external submedian lobe ( Figs. 98–99); apex of phallic guide thickened ( Fig. 100); phallus with central branch stout, somewhat coiled ( Fig. 100); ejaculatory apodeme fan-shaped ( Fig. 101)..................................................................................................  T. subvirescens  - Hind trochanter with distinct protuberance ( Figs 2, 12, 26, 42, 54, 64, 74, 84, 104–108), if no distinct protuberance, then with a small elevation having distinct ascendant and descendent margins with obtuse apex at higher point ( Fig. 122); syntergosternite 8 somewhat directed to right side ( Figs 27–28, 43–44); surstyli somewhat similar ( Figs 4–5, 14–15, 30–31, 46–47, 56–57, 66–67, 86–87, 110–111); apex of phallic guide slender ( Figs 50, 60); phallus branches with the same diameter, not coiled ( Figs 8, 18, 34, 50, 60, 70, 80, 90, 114, 129); ejaculatory apodeme generally sandal-shaped ( Figs 9, 19, 35, 51, 61, 71, 91, 115, 130) ................................................................................................... 3   3 Hind trochanter with bulging, drop-shaped protuberance (Fig. 184); both surstyli with external and internal margins somewhat straight ( Fig. 86–87).............................................................................  T. similis  - Hind trochanter different ( Figs 2, 12, 26, 42, 54, 64, 74, 94, 104–108, 122); both surstyli with external and internal margins sinuous ( Figs 4–5, 14–15, 66–67, 110–11, 125–126), in case of doubts, apex distinctly hooked ( Fig. 30–31, 56–57)....... 4   4 Apex of phallic guide distinctly angled from base, without basal spine ( Figs 8, 129)................................ 5  - Apex of phallic guide lined with or slightly above the main body of the structure, with or without basal spine ( Figs 18, 34, 50, 60, 70, 80, 90, 114).................................................................................... 6   5 Hind trochanter with short protruding protuberance with anterior margin somewhat straight and posterior margin descending, yellow-setose ( Fig. 2); surstyli similar in shape ( Figs 4–5); left surstylus with external margin sinuous; right surstylus somewhat straight dorsally in lateral view ( Figs 6–7); phallus with no projection before trifurcation and two branches with small reclined spines ( Fig. 8)............................................................................  T. galapagensis  - Hind trochanter with small triangular protuberance with obtuse apex at highest point, white-setose ( Fig. 122); Surstyli asymmetrical, left surstylus with external margin somewhat straight ( Figs 125–126), right surstylus dorsally distinctly lobate in lateral view ( Fig. 128); phallus with distinct projection before trifurcation and two branches with stout reclined spines ( Fig. 129)........................................................................................  T. venezuelana   6 Hind trochanter with protuberance somewhat squared, apex straight, yellow-setose apically ( Fig. 26)....................................................................................................  T. macarenensis  sp. nov.  - Hind trochanter with protuberance somewhat triangular having angled margins or rectangular with straight margins ( Figs 54, 64, 74, 104–108, 112).................................................................................. 7   7 Hind trochanter with protuberance somewhat rectangular, with margins straight ( Figs 64, 104–108).................... 8  - Hind trochanter with protuberance somewhat triangular with margins angled ( Figs 12, 54, 74)........................ 9   8 Surstyli setulose with high external lateral margins ( Fig. 66–67); surstyli somewhat similar in shape, robust with apexes curved inward, inner and outer margins sinuous ( Fig. 66–67); hind trochanter with long yellow-setulose protuberance at apex ( Fig. 64); apex of phallic guide simple and conical, ventrally serrated ( Fig. 70); central branch of phallus with three stout spines ( Fig. 70)........................................................................................  T. prostata  - Surstyli bare, without high external lateral margins ( Figs 110–111); surstyli similar in shape, robust with apexes curved inward, inner and outer margins sinuous ( Fig. 110–11); hind trochanter with variable stout spiniform protuberance ( Figs 104–108); apex of phallic guide short with a spine ( Fig. 114); phallus with two branches bearing a fine spine ( Fig. 114).......  T. tuberculata   9 Hind trochanter with protuberance entirely white-setose ( Fig. 12); surstylus elongated, somewhat tapered apically, both almost the same length, right surstylus slightly shorter ( Figs 14–15), downward-directed in lateral view ( Figs 16–17); apex of phallic guide short and with a fine spine ( Fig. 18), one branch of phallus with a small spine ( Fig. 18)...................  T. lynchi  - Hind trochanter with protuberance with few spaced setae not entirely covered with white setae ( Fig. 54, 74); surstylus variable as in Figs 57, 77; apex of phallic guide without spine ( Fig. 80), if spine present, then it is stout ( Fig. 60); with central branch of phallus bearing stout spines ( Fig. 80) or row of small spines ( Fig. 60)........................................... 10   10 Surstyli long, distinctly with hooked-tips, without setose basal lobe ( Fig. 56–57); apex of phallic guide with stout spine ( Fig. 60); phallus central branch with row of small spines ( Fig. 60); ejaculatory apodeme sandal-shaped ( Fig. 61).....................................................................................................  T. mexicanensis  - Surstyli robust, indistinctly with hooked-tips, with basal lobe setose ( Fig. 76–77); apex of phallic guide without spine ( Fig. 80); phallus with central branch with three distinct spines ( Fig. 80); ejaculatory apodeme with apex truncated ( Fig. 81)...................................................................................................  T. scopulata