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        <dc:title>An updated catalogue of the Histeridae (Insecta: Coleoptera) of the Arabian Peninsula, with biogeographical remarks</dc:title>
        <dc:creator>Penati, Fabio</dc:creator>
        <dc:creator>Vienna, Pierpaolo</dc:creator>
        <rdf:type rdf:resource="fabio:JournalArticle"/>
        <bibo:journal>Zootaxa</bibo:journal>
        <dc:date>2006</dc:date>
        <bibo:pubDate>2006-03-23</bibo:pubDate>
        <bibo:volume>1157</bibo:volume>
        <bibo:issue>1</bibo:issue>
        <bibo:pageStart>1</bibo:pageStart>
        <bibo:pageEnd>74</bibo:pageEnd>
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    <rdf:Description rdf:about="http://taxon-concept.plazi.org/id/3971F007FF813D20044DBB80FCEDC356">
        <rdf:type rdf:resource="http://filteredpush.org/ontologies/oa/dwcFP#Taxon"/>
        <dwc:ID-CoL>TFTV</dwc:ID-CoL>
        <dwc:baseAuthorityName>Schmidt</dwc:baseAuthorityName>
        <dwc:baseAuthorityYear>1894</dwc:baseAuthorityYear>
        <dwc:box>[308,574,1244,1270]</dwc:box>
        <dwc:class>Insecta</dwc:class>
        <dwc:family>Histeridae</dwc:family>
        <dwc:genus>Chalcionellus</dwc:genus>
        <dwc:kingdom>Animalia</dwc:kingdom>
        <dwc:order>Coleoptera</dwc:order>
        <dwc:pageId>44</dwc:pageId>
        <dwc:pageNumber>45</dwc:pageNumber>
        <dwc:phylum>Arthropoda</dwc:phylum>
        <dwc:rank>species</dwc:rank>
        <dwc:species>hauseri</dwc:species>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_1">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Turkmenistan, Iran, Kazakhstan, Mongolia, Saudi Arabia.  Chorology:Centralasiatic.</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_2">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material. Saudi Arabia:“Saudi Arabia”, sine data( CHTY).  Notes.New to the Arabian Peninsula.  48.  Chalcionellus tunisius(Marseul, 1876)( Fig. 17)</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_3">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Tunisia, North Africa, Syria, Saudi Arabia, Yemen.  Chorology:N­African.  Literature.   Saudi Arabia: Sharoura[= Sharurah, 17°34’31”N 47°09’42”E],  29.III.1979, 1 ex.( Mazur 1994).</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_4">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Saudi Arabia: Daharan, Al Jubail[= Dhahran, al Jubayl, 27°02’50”N 49°39’41”E],  4.IV.2001, 6 ex.( 4 ex. MSNTC, 1 ♂ CHFP, 1 ex. CHPV).   Yemen:near Shiakh Othman(Wadi) [=Ash Shaykh`Uthman, 12°52’53”N 44°59’42”E],  III.1987, 1 ♀( MNHUB).  Notes.New to Yemen.   FIGURE 17.Distribution of the Histeridaein the Arabian Peninsula.  Chalcionellus geminus,  Chalcionellus tunisius,  Chalcionellussp..  49.  Pholioxenus kodymiOlexa( Fig. 18)</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_5">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Iraq,  Syria, Saudi Arabia.  Chorology:SW­Asiatic.  Literature.   Saudi Arabia: 90 kmN of Riyadh,  III–IV.1985, in pitfall trapson sandy soils in several wadis and in sand dune massif Irk­Banban[ 24°58’33”N 46°34’15”E], more than 80 ex.( Kryzhanovskij 1987).  50.  Pholioxenus mesopotamicusOlexa( Fig. 18)  Distribution: Iraq, Saudi Arabia, Kuwait.  Chorology:SW­Asiatic.  Literature.   Saudi Arabia: Wadi Hurayamala[= Wadi Huraymala, 25°07’29”N 46°06’53”E],  770 m,  24.III.1988, 4 ex.( Tishechkin 2002).</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_6">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Kuwait: Al Abdaliyah, 29°03’N 47°43’E,  24.II.1996, 3 ♀and 1 ♂( CMNH).  