Carabodes
minusculus
Tegeocranus
(Flexa)
Austrocarabodes
ornatus
labyrinthicus
Austrocarabodes
castrii
granulatus
forsslundi
forsslundi
Oribates
Brasiella
Chaunoproctus
femoralis rugosior
femoralis rugosior
minusculus
Flexa
longulus
A survey of the family Carabodidae C. L. Koch, 1836 (Acari: Oribatida), II
Mahunka, S.
Acta Zoologica Hungarica
1987
33
399
434
3H8M
229788
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
2
401
Arthropoda
genus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
2
401
Arthropoda
genus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
2
401
Arthropoda
species
coriaceus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
3
402
Arthropoda
species
agenjoi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
alveolatus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
apicalis
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
3
402
Arthropoda
species
aphanicus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
areolatus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
atrichosus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
3
402
Arthropoda
species
berninii
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
canaliculatus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
3
402
Arthropoda
species
castrii
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
celisi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
cellularis
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
cephalotes
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
3
402
Arthropoda
species
chirstlus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
coronatus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
4
403
Arthropoda
species
cynocephalus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
davisi
Arachnida
Carabodidae
Carabodes
missing, matching species
1
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
dubius
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
ensifer
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
excellens
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
flagellifer
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
forsslundi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
fraterculus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
4
403
Arthropoda
species
gibbiceps
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
granosus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
haradai
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
hispanicus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
4
403
Arthropoda
species
humeratus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
hummelincki
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
4
403
Arthropoda
species
imperfectus
Arachnida
Carabodidae
Carabodes
missing, matching species
1
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
intermedius
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
4
403
Arthropoda
species
irmayi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
4
403
Arthropoda
species
jamaicaensis
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
lepidus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
longisetosus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
lounsburyi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
longulus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
longulus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
lunaris
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
luteoauratus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
magnus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
marginepunctatus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
microtrichus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
montanus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
niger
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
nitens
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
oblonga
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
5
404
Arthropoda
species
obsoletus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
ornatissimus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
5
404
Arthropoda
species
palmifer
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
papillosus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
paraspinosus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
penicillatus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
pulcher
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
rimosus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
rugosior
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
schatzi
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
schwartzi
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
scymnus
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
strinovichi
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
subnudus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
6
405
Arthropoda
species
teneriffensis
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
travei
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
6
405
Arthropoda
species
trigonosternum
Arachnida
Carabodidae
Carabodes
Animalia
Sarcoptiformes
7
406
Arthropoda
species
verrucatus
Arachnida
Carabodidae
Carabodes
missing
1
Animalia
Sarcoptiformes
7
406
Arthropoda
species
willmanni
I have made extensive use of the highly valuable manuscript of E. Piffl cataloguing the species of these genera. The ancient and for a long time the single genus of the family was the type-genus CarabodesC. L. Koch, 1836. Although, by several authors (Berlese, 1910, 1913; Traegardh, 1931; Willmann, 1936; Balogh, 1958) some supraspecific taxa were described until 1965, some of them either proved to be synonymous with or they stood so far from the type-genus that it was rather unlikely that a greater number of the older Carabodesspecies would be ranked into the new genera. But the situation changed, when erecting the genus AustrocarabodesHammer (1966: 62) included in it only his new species and a few older ones originally described in the genus Carabodes. Many of the earlier combinations with Carabodeswere wrong, as were also misconceived some with Austrocarabodes, therefore, our present-day knowledge spurs us to compile a catalogue of species for both genera. The lists of the other genera will be published at a later date. The subsequent list includes all the taxa originally described into this genus, which either still belong here today or not. The literature is restricted only to the most reliable and most important references. I always considered a reference to be useful if it gave a new combination, a new morphological feature, a note to zoogeography or to the ecology of the species. The italicized names refer to species which do not belong here today. These names are followed with reference to the new combination:
CarabodesC. L. Koch, 1836:3, 15
NeocepheusWillmann, 1936: 433 Flexa Kulijev, 1977: 64 syn. n.
Bernini 1970: 402, t: II, figs 3-4, III, 2-3 Bernini 1982: 386
Hammer 1966: 62
Hammer 1966: 62
Jacot 1937: 240 Norton 1978: 611
Bernini 1982: 387
Sellnick und Forsslund 1953: 378, fig. 5 Schweizer 1956: 293, figs 219 a-f Rajski 1968: 293 Bernini 1970: 403, t: III, 1, 4, IV: 1-3 Bernini 1982: 388
Bulanova-Zachvatkina 1975: 187, fig. 389
Norton 1978: 614, fig. 10
Oudemans 1937: 2580, fig. 1109*
* It is almost certain that Willmann's note is correct (see Oudemans), consequently, the species does not belong to the family Carabodidae.
Bernini 1970: 404 (= minusculus syn. n.)
