Review of the Spirobolida on Madagascar, with descriptions of twelve new genera, including three genera of ' fire millipedes' (Diplopoda) Wesener, Thomas Enghoff, Henrik Sierwald, Petra ZooKeys 2009 2009-09-04 19 19 1 128 X8B7 Wesener Wesener & Enghoff & Sierwald 2009 [140,598,694,721] Diplopoda Pachybolidae Colossobolus Animalia Spirobolida 36 37 Arthropoda species pseudoaculeatus sp. nov.   urn:lsid:zoobank.org:act: 2C06B20E-C62D-4CAE-9030-0761193A9DE9    Materialexamined: 1 ♂  Holotype:1 ♂ ( 119 mmlong), ZSM FGZC 1847, Madagascar, Province Antsiranana, Forêt d’Ambre, Provate Reserve Le Fontenay, leg. Köhlerand Franzen,  26.II.2008.    Differentialdiagnosis:the anterior and posterior gonopods of  C. pseudoaculeatus sp. n.can only be confused with those of  C. aculeatus. See the latter for a list of the characters distinguishing these two species.    Description. Measurements: male with 49 body rings, circa 119 mmlong, 9.55 mmwide. Female unknown.  Colorationaffected by alcohol. Head, antennae, legs and telson red ( Figs 21A, B). Median part of collum dark, margins red ( Fig. 21A). Body rings anteriorly dark brown, posteriorly wine-red. Antennaeprotruding back to ring 4. Male coxal processespresent on coxae 3 and 4, on coxae 5–7 absent. Preanal processwell-rounded, not projecting over anal valves ( Fig. 21B).  Anterior gonopodsternite elongated into a wide, broadly rounded lobe ( Fig. 21D). Mesal process of coxite weakly developed, protruding into a very short wide lobe with a well-rounded tip; lobe only slightly longer than sternite ( Fig. 21D). Telopodite process long and well-rounded, apically thicker than basally. Mesal margin laterally sharp, not projecting ( Fig. 21C).  Posterior gonopodstelopodites positioned parallel to one another, generally Ushaped ( Figs 21E, F). Coxite branch absent ( Fig. 21E), lateral branch basally slender, width apically increasing ( Figs 21E, F); tip swollen, slightly curved mesally ( Fig. 21E). Central membranous area present as a wide, well-rounded knob ( Fig. 21E). Main branch slightly shorter than lateral branch, latero-apically towards lateral branch with a large membranous area ( Figs 21E, F).    Distributionand ecology:  C. pseudoaculeatusis only known from the Forêt d’Ambre ( Fig. 10).    Etymology:  pseudoaculeatus, adjective, refers to the close similarity of the species to  Colossobolus aculeatus. 2608818273 2008-02-26 ZSM Kohler Madagascar Provate Reserve Le Fontenay 36 37 FGZC 1847 1 Antsiranana holotype