Notes.New to Kuwait.  51.  Pholioxenus olexaiKryzhanovskij( Fig. 18)</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_7">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Saudi Arabia.  Chorology:Endemic.  Literature.   Saudi Arabia: 90 kmN of Riyadh[= Riyad, 24°38’27”N 46°46’22”E],  27.III.1985, holotypeand ibidem,  26.III–14.IV1985, 11 paratypes( Kryzhanovskij 1987).   FIGURE 18.Distribution of the Histeridaein the Arabian Peninsula.  Pholioxenus kodymi,  Pholioxenus mesopotamicus,  Pholioxenus olexai.  52.  Hypocacculus( Colpellus) praecox(Erichson)( Fig. 19)  Distribution: Egypt, Mediterranean Subregion, Canary Is., Cape Verde, Afrotropical (= Ethiopian) Region, Saudi Arabia, Afghanistan, Oman.  Chorology:Afrotropico­Mediterranean.  Literature.   Saudi Arabia:“El Hahaz” [= El Hejaz] ( Reichardt 1932: 97; 2 ex.in MNHUBseen by us).</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_8">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Oman: Dhofar, Al Mughsaylenv. W, 16°53’74”N 53°48’97”E,  40 m,  5.IX.2000, 1 ♀( MSNTC).  Notes.New to Oman.   FIGURE 19.Distribution of the Histeridaein the Arabian Peninsula.  Hypocacculus ( C.) praecox,  Hypocacculus( C.) sp.,  Hypocacculus( C.) sp..  53.  Hypocacculus( Hypocacculus) harmonicus(Marseul)( Fig. 20)</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_9">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Republic of South Africa(Capland), East Africa, Sudan, Saudi Arabia, Oman.  Chorology:Afrotropical (spread in E and S Africa).  Literature.   Saudi Arabia:“El Hahaz” [= El Hejaz], 8 ex.( Reichardt 1932: 99; 5 ex.in MNHUBexamined by us).</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_10">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Oman: Dhofar, Salalahenv. W [ 17°01’03”N 54°04’58”E], near Veterinary Hospital,  27.VIII.2000, 1 ex.( MSNTC);  Dhofar, Rd. Taqahto Madnatal Haq, 2 kmafter jct.,  5.IX.2002, 3 ex.( MSNTC);  Dhofar, Rd.41 tra Raysute Adwnab, 16°57’72”N 53°52’73”E,  90 m,  23.IX.2001, 2 ex.( 1 ex. MSNTC, 1 ex. CHPV);  Dhofar, Khor Ron, 17°02’27”N 54°26’16”E,  35 m,  24.VIII.2000, 3 ex.( MSNTC);  ibidem,   12.IX.2000, 2 ex.( CHFP); Dhofar, Rd.49 near Ain Sahanawt, 17°07’40”N 54°09’44”E,   24.IX.2001, 2 ex.( MSNTC); Dhofar, Ain Razat, 17°07’75”N 54°14’42”E,  11.IX.2002, 2 ex.( MSNTC); Dhofar, Hajaif env., 17°14’53”N54°01’67”E,  900 m,   4.X.2001, 2 ex.( MSNTC); Dhofar, Rd.47 ante Al Mughsayl, 16°53’45”N 53°48’57”E,  30 m,   1.III.2004, 1 ♀( MSNTC); Dhofar, Ain Garzizenv., 17°06’29”N 54°04’35”E,  110 m,   16.III.2004, 1 ♂( MSNTC).  Notes.New to Oman.   FIGURE 20.Distribution of the Histeridaein the Arabian Peninsula.  Hypocacculus( s. str.) harmonicus,  H.( s. str.) metallescens,  H.( s. str.) simulans,  H.( s. str.) spretulus.  54.  Hypocacculus( Hypocacculus) metallescens(Erichson)( Fig. 20)</spm:hasContent>
    </rdf:Description>
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        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Italy( Sardinia), Hungary, Romania, Mediterranean Subregion, South Russia, Central Asia, Africa, Mascarene Is., Saudi Arabia, Oman.  Chorology:undefined.  Literature. Saudi Arabia:“El Hahaz” [= El Hejaz] ( Reichardt 1932: 99; 2 ex.in MNHUB seen by us); Khulais valley [= Khulays, 22°09’14”N 39°19’06”E], 80 kmNE of Jeddah, 1997–1998 ( Al­Ghamdi 1999).</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_12">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Saudi Arabia:“ Saudi Arabia”, sine data( CHTY).   