Hendel, 1933 : 217 Plocoscelus conifer (Hendel) Hennig 1937 : 49 Hennig 1935 : 51 Hennig 1937 : 49 A revision of the Neotropical ant-like genus Cardiacephala Macquart, including Plocoscelus Enderlein syn. nov. (Diptera: Micropezidae, Taeniapterinae) Ferro, Gustavo Borges Marshall, Stephen A. Zootaxa 2018 2018-06-07 4429 3 401 458 8BBWV Hendel, 1933 Hendel 1933 [151,607,1756,1782] Insecta Micropezidae Cardiacephala GBIF Animalia Diptera 23 424 Arthropoda species conifera     Cardiacephala conifera  Hendel, 1933: 217.    Plocoscelus conifer(Hendel),  Hennig 1937: 49.    Plocoscelus haedulus  Hennig 1935: 51,  Hennig 1937: 49.   Description:Body length 8 mm, wing 5 mm. Palpus mostly dark brown, white on outer face. Clypeus brown with pale microsetulae. First flagellomere brown and covered with pale microsetulae. Row of katepisternal setae brownish. Fore tibia dark brown. Hind femur narrowly white basally. Fore tarsus dark brown with gold/whitish pubescence ventrally at base of tarsomere one. Oviscape brown with apex dark brown. Epandrium yellow. Head: Palpus broadened basally and gradually tapering to apex. Clypeus short, length less than half of width. Two frontal setae. Vertex flattened. Frontal vitta slightly elevated until vertex. Inner vertical setae inserted in strong tubercles. Paracephalon swollen, conical. Black supracervical setae sparse, weak.   FIGURES 33–36.  Cardiacephala conifera.33, female, dorsal view; 34, female genitalia, lateral view; 35, female, lateral view; 36, male genitalia, lateral view. Thorax: Mesonotum with golden pruinosity. Notopleural setae strong. Posterior notopleural seta insertion slightly protuberant. Scutellum upturned, triangular. Femora with sparse black setulae. Mid and hind tibia flattened, but sulcus on outer face weak. Wing with incomplete apical pigmentation and three preapical darkened spots. Crossvein r-m with dark spot. R4+5 and M1+2 ending separately in wing margin. A1+CuA2 less than half as long as CuA2. Female cervical sclerite smooth, without swelling. Abdomen: Tergites with sparse brownish setae. Female abdomen: Single spermathecal duct apically broad, basal half slightly tapered. Single spermatheca cylindrical. Paired spermathecal duct divided into a narrow basal half and a broader distal half. Paired spermathecal duct stems thin and smooth on basal third, apical two thirds with a large rounded swelling ( Fig. 34). Paired spermathecae spherical, somewhat flattened basally (almost disc-shaped) and deeply invaginated from apex to base. Male abdomen: Ejaculatory apodeme reduced, blade small and short, sperm pump wider than blade. Phallapodeme extending to anterior limit of hypandrium. Distiphallus long, longer than epandrium, ending in a phallic bulb, phallic bulb subequal in length and width.    Typematerial: Syntypesof  C. coniferaHendel( 2 ♀, NMW, probably lost):  PARAGUAY. Aregua. No information about collector and date;  Plocoscelus haedulusHennig( 1 ♀, MLUH): BRAZIL, Amazonas, Tefé. No information about collector.  Material examined: BRAZIL. Amazonas, 45 kmE Tapuruquara, 23.I.1978, Norman D. Penny ( 1 ♂, INPA); Amazonas, Manaus, Reserva Ducke, 9.IX.1986, L.S. Aquino & U. Barbosa ( 1 ♀, INPA); Amazonas, Manaus, Est. AM 1 km64, 28.VII.1970, A. Faustino ( INPA); Amazonas, Reserva Ducke, 14.II.1978, Jorge Arias ( INPA); Mato Grosso do Sul, Bodoquena, Fazenda California, 20º41’55.9” S 58º52’49.4” W, 6.XI.2011( 1 ♀, DZUP341592, sequence MYCRO237–16); COLOMBIA. Bolivar, SFF Los Colorados, La Suiris, 9º54’ N 75º07’ W, 126 m, 16– 30.XI.2000, E. Deulufeut M. 921 ( 1 ♀, IAVH); Magdalena, Tayrona Natl. Pk., Pueblito, 225 m, malaise trap, 4– 20.XII.2000, M. 965, R. Henriquez ( 1 ♀, IAVH); Magdalena, PNN Tayrona Pueblito, 225 m, 11º20’ N 74º02’ W, 1–19.IX.2000, M. 626, R. Henriquez ( 1 ♀, USNM); Vichada, PNN Tuparro CentroAdministrativo, 5º21’ N 67º51’ W, 100 m, Malaise, 17–26.XII.2000, M. 1382, W. Villalba ( 1 ♀, USNM).   Distribution: Brazil, Colombia, Guiana,  Paraguay.   Comments:The swelling on the inner vertical seta insertion and the paracephalon, in combination with three preapical dark spots and short A1+CuA2 on the wing, makes this species easily distinguishable. Hendel (1936: 65, Fig. 1)also provides an illustration of a  C. coniferahead.