Studies on the Australian Muscidae (Diptera). VIII. The genus Lispe Latreille, 1797 Pont, Adrian C. Zootaxa 2019 2019-02-18 4557 1 1 232 8CMT5 Pont, 2019 Pont 2019 [151,397,151,177] Insecta Anthomyiidae Lispe Animalia Diptera 41 42 Arthropoda species nigrimanoides sp. nov.  ( Figs 77–82)   Diagnosis.In the ♂of this species the armature of the mid and hind tarsomeres ( Figs 80–82) is unique. The ♂is similar to that of  L. nigrimana, but lacks the very long pvsetae on hind femur so characteristic of that species; the ventral surfaces of mid and hind femora are entirely without setae.   Etymology.The name indicates the close similarity between the new species and  L. nigrimana.   Type Material Examined.  Holotype ♂. AUSTRALIA: Western Australia: Millstream,  24.x.1970, D.H. Colless( ANIC).  Paratypes 1♂ 7♀. AUSTRALIA: Western Australia: same data as holotype,  1♀( ANIC); Millstream,  22.x.1970, D.H. Colless,  5♀(3 ANIC, 1 BMNH, 1 OUMNH); Millstream, at light,  23.x.1970, D.H. Colless  1♀( ANIC); Shothole Canyon, Cape Range, Exmouth, beside water,  12.viii.1970, P. Ferrar,  1♂( ANIC).   Description ♂ ♀. Head( Fig. 78). Ground-colour black. Frons at broadest point 0.33 ( ♂) or 0.43 ( ♀), at lunula 0.25 ( ♂) or 0.37 ( ♀), of maximum head-width (frontal view). Eye with an area of facets alongside lunula conspicuously enlarged. Ocellar setae very short and fine. Fronto-orbital plate with upper third matt to subshining, lower part brown ( ♂) to brownish-yellow ( ♀) pruinose, extreme lower part together with parafacial, face and gena yellowish-white pruinose, almost golden; lower 2/3 of occiput white pruinose, upper 1/3 thinly so but not shining, except weakly so at vertex. Lower reclinate orbital seta usually present. Frontal triangle slender ( ♂) to moderate ( ♀), shining black. Parafacial narrow, at middle 1/4 ( ♂) or 1/2 ( ♀) width of postpedicel; setulae short, fine, in 1 row. Antenna black, except tip of pedicel and base of postpedicel which are orange; postpedicel 2.0 times as long as broad, falling short of mouth-margin by 1.4 ( ♂) or 0.8 ( ♀) times its length. Arista plumose, the hairing at widest point 1.7 ( ♂) to 1.5 ( ♀) times width of postpedicel. Vibrissa strong but short, only just crossing, set high above mouth-margin, especially in ♂( Fig. 83), vibrissal area otherwise with few setulae. Gena moderate, 0.11 ( ♂) to 0.18 ( ♀) of vertical eye-length; genal setae weak. Palpus yellow, swollen at tip (as in  L. nigrimana, Fig. 59).  Thorax. Ground-colour black. Scutum brownish-grey dusted, with 3 inconspicuous darker brown vittae along the dcand acrlines, the acrvitta sometimes enlarged, sometimes all three rather enlarged, and a broader dark brown postpatch on each side outside the ialines; scutellum dark brown dusted on disc and at tip, the sides yellowish-grey in basal half; all pleura (including notopleuron) dull grey dusted. Anterior spiracle brown. Scutal ground-setulae short and decumbent. Acrsetulae in 4–5 irregular rows at suture. Dc0+2, the anterior postseta half as long as the posterior one. 1 pprn. 0+0 ia. 1+1 sa, the prstseta short and weak. 1 proepisternal seta, with an adjacent setula. 1 proepimeral seta, with 8-11 tiny adjacent setulae. Anepisternum without a short seta in upper anterior corner; posterior row with 1 strong and 4 weak setae. 3 kepst, lower one closer to posterior than to anterior one, anterior and lower setae well developed. Meron bare below spiracle, with a few tiny setulae above hind coxa. Scutellum with disc sparsely setulose, bare on sides; with some soft pale hairs on ventral surface at tip.  