Oman: Dhofar, Al Mughsaylenv. E, 16°52’53”N 53°43’51”E,  8.III.2004, 1 ♂( CHFP);  Dhofar, 7 kmafter Tawi Atayr, 17°03’46”N 54°36’15”E,  600 m,  9.III.2004, 1 ex.( MSNTC);  Dhofar, Ain Razatenv., 17°07’N 54°14’E,  16.III.2004, 1 ex.( MSNTC).  Notes.New to Oman. The chorology of this species is undefined, because documentation of its supposed wide distribution preceded the description of the two almost identical afrotropical species  H. simillimusVienna and  H. simulansVienna with which  H. metallescenscould have been confused in the past.  55.  Hypocacculus( Hypocacculus) simulans Viennain Penati et Vienna( Fig. 20)</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_13">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Republic of South Africa, Malawi, Yemen ( Gomy, 2004).  Chorology:Afrotropical (spread in E and S Africa).  Literature. Yemen:Wadi Sharez [a few km east of Hajjah, 15°41’36”N 43°36’11”E], II.1985, 1 ♀( Gomy 2004); ibidem, 24.III.1985, cow dung, 6 ♂and 4 ♀( Gomy 2004).  56.  Hypocacculus( Hypocacculus) spretulus(Erichson)( Fig. 20)  Distribution: Portugal, South Europe, Syria, Crimea, Caucasus, Central Asia, Iran, Saudi Arabia, Sri Lanka, Burma, Vietnam.  Chorology:undefined.  Literature.   Saudi Arabia: Fifanear Gizan[= Faifa, 17°08’38”N 43°07’09”E],  1240 m,  27–31.III.1983, 1 ex.( Mazur 1994).  57.  Hypocacculus( Nessus) controversusMüller( Fig. 21)  Distribution: Balkans, Spain, Greece, Turkey, Jordan, Saudi Arabia.  Chorology:Mediterranean.  Literature.   Saudi Arabia: Wadi Hurayamala[= Wadi Huraymala, 25°07’29”N 46°06’53”E],  770 m,  10.III.1988, 1 ex.( Tishechkin 2002).  58.  Hypocacculus( Nessus) hosseinius(Théry)( Fig. 21)  Distribution:  Morocco, Egypt, Iran, Saudi Arabia, Oman.  Chorology:Saharo­Sahelo­Sindian.</spm:hasContent>
    </rdf:Description>
    <rdf:Description rdf:about="http://treatment.plazi.org/id/3971F007FF813D20044DBB80FCEDC356#section_14">
        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Material.   Saudi Arabia: Kushm Dibi[?],  20.IV.1978, 1 ex.( CHSM).   Oman: Dhofar, Al Mughsaylenv., 16°53’90”N 53°46’30”E,  9.IX.2002, 1 ♂( MSNTC).  Notes.New to the Arabian Peninsula. This species seems to be closely related to a few other Palearctic Nessuscharacterized by striking psammophilous features (body convex, pronotal hypomeron setose, pronotal and elytral surface almost entirely and coarsely punctate, mid and hind tibiae enlarged bearing long spinulae, etc.):  H. tigris(Marseul),  H. orbusReichardt,  H. occatorReichardt,  H. eremobiusReichardt,  H. baluxReichardtand  H. neftensisOlexa. Dueto their relative rarity, these species are poorly known and for this reason a future comprehensive revision of this group might ascribe the two specimensof  H. hosseinius(Théry)from the Arabian Peninsula to another species.  59.  Hypocacculus( Nessus) rubripes(Erichson)( Fig. 21)</spm:hasContent>
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        <rdf:type rdf:resource="spm:InfoItem"/>
        <spm:hasContent>  Distribution: Portugal, Mediterranean Subregion, Netherlands, Central Europe, Crimea, Caucasus, Iran, Central Asia, Mongolia, Russian Far East: Ussuriyskiy Kray, India, tropical Africa.  Chorology:undefined.  Literature.   Saudi Arabia: Khulais valley[= Khulays, 22°09’14”N 39°19’06”E],  80 kmNE of Jeddah, 1997–1998 ( Al­Ghamdi 1999).</spm:hasContent>
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