Carabodes minusculus Tegeocranus (Flexa) Austrocarabodes ornatus labyrinthicus Austrocarabodes castrii granulatus forsslundi forsslundi Oribates Brasiella Chaunoproctus femoralis rugosior femoralis rugosior minusculus Flexa longulus A survey of the family Carabodidae C. L. Koch, 1836 (Acari: Oribatida), II Mahunka, S. Acta Zoologica Hungarica 1987 33 399 434 3H8M 229788 Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 2 401 Arthropoda genus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 2 401 Arthropoda genus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 2 401 Arthropoda species coriaceus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 3 402 Arthropoda species agenjoi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species alveolatus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species apicalis Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 3 402 Arthropoda species aphanicus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species areolatus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species atrichosus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 3 402 Arthropoda species berninii Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species canaliculatus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 3 402 Arthropoda species castrii Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species celisi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species cellularis Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species cephalotes Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 3 402 Arthropoda species chirstlus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species coronatus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 4 403 Arthropoda species cynocephalus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species davisi Arachnida Carabodidae Carabodes missing, matching species 1 1 Animalia Sarcoptiformes 4 403 Arthropoda species dubius Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species ensifer Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species excellens Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species flagellifer Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species forsslundi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species fraterculus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 4 403 Arthropoda species gibbiceps Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species granosus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species haradai Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species hispanicus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 4 403 Arthropoda species humeratus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species hummelincki Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 4 403 Arthropoda species imperfectus Arachnida Carabodidae Carabodes missing, matching species 1 1 Animalia Sarcoptiformes 4 403 Arthropoda species intermedius Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 4 403 Arthropoda species irmayi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 4 403 Arthropoda species jamaicaensis Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species lepidus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species longisetosus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species lounsburyi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species longulus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species longulus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species lunaris Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species luteoauratus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species magnus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species marginepunctatus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species microtrichus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species montanus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species niger Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species nitens Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species oblonga Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 5 404 Arthropoda species obsoletus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species ornatissimus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 5 404 Arthropoda species palmifer Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species papillosus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species paraspinosus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species penicillatus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species pulcher Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species rimosus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species rugosior Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species schatzi Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species schwartzi Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species scymnus Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species strinovichi Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species subnudus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 6 405 Arthropoda species teneriffensis Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species travei Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 6 405 Arthropoda species trigonosternum Arachnida Carabodidae Carabodes Animalia Sarcoptiformes 7 406 Arthropoda species verrucatus Arachnida Carabodidae Carabodes missing 1 Animalia Sarcoptiformes 7 406 Arthropoda species willmanni  I have made extensive use of the highly valuable manuscript of E. Piffl cataloguing the species of these genera. The ancient and for a long time the single genus of the family was the type-genus CarabodesC. L. Koch, 1836. Although, by several authors (Berlese, 1910, 