Isoperla fulva Claassen 1937
The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species
Sandberg, John B.
Kondratieff, Boris C.
Illiesia
2013
9
5
34
64
6MZWW
Claassen
Claassen
1937
[998,1279,366,391]
Insecta
Perlodidae
Isoperla
Animalia
Plecoptera
12
46
Arthropoda
species
fulva
( Figs. 6a-d, 20b)
Isoperla fulva Claassen 1937.
Holotype ♂, Utah, Logan River.
Male. Aedeagus: sclerotized posterior process present; body with one posterior lobe, one small dorsal lobe, and one large anterior lobe ( Fig. 6a); one small bulbous patch of spinulae above sclerotized process ( Figs. 6a-c); sclerotized process length approaching 0.5 mmwhen fully extended, blade-like in lateral view ( Fig. 6a), distal margin expanded and slightly membranous in posterior view ( Fig. 6b), and narrow proximal portion of sclerotized process sometimes extended past posterior aedeagal membrane ( Fig. 6a). Abdominal terga 8-9, 9, 9-10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering abruptly to blunt apices ( Fig. 6a). Vesicle: rounded lobe, widest at base with broadly rounded apical margin ( Fig. 6d).
Isoperla fulvais most similar to the two other I. marmoratacomplex species, I. marmorataand I. roguensis. These species differ in paraproct shape and aedeagal characters. The paraprocts taper abruptly to blunt apices in I. fulva, taper gradually to blunt apices in I. marmorata, and taper abruptly to tiny sharp apices in I. roguensis( Table 1). The distal margin of blade-like sclerotized processes is thin in I. marmorata, expanded in I. fulvaand expanded and variably-shaped in I. roguensis. The aedeagus of this species was described with an additional pair of long narrow tubular lobes extending from the base of the anterodorsal lobe ( Szczytko & Stewart 1979, fig. 138C). When using the live eversion method, these processes remain inverted ( Fig. 6a); however two small areas in inverted membrane are present on anterior lobe C of this study (not discernible in Fig. 6a). The live everted aedeagus was cleared in KOH, which allowed observation of an inverted pair of tube-like processes within the aedeagus, and openings along the anteroapical margin ( Fig. 20b). The small dorsal lobe B ( Figs. 6a & c) in the current study possesses a pair of minute, closely placed holes from which seminal fluid streamed after live eversion and fixing. These dorsal aedeagal openings require further study before their function can be determined.
Isoperla fulvais putatively known from Californiabased on three malesand one female(El Dorado and Modoc counties - Jewett 1960) and Plumas County ( Szczytko & Stewart 1979) collected in the 1940’s and 1950’s. No recent material (everted males) from Californiacompares well with the Coloradoand Utahspecimens examined for this study, and the California I. fulvarecords were determined before the description of I. roguensis. The El Dorado County male was likely collected from Weber Creek, south of Camino, 10 miles( 16 km) north of Placerville, where Sandberg (2011b)collected I. marmorataand I. mormona.
Additional Material Examined. COLORADO, Larimer Co., Sand Creek, Ruby Wash, Red Mountain Open Space, 24/ V/2012, B.C. Kondratieff& D. Givens, 5♂, 2♀( CSUC); 17/ VI/2012, B.C. Kondratieff& R. Stokes, 10♂, 3♀( CSUC). OREGON, Harney Co., Trout Creek, Whitehorse Ranch Rd 18 mi( 29 km) SE Fields, 42.17439°N, 118.43199°W, 20-21/ V/2013, J.B. Sandberg, B.C. Kondratieff, 2♂, 1♀( JBSC); Union Co., Grande Ronde River, Hwy244, Hilgard Junction State Park( Hwy84), 45.34323°N, 118.24013°W, 22/ V/2013, J.B. Sandberg, B.C. Kondratieff, 2♂, 3♀( JBSC). UTAH, Juab Co., Pole Creek, FR016 below confluence Pole Canyon, 14/ VI/2012, B.C. Kondratieff& R. W. Baumann, 10♂, 23♀( CSUC); Salt Creek, Ponderosa Campground, 14/ VI/2012, B.C. Kondratieff& R. W. Baumann, 2♂, 2♀( CSUC); Utah Co., Nebo Creek, FR405, 13/ VI/2012, B.C. Kondratieff& R.W. Baumann, 1♂, 8♀( CSUC).
3111810344
Logan River
Logan River
12
46
1
1
Utah
holotype
3111810304
V
Ruby Wash & Red Mountain Open Space
Larimer Co.
Sand Creek
14
48
1
2
5
Colorado
3111810341
V
Ruby Wash & Red Mountain Open Space
Larimer Co.
Sand Creek
14
48
1
2
5
Colorado
3111810353
V
Whitehorse Ranch Rd & Fields
Harney Co.
42.17439
Trout Creek
1
-118.43199
Grande Ronde River
14
48
1
1
2
Oregon
3111810329
JBSC, V
J. B. Sandberg & B. C. Kondratieff
Union Co.
45.34323
Hwy
1
-118.24013
Grande Ronde River
14
48
3
1
2
Oregon
3111810323
VI
Pole Creek, FR & Pole Canyon
Juab Co.
Juab Co.
Salt Creek
14
48
1
23
10
Utah
3111810354
VI
Pole Creek, FR & Pole Canyon
Juab Co.
Juab Co.
Salt Creek
14
48
1
23
10
Utah
3111810301
[243,598,1503,1528]
CSUC
Baumann
14
48
4
2
2
Utah
3111810303
VI
Co. & Nebo Creek
Utah
14
48
1
8
1
Utah