Legs. Coxae black; trochanters mostly reddish; fore leg black except for knee and basal 1/4 ( ♂) or 1/3 ( ♀) of tibia; mid and hind femora black in basal half and yellow in apical half or a little more; mid and hind tibiae yellow; tarsomeres black. Fore coxa bare behind. Fore femur without avsetae; pvsurface with 3–4 short setae in apical half, otherwise short setulose; pdrow complete. Fore tibia not expanded or flattened; without submedian setae; with dand short pvapical setae, the pvabsent in ♂. Fore tarsus without modifications. Mid femur without av, pvor asetae; without preapical ad; 1 ppreapical. Mid tibia with 0 adand 1 p; with moderate a, vand pvapicals, and a short dand p. Mid tarsus of ♂modified ( Fig. 80): basal tarsomere elongated, and tarsomeres 1–3 with sparse semierect vsetulae; not modified in ♀. Hind coxa without a seta on posterior apical margin. Hind femur without avor pvsetae; adrow complete, weak; 0 pdpreapical. Hind tibia with 1 ad, without avor pd; 1 dbut no adpreapical seta; 1 short avapical but no pv. Hind tarsus of ♂( Figs 81–82) with basal tarsomere slightly bent, concave, with a vseta near base (also present in ♀), a row of asetae, long at base, becoming shorter and continuing on to tarsomere 2, avto vsurfaces finely setulose; not modified in ♀.  Wing. Narrow. Very weakly smoky. Tegula brown, basicosta orange. Crossvein r-m placed below the point where vein R1 enters costa; dm-cu straight, longer than apical section of vein CuA1. Vein M running straight to wing-margin. Calypters white to creamy, with creamy margins. Haltere yellow.  Abdomen. Ground-colour black. Segments 4 and 5 of ♂modified as in  L. nigrimana( Figs 67–68). Tergites subshining black with very thin dark brown dust; ♂with white dusted spots as in  L. nigrimana( Figs 64, 66), except that the lateral and dorsal areas of dust on syntergite 1+2 are connected by a hind-marginal band of dust; more yellowish-grey in ♀, the pattern as in  L. nigrimana( Fig. 65) except that syntergite 1+2 is more extensively dusted. ♂epandrium dusted white. Sternites 1–4 ( ♂) or 1–5 ( ♀) light grey dusted. Setae on tergites very weak, tergites 3 to 5 each with 1 pair of weak lateral marginal setae, tergite 5 with a few additional shorter marginals that form a row, without discals; these all very short and inconspicuous in ♀. ♂sternites 2–4 with sparse setulae.  Ƌ terminalia. Not differing in any significant way from those of  L. nigrimana(see Figs 70–72). Cercal plate more indented on lower outer edge ( Fig. 79).   ♀ovipositor. Not differing from that of  L. nigrimana(see Figs 73–75). Ventral membrane of segment 8 sometimes with a bristle on one or both sides, indicating the former presence of 2 small plates representing sternite 8.   FIGURE 77.  Lispe nigrimanoides  sp. nov., ♂ habitus (holotype).  Measurements. Wing-length, 3.0– 3.5 mm( ♂), 3.5–4.0 mm ( ♀). Body-length, 4.5–5.0 mm ( ♂), 4.5–5.0 mm ( ♀).   Biology.Nothing known.   Distribution.Known only from the type-series from Australia(WA). 2004852036 1970-10-24 ANIC D. H. Colless Australia Millstream 41 42 1 1 Western Australia holotype 2004852033 [357,1273,438,463] 1970-10-24 ANIC D. H. Colless Australia Millstream 41 42 8 7 1 Western Australia paratype 2004852038 1970-10-22 ANIC D. H. Colless Australia Millstream 41 42 1 1 paratype 2004852049 1970-10-23 ANIC, BMNH, OUMNH D. H. Colless Australia Millstream 41 42 5 5 paratype 2004852059 [240,1258,510,535] 1970-08-12 ANIC P. Ferrar Australia Shothole Canyon Exmouth Cape Range 41 42 1 1 paratype 2004852050 [1269,1402,511,535] ANIC Australia 41 42 1 1 paratype