Balogh 1961: 276, t. 15: 1-2 Hammer 1966: 62
Hammer 1966: 62
Oudemans 1937: 2632, fig. 1136
** It is almost certain that it does not belong to the genus Carabodes, nor even to the family Carabodidae.
Michael 1884: 316, t: XX, 1-8, XXII, 11 ( Tegeocranus) Oudemans 1937: 2640, fig. 1141 Sellnick und Forsslund, 1953: 381, fig. 7 Rajski 1968: 292 Bernini 1970: t: VIII, 3-4 Perez-Inigo1971: 281, fig. 16 Mahunka 1986: 117, figs 13-15, 84
Balogh 1960: 92, figs 6 - 8 Hammer 1966: 62
Oudemans 1937: 2646, fig. 1142
* The combination of Willmann and Oudemans is rather uncertain. Considering the shape of the legs, it rather belongs to the groups of Oppoidea.
Jacot 1937: 238, fig. 5 Norton 1978: 611
Kulijev 1977: 66, figs 1: 1-6, 2: 1-3 (Flexa)
Hammer 1966: 59 (> to Austrocarabodes comb. n.)
Michael 1884: 318, t: XX, 9 Hull 1914: 282 Sellnick und Forsslund 1953: 386, fig. 10 Bernini 1970: 404, t: X, 3-4, XI, 3-4, XX, 1
** Cepheus flavus Ewing, 1918: Woolley (1957: 113, figs 1 - 2) erroneously placed it in the genus Carabodes. I think that it belongs to the family Cepheidae.
Rajski 1968: 297 (syn. with ornatus)
Balogh 1965: 59, t: 9, fig. 7
Bernini 1982: 389
Norton, 1978: 614, fig. 11
Perez-Inigo, 1971: 284 (syn. with labyrinthicus)
Perez-Inigo1971: 284
Balogh 1970: 299 ( Austrocarabodes)
Bernini 1981: 27, figs 1-8
Hammer 1966: 62
Schweizer 1956: 295, figs 222: a -c
Mahunka 1986: 109
Michael 1884: 319, figs t: XXI, 1-8, XXII, 3 Hammer1952: 42, fig. 62 Sellnick und Forsslund 1953: 387, fig. 12 Schweizer 1956: 294, figs 221: a - c Rajski 1968: 294 Perez-Inigo1971: 287, figs 19-20
Aoki 1982: 175
Bulanova-Zachvatkina 1975: 185
Balogh 1965: 59 Mahunka 1986: 132, figs 147 - 151
Hammer 1966: 62
Sellnick und Forsslund 1953: 373 (as nom. nud.)
Hammer 1966: 62
Hammer 1973: 17
Sellnick und Forsslund 1953: 382, fig. 8 Rajski 1968: 296 Bernini 1970: 404, t: VI, 3-4, VII, 1-2, VIII, 2 Bernini 1974: 51, figs 1-4
Traegardh1910: 515, figs 284-289 Sellnick et Forsslund 1953: 388
Sellnick und Forsslund 1953: 388, fig. 13 Mahunka 1966: 379, fig. 12 ( castrii) Bernini 1970: 404, t: IV, 4-5, V, 1-4 Perez-Inigo1971: 282, figs 17-18 Bernini 1976: 3, figs I: a-h, II: a-j, III: a-j, IV: a-d; 39, 45
Sellnick und Forsslund 1953: 380, 385
Norton 1978: 613, figs 3-9
* On the basis of Piffl's catalogue. I was unable to check it myself.
Johnston 1965: 54 Norton 1978: 613
Norton 1978: 615 (proposed synonymy with granulatus)
Sellnick und Forsslund 1953: 383, fig. 9 ( forsslundi) Schweizer 1956: 294, figs 220: a -c ( forsslundi) Rajski 1968: 297 Bernini 1982: 390
Bernini 1970: 404, figs 5: e-f Bernini 1982: 390
Michael 1898: 39 Oudemans, 1937 2647, fig. 1143 ( Oribates)
Bulanova-Zachvatkina 1975: 185
Bernini 1970: 407, t: VI, 1-2, VII, 3-4, VIII, 1
Sellnick und Forsslund 1953: 379, fig. 6 Bernini 1970: 409, t: IX, 1-5 Oudemans 1915: 194> to Brasiella comb. n. Oudemans 1917: 55, figs 96 - 97 Sellnick und Forsslund 1953: 79, fig. 6 ( Chaunoproctus) Balogh 1970: 311, figs 54-55
Aoki 1970: 417, figs 44-46
Sellnick und Forsslund 1953: 387, fig. 11 ( femoralis rugosior) Rajski 1968: 294 ( femoralis rugosior) Bernini 1970: 410, t: X, 1, XI, 1-2, XII, 2 Bernini 1982: 392
Schweizer 1956: 295, fig. 223 ( minusculus)
Kulijev 1977: 66, figs 3: 1-3 ( Flexa)
Hammer 1966: 62
Rajski 1968: 298 Sellnick und Forsslund 1953: 375, fig. 4
Oudemans, 1917: 57, figs 98 - 107
Forsslund 1943: 190 nom. nud. Willmann 1944:* ( longulusWillmann nom. nud.) Rajski 1968: 299 Hammer 1977: 30 * I have not seen the publication.
** On the basis of Piffl's catalogue.
Bernini 1970: 402
* On the basis of the figure, it definitely possesses more than 10 notogastral setae, though, as far as its habitus is concerned, a relationship with Diplobodes- Kalloiais more probable. Further research is necessary.
Perez-Inigo1971: 282, figs 17-18 Bernini 1976: 21