Prachynella lodo J.L. Barnard, 1964: 233 P. oculata Barnard, 1966a: 70 Barnard, 1966b: 26 P. oculata Barnard, 1967: 69 The Pachynidae fam. nov. (Crustacea: Amphipoda: Lysianassoidea) 3246 Lowry, J. K. Stoddart, H. E. Zootaxa 2012 2012-03-23 3246 1 1 69 4M8BT J. L. Barnard, 1964 J. L. Barnard 1964 [151,600,1729,1755] Malacostraca Pakynidae Prachynella Animalia Amphipoda 45 46 Arthropoda species lodo  ( Fig. 34)      Prachynella lodoJ.L. Barnard, 1964: 233(in part, illustrated specimen =  P. oculata  sp. nov.). ―J.L.  Barnard, 1966a: 70(possibly in part). ―J.L.  Barnard, 1966b: 26(in part, part =  P. oculata  sp. nov.). ―J.L.  Barnard, 1967: 69(in part, not figs 29, 30) [see Remarks]. ― Lowry 1984: 72, figs 13–15. ― Austin, 1985: 601. ― Barnard & Karaman, 1991: 520. ―García-  Madrigal, 2007: 108. not  Prachynella lado(sic). ― Hirayama, 1985: 182, figs 175–178 (=  P. shijiki  sp. nov.). not  Prachynella lodo. ― Ishimaru, 1994: 59(=  P. shijiki  sp. nov.).   Material examined.  Holotype, female, 5.8 mm, LACM 57- 210.1, south-east of San Mateo Point, southern California, USA, 33°17’35”N 117°31’00”W,  24 m,  20 February 1957,  Velero IVstation 4868. – 1 female, 7.6 mm, LACM, off Santa Barbara, southern California, station 5261. – about 45 specimens, LACM, southern California, USA,  12– 157 m, 1956–1959, from 40  Velero IVstations (all identified by J.L. Barnardas  P. lodo). –1 ovigerous female, as 1 slide, NMCC 1990-0550, McCauley Point, Vancouver Island, British Columbia, Canada, eastern North Pacific Ocean, [approx. 48°24'N 123°24'W],  49 m,  July 1977, G.W. O'Connell, stn W3-AO.   Description. Headlateral cephalic lobe large, broadly rounded; ventral margin curved. Antenna 1peduncular article 1 with posterodistal spine; accessory flagellum 2-articulate, article 1 short; flagellum with strong 1-field callynophore in female, strong 2-field in male. Mandiblelacinia mobilis a simple robust seta; lamina dentata vestigial; accessory setal row absent; molar absent; palp attached distally; palp article 2 broad, subequal to article 3; article 3 with subparallel margins.  Maxilla 1outer plate with 10 setal-teethin 7/3 arrangement, most outer row setalteeth with medial cusps; palp vestigial, 1-articulate, apical setae absent. Maxillipedinner plate small; outer plate large; palp 3-articulate (articles 1 and 2 fused). Gnathopod 1chelate; ischium smaller than propodus, not swollen posteroproximally; carpus extremely compressed; propodus length about 1.9 × breadth, posterior margin well developed, plam with produced corner, pollex length about 1.2 × breadth, corner of palm with complex robust seta; dactylus reaching to corner of palm.  Gnathopod 2palm minute,obtuse. Pereonite 4without dorsal spines.  Pereonite 5with 1 acute dorsal spine. Pereopod 5merus with posteroproximal shoulder. Pereopod 6basis with posteroventral lobe weakly produced; merus longer than broad, with posteroproximal shoulder. Pereopod 7basis subovate with posterior margin smooth or weakly dentate. Epimeron 3posterior margin without projections, posteroventral corner broadly rounded. Uropod 3biramous, inner ramus extending more than halfway along outer ramus but not reaching base of article 2.  Telsonshorter than broad, weakly emarginate.   Remarks.As pointed out by Lowry (1984: 72), when J.L. Barnard (1964)established the species  P. lodohe described and illustrated one specimenbut designated another specimen as the holotype. Lowry (1984: figs 13–15) illustrated the holotypeof  P. lodoand noted some differences between it and the material illustrated by J.L. Barnard. We have examined a large amount of material from southern Californiaidentified as  P. lodoby J.L. Barnard. This material contains two species:  P. lodoand the species actually illustrated by J.L. Barnard (1964, fig. 7 and 1967, fig. 29) which we establish below as  P. oculata  sp. nov. Although extremely similar,  P. lodoand  P. oculatacan be distinguished by the shape of the dorsal spine on pereonite 5 (posteriorly acute in  P. lodo, rounded in  P. oculata) and by the shape of gnathopod 1 propodus: the pollex (or fixed finger) in  P. lodois considerably stockier than that in  P. oculata, the length-to-breadth ratio being about 1.2 in  P. lodoand 1.8 in  P. oculata. We have re-examined the holotypeof  P. lodoand find that the left mandible has a lacinia mobilis and a vestigial ridge possibly representing the accessory robust setae or a lamina dentata. The single specimen from Vancouver Island also has a poorly developed ridge on the left mandible. J.L. Barnard (1967)recorded one specimenfrom Cedros Trench station 7234 as an “anoculate form” of  P. lodo, and illustrated some parts in fig. 30a–g. The vial now contains only a heavily-dissected carcass and no loose parts. The body is intact but there are no mouthparts, no right-side limbs and no left pereopods below the coxae. There is definitely no dorsal spine on pereonite 4 or 5. We cannot place this specimen, but can say that it is not  P. lodoor  P. oculata.   Distribution.Eastern Pacific Ocean: Vancouver Island, British Columbia, Canada; southern California, USA; Baja California, Mexico. 1957-02-20 1977-07 1957-02-20 LACM Velero IV & J. L. Barnard & G. W. O'Connell United States of America eastern North Pacific Ocean 24 48.4 San Mateo Point 1110 -123.4 Vancouver Island 48 49 LACM 57, 210.1, NMCC 1990-0550 46 1 California holotype