Digitonthophagus Balthasar, 1959: taxonomy, systematics, and morphological phylogeny of the genus revealing an African species complex (Coleoptera: Scarabaeidae: Scarabaeinae) Génier, François Moretto, Philippe Zootaxa 2017 4248 1 1 110 363WL [151,455,799,824] Insecta Scarabaeidae Digitonthophagus Animalia Coleoptera 73 74 Arthropoda species falciger  Distribution: Benin, Burkina Faso, Cameroon, Chad,  Coted’Ivoire, Gambia, Ghana, Guinea, Guinea-Bissau, Mauritania,  Niger, Nigeria, Senegal, Sierra Leone,? Uganda