1913; Traegardh, 1931; Willmann, 1936; Balogh, 1958) some supraspecific taxa were described until 1965, some of them either proved to be synonymous with or they stood so far from the type-genus that it was rather unlikely that a greater number of the older Carabodesspecies would be ranked into the new genera. But the situation changed, when erecting the genus AustrocarabodesHammer (1966: 62) included in it only his new species and a few older ones originally described in the genus Carabodes. Many of the earlier combinations with Carabodeswere wrong, as were also misconceived some with Austrocarabodes, therefore, our present-day knowledge spurs us to compile a catalogue of species for both genera. The lists of the other genera will be published at a later date. The subsequent list includes all the taxa originally described into this genus, which either still belong here today or not. The literature is restricted only to the most reliable and most important references. I always considered a reference to be useful if it gave a new combination, a new morphological feature, a note to zoogeography or to the ecology of the species. The italicized names refer to species which do not belong here today. These names are followed with reference to the new combination:   CarabodesC. L. Koch, 1836:3, 15   NeocepheusWillmann, 1936: 433  Flexa Kulijev, 1977: 64 syn. n.  Bernini 1970: 402, t: II, figs 3-4, III, 2-3 Bernini 1982: 386  Hammer 1966: 62  Hammer 1966: 62  Jacot 1937: 240 Norton 1978: 611  Bernini 1982: 387  Sellnick und Forsslund 1953: 378, fig. 5 Schweizer 1956: 293, figs 219 a-f Rajski 1968: 293 Bernini 1970: 403, t: III, 1, 4, IV: 1-3 Bernini 1982: 388  Bulanova-Zachvatkina 1975: 187, fig. 389  Norton 1978: 614, fig. 10  Oudemans 1937: 2580, fig. 1109*  * It is almost certain that Willmann's note is correct (see Oudemans), consequently, the species does not belong to the family Carabodidae.  Bernini 1970: 404 (= minusculus syn. n.)  Balogh 1961: 276, t. 15: 1-2 Hammer 1966: 62  Hammer 1966: 62  Oudemans 1937: 2632, fig. 1136  ** It is almost certain that it does not belong to the genus Carabodes, nor even to the family Carabodidae.  Michael 1884: 316, t: XX, 1-8, XXII, 11 ( Tegeocranus) Oudemans 1937: 2640, fig. 1141 Sellnick und Forsslund, 1953: 381, fig. 7 Rajski 1968: 292 Bernini 1970: t: VIII, 3-4  Perez-Inigo1971: 281, fig. 16 Mahunka 1986: 117, figs 13-15, 84  Balogh 1960: 92, figs 6 - 8 Hammer 1966: 62  Oudemans 1937: 2646, fig. 1142  * The combination of Willmann and Oudemans is rather uncertain. Considering the shape of the legs, it rather belongs to the groups of Oppoidea.  Jacot 1937: 238, fig. 5 Norton 1978: 611  Kulijev 1977: 66, figs 1: 1-6, 2: 1-3 (Flexa)  Hammer 1966: 59 (> to Austrocarabodes comb. n.)  Michael 1884: 318, t: XX, 9 Hull 1914: 282 Sellnick und Forsslund 1953: 386, fig. 10 Bernini 1970: 404, t: X, 3-4, XI, 3-4, XX, 1  ** Cepheus flavus Ewing, 1918: Woolley (1957: 113, figs 1 - 2) erroneously placed it in the genus Carabodes. I think that it belongs to the family Cepheidae.  Rajski 1968: 297 (syn. with ornatus)  Balogh 1965: 59, t: 9, fig. 7  Bernini 1982: 389  Norton, 1978: 614, fig. 11   Perez-Inigo, 1971: 284 (syn. with labyrinthicus)   Perez-Inigo1971: 284  Balogh 1970: 299 ( Austrocarabodes)  Bernini 1981: 27, figs 1-8  Hammer 1966: 62  Schweizer 1956: 295, figs 222: a -c  Mahunka 1986: 109  Michael 1884: 319, figs t: XXI, 1-8, XXII, 3  Hammer1952: 42, fig. 62 Sellnick und Forsslund 1953: 387, fig. 12 Schweizer 1956: 294, figs 221: a - c Rajski 1968: 294  Perez-Inigo1971: 287, figs 19-20  Aoki 1982: 175  Bulanova-Zachvatkina 1975: 185  Balogh 1965: 59 Mahunka 1986: 132, figs 147 - 151  Hammer 1966: 62  Sellnick und Forsslund 1953: 373 (as nom. nud.)  Hammer 1966: 62  Hammer 1973: 17  Sellnick und Forsslund 1953: 382, fig. 8 Rajski 1968: 296 Bernini 1970: 404, t: VI, 3-4, VII, 1-2, VIII, 2 Bernini 1974: 51, figs 1-4   Traegardh1910: 515, figs 284-289 Sellnick et Forsslund 1953: 388  Sellnick und Forsslund 1953: 388, fig. 13 Mahunka 1966: 379, fig. 12 ( castrii) Bernini 1970: 404, t: IV, 4-5, V, 1-4  Perez-Inigo1971: 282, figs 17-18 Bernini 1976: 3, figs I: a-h, II: a-j, III: a-j, IV: a-d; 39, 45  Sellnick und Forsslund 1953: 380, 385  Norton 1978: 613, figs 3-9  * On the basis of Piffl's catalogue. I was unable to check it myself.  Johnston 1965: 54 Norton 1978: 613  Norton 1978: 615 (proposed synonymy with granulatus)  Sellnick und Forsslund 1953: 383, fig. 9 ( forsslundi) Schweizer 1956: 294, figs 220: a -c ( forsslundi) Rajski 1968: 297 Bernini 1982: 390  Bernini 1970: 404, figs 5: e-f Bernini 1982: 390  Michael 1898: 39 Oudemans, 1937 2647, fig. 1143 ( Oribates)  Bulanova-Zachvatkina 1975: 185  Bernini 1970: 407, t: VI, 1-2, VII, 3-4, VIII, 1  Sellnick und Forsslund 1953: 379, fig. 6 Bernini 1970: 409, t: IX, 1-5 Oudemans 1915: 194> to Brasiella comb. n. Oudemans 1917: 55, figs 96 - 97 Sellnick und Forsslund 1953: 79, fig. 6 ( Chaunoproctus) Balogh 1970: 311, figs 54-55  Aoki 1970: 417, figs 44-46  Sellnick und Forsslund 1953: 387, fig. 11 ( femoralis rugosior) Rajski 1968: 294 ( femoralis rugosior) Bernini 1970: 410, t: X, 1, XI, 1-2, XII, 2 Bernini 1982: 392  Schweizer 1956: 295, fig. 223 ( minusculus)  Kulijev 1977: 66, figs 3: 1-3 ( Flexa)  Hammer 1966: 62  Rajski 1968: 298 Sellnick und Forsslund 1953: 375, fig. 4  Oudemans, 1917: 57, figs 98 - 107  Forsslund 1943: 190 nom. nud. Willmann 1944:* ( longulusWillmann nom. nud.) Rajski 1968: 299 Hammer 1977: 30 * I have not seen the publication.  ** On the basis of Piffl's catalogue.  Bernini 1970: 402  * On the basis of the figure, it definitely possesses more than 10 notogastral setae, though, as far as its habitus is concerned, a relationship with Diplobodes- Kalloiais more probable. Further research is necessary.   Perez-Inigo1971: 282, figs 17-18 Bernini 1976: 21