Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda) Wesener, Thomas Enghoff, Henrik Sierwald, Petra ZooKeys 2009 2009-09-04 19 19 1 128 X8B2 Wesener Wesener & Enghoff & Sierwald 2009 [140,516,905,932] Diplopoda Pachybolidae Colossobolus Animalia Spirobolida 34 35 Arthropoda species aculeatus sp. nov.   urn:lsid:zoobank.org:act: FA93B3D8-7AD7-4E8F-93E9-CA4B329D4BB9    Materialexamined: 2 ♂   Holotype: 1 ♂( 122 mmlong), CAS BLF 9555, Madagascar, Province Antsiranana, Forêt de Binara,  7.5 km230° SW Daraina,  375 m, tropical dry forest, 13°15’18”S, 049°37’00”E, leg. B. L. Fisher,  1.XII.2003.   Paratype: 1 ♂, same data as holotype.    Similarspecies:the special U-shaped posterior gonopods are unique for  Colossobolusexcept for  C. pseudoaculeatus sp. n.  C. aculeatusdiffers from the latter in the presence of a sharp-ending preanal process, which is well-rounded in  C. pseudoaculeatus sp. n.(compare Fig. 20Cwith Fig. 21B). The lateral branch of the posterior gonopod telopodite is of equal width all along in  C. aculeatuswhile it is basally slen- der with a swollen tip in  C. pseudoaculeatus sp. n.(compare Fig. 20Ewith Fig. 21E).    Description. Measurements: males with 51 body rings, circa 122 mmlong, 9.5 mmwide. Female unknown.  Colorationaffected by alcohol. Head, antennae, legs and telson red ( Figs 20A–C). Median part of collum dark, margins red ( Fig. 20A). Pro- and anterior part of mesozonites dark brown, posterior part of meso- and metazonites red. Antennaeprotruding back to ring 3. Male coxal processeson coxae 3 and 4 barely visible, on coxae 5–7 absent ( Fig. 20B). Preanal processsharp-edged, but not projecting over anal valves ( Fig. 20C).  Anterior gonopodsternite elongated into a wide, broadly rounded lobe ( Fig. 20D). Mesal process of coxite weakly developed, protruding into a very short wide lobe with   Figure 21.  Colossobolus pseudoaculeatus sp. n., male holotype. Ahead, lateral Btelson Canterior gonopods, posterior view Danterior gonopods, anterior view Eright posterior gonopod, posterior view Fright posterior gonopod, anterior view. Apo = apodeme; av = anal valves; Co = collum; Cx = coxite; ma = membranous area; Md = mandible; Pre = preanal ring; St = sternite; sub = subanal scale; T = telopodite; T-Lbr = telopodite lateral branch; T-mbr = telopodite main branch; Tp = telopodite process. Not to same scale. a well-rounded tip. Lobe only slightly longer than sternite ( Fig. 20D). Telopodite process long, becoming apically thicker than basally. Mesal margin laterally sharp, not projecting ( Fig. 20F).  Posterior gonopodstelopodites positioned parallel to one another, generally Ushaped ( Figs 20E, G). Coxite branch absent ( Fig. 20E). Lateral branch of regular width, tip wide, well-rounded, not thicker than median or basal parts ( Fig. 20E). Central membranous area present as a wide, well-rounded knob ( Fig. 20G). Main branch slightly longer than lateral branch, latero-apically towards lateral branch with a large membranous area (Fig, 20G).    Distributionand ecology:  C. aculeatusis only known from the Forêt Binara ( Fig. 10) where it occurs in close proximity to  C. semicyclus.    Etymology:  aculeatus, adjective, refers to the sharp-edged preanal process, unique for  Colossobolus. 2620669307 2003-12-01 de Binara & B. L. Fisher Madagascar 375 -13.255 Foret de Binara 21 49.616665 Province 34 35 1 1 Antsiranana holotype 2620669301 2003-12-01 de Binara & B. L. Fisher Madagascar 375 -13.255 Foret de Binara 21 49.616665 Province 34 35 1 1 Antsiranana paratype