A mountain of millipedes I: An endemic species-group of the genus Chaleponcus Attems, 1914, from the Udzungwa Mountains, Tanzania (Diplopoda, Spirostreptida, Odontopygidae) Enghoff, Henrik European Journal of Taxonomy 2014 2014-10-24 100 100 1 75 TH7H Enghoff, 2014 Enghoff 2014 [633,855,267,294] Diplopoda Odontopygidae Chaleponcus Animalia Spirostreptida 50 51 Arthropoda species teres sp. nov.   urn:lsid:zoobank.org:act: F4EB8A22-2F19-438F-8C65-DB6091867337  Figs 4, 6, 9, 25    Diagnosis Medium-sized. Differs from all other group members, except  C. malleolus sp. nov., by the absence of a dorsal spine on each anal valve. Further characterized by the subquadratic outline of the cucullus and the structure of the telomere, especially the presence of a slender, sinuous, spine-like branch. Gonopod coxa ( Fig. 9) without a lateral process; metaplical shelf-spine long.    Etymology The name is a Latin adjective meaning “smooth” and refers to the spineless anal valves.   Material studied(total: 29 ♂♂)    Holotype  TANZANIA: ♂, Iringa Region, Iringa District, Udzungwa Mts, New Dabaga/ UlangambiFR, 08°04’ 05.7” S, 35°54’ 18.8” E, montane,  1930–1950 masl, plot 1, casual,  16–19 Oct. 2000, Frontier Tanzanialeg. ( ZMUC).    Paratypes All from TANZANIA, Iringa Region, Iringa District, Udzungwa Mts, New Dabaga/ UlangambiFR, all collected by Frontier Tanzania, all in ZMUC: 27 ♂♂, 08°04’ 05.7” S, 35°54’ 18.8” E, montane,  1930– 1950 masl, plot 1, casual,  16–19 Oct. 2000;  1 ♂, 08°04’ 05.7” S, 35°54’ 13.8” E, montane,  1940 masl, plot 1, sample 3 of 3,  19 Oct. 2000.   Type locality  TANZANIA: Iringa Region, Iringa District, Udzungwa Mts, New Dabaga/Ulangambi FR, 08°04’ 05.7” S, 35°54’ 18.8” E, montane, 1930–1950 masl.   Description(male) DIAMETER. 1.9–2.2 mm, 45–47 podous rings. COLOUR. After 12 years in alcohol yellowish, with dark spots at ozopores and paramedian dark dorsal bands flanking a mid-dorsal light stripe; paramedian dark colouration sometimes extending down flanks on metazonites, giving the body a ringed appearance; body rings 6 and 7 darker than the others. ANAL VALVES ( Fig. 6D). Entirely withoutdorsal and ventral spines; marginal rim not raised, setae not on tubercles. LIMBUS ( Fig. 4H). With low, broad, rounded lobes, separated by parabolic sinuses; each lobe ca.4 times broader than long, margin densely and finely serrate, lobes indistinctly striate on external surface. TARSAL SETATION. Normal. GONOPOD COXA ( Figs 9, 25). About 3½ × as long as wide. Lateral margin straight in basal 2/3, then shallowly concave, angle between lateral and apical margins 90°, sharp; cucullus ( cu)hence subquadratic. Metaplical flange ( mf) smoothly rounded, no angle, no process; metaplical mesal margin straight up to level of arculus. Metaplical shelf ( ms) forming subrectangular process just distal to arculus, followed by very deep incision ( in) separating cucullus from basal part of metaplica; msproximally stout, continuing   Fig. 25.  Chaleponcus teres sp. nov., gonopods. A. Gonopods in situin seventh body ring, anterior view. B. Left gonopod, mesal view. C. Right gonopod, base of solenomere and telomere, ventral view. D. Right gonopod, posterior view. E. Left gonopod, anterior(-lateral-ventral) view. F. Right telopodite, mesalanterior-ventral view. G. Right telopodite, mesal-anterior view. al1, al2= branches of apical telomeral lamella, bal, bal’= basal lamella of telomere, cu= cucullus, in= metaplical incision, mf= metaplical flange, ms =metaplical shelf, mss= metaplical shelf-spine, ps= proximal spine of solenomere, slm= solenomere, spl= spine-like lamella of telomere, st, st’= main stem of telomere. Scales 0.1 mm. in metaplical shelf-spine ( mss) with no clear demarcation. mssmedium long, originating from anteriordistal surface of ms, first projecting distad, then curving mesad, and then slightly basad. GONOPOD TELOPODITE ( Fig. 25). Solenomere with a small, inconspicuous, irregular proximal spine ( ps). Telomere without a separate proximal lobe but posterior margin of proximal part of main stem ( st’) lying closely against base of solenomere ( slm). Telomere distally with three branches: • A large, roughly rectangular basal lamella ( bal) carrying a secondary lamella ( bal’) on its lateral side, • A distal spine-like branch, slender and sinuous ( spl), • An irregular apical lamella dividing in two branches, proximal branch ( al1) with smooth margins, the distal branch ( al2) with irregularly serrated margin. al1and bal in situcurving around opposite coxa ( Fig. 25A).    Distribution and habitat Known only from Udzungwa Mts, New Dabaga/Ulangambi FR. Altitudinal range: 1930–1950 masl. Habitat: montane forest.    Coexisting species   C. termini sp. nov.and  C. vilici sp. nov.were found in the same samples as  C. teres sp. nov.In addition,  C. dabagaensis, C. gracilior sp. nov.,  C. krai sp. nov.,  C. malleolus sp. nov.,  C. mwabvui sp. nov.,  C. nectarinia sp. nov.,  C. netus sp. nov.and  C. vandenspiegeli sp. nov.occur in New Dabaga/ Ulangambi FR.   Notes The gonopods somewhat resemble those of  C. hamerae sp. nov. 2626747319 2000-10-16 2000-10-19 2000-10-16 ZMUC Frontier Tanzania Tanzania Udzungwa Mts 1940 -8.06825 Udzungwa Mts 1 35.905224 Iringa District 50 51 1 1 Iringa Region holotype 5003826347 2000-10-16 2000-10-19 2000-10-16 ZMUC Frontier Tanzania Tanzania 1940 -8.06825 Udzungwa Mts 1 35.905224 Iringa District 50 51 27 27 Iringa Region paratype 5003826338 2000-10-19 2000-10-19 2000-10-16 ZMUC Frontier Tanzania Tanzania 1940 -8.06825 Udzungwa Mts 1 35.903835 Iringa District 50 51 1 1 Iringa Region paratype