Hennig, 1935 : 52 A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae) Ferro, Gustavo Borges Marshall, Stephen A. Zootaxa 2018 2018-06-07 4429 3 401 458 5WZNV Hennig, 1935 Hennig 1935 [151,607,1249,1275] Insecta Micropezidae Cardiacephala GBIF Animalia Diptera 46 447 Arthropoda species secunda     Cardiacephala secunda  Hennig, 1935: 52.   Description:Body length 6 mm, wing 4 mm. Palpus mostly dark brown, white on outer face. Clypeus black with sparse and pale microsetulae. First flagellomere dark brown covered with pale microsetulae. Frontal vitta dark brown and with a black spot between the anterior pair of frontal setae. Fronto-orbital plates shiny. Row of katepisternal setae black. Mid and hind femur narrowly yellow basally and apical third white. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape black. Epandrium yellow. Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Frontal vitta with a narrowed depression centrally, reaching ocellar triangle. Inner vertical seta insertion flat, without tubercle. Vertex rounded. Paracephalon smooth, without swelling (not extending beyond the vertex). Line of black supracervical setae strong. Thorax: Mesonotum with white pruinosity. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing mostly black with rounded hyaline spots, discal band with four hyaline spots and apical with three ( Fig. 82, 84). R4+5 and M1+2 ending separately in wing margin. A1+CuA2 as long as CuA2. Female cervical sclerite smooth, without swelling. Abdomen: Tergites with brownish setulae. Female abdomen: Single spermathecal duct uniformly thick, ending in a sac-like spermatheca (tapered basally and distally swollen). Paired spermathecal duct divided into a narrow basal half and a broader distal half. Paired spermathecal duct stems thin and smooth in basal third, followed by a small rounded swelling and a small appendix ( Fig. 83). Paired spermathecae spherical, somewhat flattened basally and deeply invaginated from apex to base. Male abdomen: Ejaculatory apodeme bigger than epandrium and with a broad blade, wider than sperm pump ( Fig. 85). Phallapodeme extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium, ending in a phallic bulb, phallic bulb subequal in length and width.    Typematerial: Holotype( 1 ♀, MLUH, not examined): ECUADOR. Cordillera. Date and collector not cited.  Material examined: ECUADOR. Pichincha, Maquipucuna Biol. Res., river trail, 1200 m, 0º7’34” N 78º37’57” W, 28.IV.2002, S. A. Marshall ( 3 ♀, 5 ♂, DEBU00178603, DEBU00178617, DEBU00178628, DEBU00178630, DEBU00178631, DEBU00178638, DEBU00195845, DEBU00195848); Pichincha, Maquipucuna Biol. Res., river trail, 1200 m, 0º7’34” N 78º37’57” W, 28.IV.2002, S. A. Marshall ( 1 ♂, DEBU00178614, sequence MYCRO261–16); Pichincha, Maquipucuna Biol. Res., main trail, 1500 m, 27.IV.2002, S. A. Marshall ( 1 ♀, DEBU, sequence MYCRO260–16); Pichincha, Maquipucuna Biol. Res., river trail, 1200 m, 0º7’34” N, 78º37’57” W, 27.X.1999, S. A. Marshall ( 1 ♀, DEBU00100042); Pich., Tinalandia, 800 m, II.1983, Malaise trap, M. Sharkey ( 1 ♀, DEBU).   Distribution: Ecuador.   Comments:  Cardiacephala secundais one of the most distinctive species in  Cardiacephaladue to the striking black wing with rounded hyaline spots (four on discal and three on apical band). Hennig (1934: 107, Fig. 2)provides an illustration of a  C. secundahead.