Isoperla fulva Claassen 1937 The Isoperla Of California (Plecoptera: Perlodidae); Updated Male Descriptions And Adult Keys For 18 Western Nearctic Species Sandberg, John B. Kondratieff, Boris C. Illiesia 2013 9 5 34 64 6MZWW Claassen Claassen 1937 [998,1279,366,391] Insecta Perlodidae Isoperla Animalia Plecoptera 12 46 Arthropoda species fulva  ( Figs. 6a-d, 20b)     Isoperla fulva Claassen 1937.    Holotype ♂, Utah, Logan River.   Male. Aedeagus: sclerotized posterior process present; body with one posterior lobe, one small dorsal lobe, and one large anterior lobe ( Fig. 6a); one small bulbous patch of spinulae above sclerotized process ( Figs. 6a-c); sclerotized process length approaching 0.5 mmwhen fully extended, blade-like in lateral view ( Fig. 6a), distal margin expanded and slightly membranous in posterior view ( Fig. 6b), and narrow proximal portion of sclerotized process sometimes extended past posterior aedeagal membrane ( Fig. 6a). Abdominal terga 8-9, 9, 9-10: without stout spinulae or long stout setae. Posterolateral margins of at least abdominal segment 8 with scale-like setae clustered in brushes of several setae. Paraprocts: curved dorsally, length if straightened subequal to combined first and second cercal segments, tapering abruptly to blunt apices ( Fig. 6a). Vesicle: rounded lobe, widest at base with broadly rounded apical margin ( Fig. 6d).    Isoperla fulvais most similar to the two other  I. marmoratacomplex species,  I. marmorataand  I. roguensis. These species differ in paraproct shape and aedeagal characters. The paraprocts taper abruptly to blunt apices in  I. fulva, taper gradually to blunt apices in  I. marmorata, and taper abruptly to tiny sharp apices in  I. roguensis( Table 1). The distal margin of blade-like sclerotized processes is thin in  I. marmorata, expanded in  I. fulvaand expanded and variably-shaped in  I. roguensis. The aedeagus of this species was described with an additional pair of long narrow tubular lobes extending from the base of the anterodorsal lobe ( Szczytko & Stewart 1979, fig. 138C). When using the live eversion method, these processes remain inverted ( Fig. 6a); however two small areas in inverted membrane are present on anterior lobe C of this study (not discernible in Fig. 6a). The live everted aedeagus was cleared in KOH, which allowed observation of an inverted pair of tube-like processes within the aedeagus, and openings along the anteroapical margin ( Fig. 20b). The small dorsal lobe B ( Figs. 6a & c) in the current study possesses a pair of minute, closely placed holes from which seminal fluid streamed after live eversion and fixing. These dorsal aedeagal openings require further study before their function can be determined.    Isoperla fulvais putatively known from Californiabased on three malesand one female(El Dorado and Modoc counties - Jewett 1960) and Plumas County ( Szczytko & Stewart 1979) collected in the 1940’s and 1950’s. No recent material (everted males) from Californiacompares well with the Coloradoand Utahspecimens examined for this study, and the California  I. fulvarecords were determined before the description of  I. roguensis. The El Dorado County male was likely collected from Weber Creek, south of Camino, 10 miles( 16 km) north of Placerville, where Sandberg (2011b)collected  I. marmorataand  I. mormona.   Additional Material Examined.  COLORADO, Larimer Co., Sand Creek, Ruby Wash, Red Mountain Open Space, 24/ V/2012, B.C. Kondratieff& D. Givens, 5♂, 2♀( CSUC);  17/ VI/2012, B.C. Kondratieff& R. Stokes, 10♂, 3♀( CSUC).  OREGON, Harney Co., Trout Creek, Whitehorse Ranch Rd 18 mi( 29 km) SE Fields, 42.17439°N, 118.43199°W, 20-21/ V/2013, J.B. Sandberg, B.C. Kondratieff, 2♂, 1♀( JBSC);  Union Co., Grande Ronde River, Hwy244, Hilgard Junction State Park( Hwy84), 45.34323°N, 118.24013°W, 22/ V/2013, J.B. Sandberg, B.C. Kondratieff, 2♂, 3♀( JBSC).  UTAH, Juab Co., Pole Creek, FR016 below confluence Pole Canyon, 14/ VI/2012, B.C. Kondratieff& R. W. Baumann, 10♂, 23♀( CSUC);  Salt Creek, Ponderosa Campground, 14/ VI/2012, B.C. Kondratieff& R. W. Baumann, 2♂, 2♀( CSUC);  Utah Co., Nebo Creek, FR405, 13/ VI/2012, B.C. Kondratieff& R.W. Baumann, 1♂, 8♀( CSUC). 3111810344 Logan River Logan River 12 46 1 1 Utah holotype 3111810304 V Ruby Wash & Red Mountain Open Space Larimer Co. Sand Creek 14 48 1 2 5 Colorado 3111810341 V Ruby Wash & Red Mountain Open Space Larimer Co. Sand Creek 14 48 1 2 5 Colorado 3111810353 V Whitehorse Ranch Rd & Fields Harney Co. 42.17439 Trout Creek 1 -118.43199 Grande Ronde River 14 48 1 1 2 Oregon 3111810329 JBSC, V J. B. Sandberg & B. C. Kondratieff Union Co. 45.34323 Hwy 1 -118.24013 Grande Ronde River 14 48 3 1 2 Oregon 3111810323 VI Pole Creek, FR & Pole Canyon Juab Co. Juab Co. Salt Creek 14 48 1 23 10 Utah 3111810354 VI Pole Creek, FR & Pole Canyon Juab Co. Juab Co. Salt Creek 14 48 1 23 10 Utah 3111810301 [243,598,1503,1528] CSUC Baumann 14 48 4 2 2 Utah 3111810303 VI Co. & Nebo Creek Utah 14 48 1 8 1 Utah