Coptorhina davidi Janssens, 1939:35 Ferreira, 1972:357 Revision of the obligate mushroom-feeding African ‘‘ dung beetle’ ’ genus Coptorhina Hope (Coleoptera: Scarabaeidae: Scarabaeinae) Frolov, Andrey V. Akhmetova, Lilia A. Scholtz, Clarke H. Journal of Natural History 2008 2008-06-30 42 21 - 24 1477 1508 Y87Y Janssens Janssens 1939 [471,777,1370,1394] Insecta Scarabaeidae Coptorhina Animalia Coleoptera 23 1498 Arthropoda species davidi  ( Figures 13G, 19)      Coptorhina davidi Janssens, 1939:35, 38;  Ferreira, 1972:357.   Type material examined  Holotypewith labels ‘‘Coll. R.I.Sc.N.B. Congo Belge KatangaAnkoro 7.I-1939Dr David’’, ‘‘A Janssens det., 1939: Coptorhina Davidinsp. Type’’, ‘‘Type’’ ( IRSNB). 11 paratypes: 1 spm. with labels ‘‘Paratypus’’, ‘‘ ParatypeC. Davidi’’, ‘‘Musee du Congo belgeKundelungu 9-22. XII.1910’’, ‘‘R. DÉT. 432’’, ‘‘Paratype’’, ‘‘438’’, ‘‘ optata Pér.’’, ‘‘A. Janssens det., 1939: Coptorhina Davidi n. sp.’’, ‘‘R. DÉT. L 3943’’ ( MRAC). 1 spm. with labels ‘‘Paratypus’’, ‘‘ ParatypeC. Davidi’’, ‘‘Musee du Congo DrSheffield Neave MADONA-BANG-WEOLO XII.07’’, ‘‘R. DÉT. 264’’, ‘‘Paratype’’, ‘‘A. Janssens det., 1939: Coptorhina Davidi n. sp.’’, ‘‘R. DÉT. M 3943’’ ( MRAC). 5 spm. with labels ‘‘Coll. R.I.Sc.N.B. ZambiaDr Sheffield Neave MPIKA XII. 07’’, ‘‘A. Janssens det., 1939: Coptorhina Davidi n. sp.’’, ‘‘Paratype’’ ( IRSNB); 3 of them with additional labels ‘‘R. DÉT. 264’’, ‘‘A. Janssens det., 1939: Coptorhina Davidi n. sp.’’, ‘‘R. DÉT. M 3943’’; 1 spm. with additional label ‘‘ Coptorhina auspicata Per. Det. J.J.E.Gillet’’. 4 spm. with labels ‘‘Coll. R.I.Sc.N.B. ZambiaDr Sheffield Neave MPIKA I.08’’, ‘‘C. J. Arrow det. Coptorhina auspicata Pér.’’, ‘‘A. Janssens det., 1939: Coptorhina Davidi n. sp.’’, ‘‘Paratype’’ ( IRSNB); three of them with additional label ‘‘J.J. Gillet det., vend.: Coptorhina auspicata Pering R.M.H.N. Belg.10.640’’.  Additional material examined    Tanzania: 1 spm. ( NMPC).   Democratic Republic of Congo: Katanga[09 °419 S 23 °449 E],  November 1937, F.G. Overlaetleg. 1 spm. ( IRSNB).   Zambia: N’Changa[12 °319 S 27 °529 E],  December 1930, C.T. Macnamaraleg. 1 „( TMSA); Serenje District [13 °149 S 30 °149 E],  19–22 December 1907,  4500 ft, Neaveleg. 1 „( BMNH);   23–26 December 1907,  4500 ft, 1 spm. ( BMNH); N of Broken Hill, Kashitu[14 °269 S 28 °279 E],  December 1914, H.C. Dollmanleg. 1 spm. ( SANC), 1 spm. ( BMNH).  Diagnosis This species is most similar to  C. klugiiand  C. angolensisbut differs from them in the sculpture of the pronotum: disc basad of transverse carina densely punctate, sometimes rugose, even in larger specimens; transverse carina with two distinct medial tubercles and two indistinct or absent lateral ones ( Figure 8). From other species it differs in having narrow lateral borders of the pronotum and evenly rounded lateral sides of the pronotum.  Description Strongly convex, black to dark-brown, subopaque beetle. Body length 10.2– 17.0 mm, width 7.0–11.0 mm. Dorsal surface without visible setae.  Head. Clypeus with protruding anterior angles separated by deep sinuation ( Figure 8). Sinuation right-angled to broadly rounded. Protruding angles acute to narrowly rounded apically, with short longitudinal carinae on the upper side. Genae obtusely rounded. Genal sutures feebly distinct. Lateral margins of clypeus sinuate near genal sutures in some individuals. Frontal suture feebly visible, broadly interrupted in the middle. Head very densely and evenly punctuate, punctures sometimes adjacent and sculpture appears rugose. Antennal club dark-brown.  Pronotum. Trapezoidal, two times wider than long. Anterior margin and base not bordered. Sides more or less evenly rounded. Lateral border relatively narrow, evenly tapering basally and apically. Pronotum divided into larger anterior part and smaller posterior part by a transverse carina ( Figure 8). The carina is more developed in larger specimens where it has two distinct medial tubercles and two indistinct or absent lateral ones ( Figure 8). Anterior part of pronotum rugose to granulate, excavated laterally in larger specimens. Posterior part in specimens with well developed carina densely punctate to rugose.  Elytra. Striae fine but distinct, punctate (punctures separated by 3–4 puncture diameter). Intervals flat, very densely punctuate, punctures separated by about 1 puncture diameter, sometimes almost adjacent; their margins sometimes indistinct especially in apical part.  Underside. Pygidium with fine border, its disc densely punctate. Abdominal and thoracic sternites coarsely punctate except for disc of metasternum which is very sparsely to almost indistinctly punctuate.  Aedeagus. Parameres with widely rounded apices in lateral view.  Variability Except for the body size variability the beetles differ in shape and sculpture of pronotum. Smaller specimens have less developed transverse pronotal carina and little excavated anterolateral sides of pronotum.  Distribution The species is distributed west and north of Tanganyika Lake ( Figure 19). It is known from a few localities in Zambiaand Democratic Republic of Congoand from an unknown locality in Tanzania. [213,544,640,664] NMPC Tanzania Tanzania 24 1499 1 1937-11 IRSNB F. G. Overlaet Democratic Republic of the Congo Katanga 24 1499 1 Katanga 1907-12-19 1930-12 1907-12-19 TMSA, BMNH C. T. Macnamara & Neave Zambia 1372 N'Changa 24 1499 1 [213,755,768,792] 1907-12-23 1907-12-26 1907-12-23 BMNH Zambia 1372 24 1499 1 1914-12 SANC H. C. Dollman Zambia Kashitu Broken Hill 24 1499 1