Thoracostrongylus formosanus Shibata, 1982: 71 Thoracostrongylus malaisei Scheerpeltz, 1965 Thoracostrongylus malaisei Scheerpeltz, 1965: 245 Taxonomy of the genus Thoracostrongylus Bernhauer (Coleoptera: Staphylinidae) with descriptions of five new species from China Yang, Zhuo Zhou, Hong-Zhang Schillhammer, Harald Journal of Natural History 2011 2011-02-08 45 7 - 8 407 433 56KHB Shibata, 1982 Shibata 1982 [374,871,1223,1247] Insecta Staphylinidae Thoracostrongylus Animalia Coleoptera 19 424 Arthropoda species formosanus  ( Figure 8A–G)      Thoracostrongylus formosanus  Shibata, 1982: 71( typelocality: Taiwan, near Tsuifeng, about  2200 malt., Nantou Hsien).  Smetana and Davies, 2000: 39(checklist); Herman, 2001: 3556(catalogue); Smetana, 2004: 686(catalogue of Palaearctic region).   Material   9 males, 6 females, CHINA: Sichuan, south of Danba County, Wujiapeng village[30 ◦52’N, 101 ◦53’E],  2620 m, pitfall trap,  2001.VII.19–21, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  13 males, 8 females, Sichuan, south of Danba County, Wujiapeng village[30 ◦52’N, 101 ◦53’E],  2410 m, pitfall trap,  2001.VII.19–21, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  8 males, 1 female, Sichuan, south of Danba County, Wujiapeng village[30 ◦52’N, 101 ◦53’E],  2320 m, pitfall trap,  2001.VII.19–21, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  2 males, Sichuan, Danba County, Xigeshenzha town[30 ◦52’N, 101 ◦53’E],  2620 m, pitfall trap,  2001.VII.20 –22. Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  1 female, Sichuan, Danba County, Xigeshenzha town[30 ◦52’N, 101 ◦53’E],  2640 m, pitfall trap,  2001.VII.20–22, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  2 males, 5 females, Sichuan, Danba County, Xigeshenzha town[30 ◦52’N, 101 ◦53’E],  2610 m, pitfall trap,  2001.VII.20–22, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  1 male, Sichuan, Wolong, Gengda[30 ◦20’N, 101 ◦25’E],  1515 m, pitfall trap,  2004.VI.20–23, Xiaodong Yucoll. (IZCAS);  1 male, 1 female, Hunan, Shennonggu, Jiaoshi[26 ◦49’N, 114 ◦04’E],  880 m,  2008.VII.6. Zhuo Yangcoll. (IZCAS);  1 male, Sichuan, Baoxing, Muping[30 ◦22’N, 102 ◦48’E], pitfall trap,  1740 m,  2003.VIII.2–5, Xiaodong Yuand Jie Wucoll (IZCAS);  14 males, 19 females, Sichuan, Baoxing, Fengtongzhai[30 ◦35’N, 102 ◦52’E],  1997.VI.9. Hongzhang Zhoucoll. (IZCAS);  8 males, 6 females, Sichuan, Baoxing, Fengtongzhai[30 ◦35’N, 102 ◦52’E],  1997.VI.8. Haisheng Zhoucoll. (IZCAS);  1 male, Sichuan, Baoxing, Fengtongzhai[30 ◦35’N, 102 ◦52’E],  1795 m, pitfall trap,  2001.VI.30–VII.3, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  3 females, Sichuan, Baoxing, Fengtongzhai[30 ◦35’N, 102 ◦52’E],  1680 m, pitfall trap,  2001.VI.30–VII.3, Xiaodong Yuand Hongzhang Zhoucoll. (IZCAS);  6 males, 19 females, Zhejiang, Longquan City, Fengyang Mountain[27 ◦55’N, 119 ◦11’E],  1400 m, pitfall trap,  2007.VII.28–31, Zongyi Zhaoand Zhuo Yangcoll. (IZCAS);  5 males, 2 females, Zhejiang, Longquan City, Fengyang Mountain[27 ◦55’N, 119 ◦11’E],  1400 m,  2007.VIII.1, Zongyi Zhaocoll. (IZCAS);  1 female, Zhejiang, Longquan City, Fengyang Mountain[27 ◦55’N, 119 ◦11’E],  1400 m, Malaise trap,  2007.VII.31, Zongyi Zhaocoll. (IZCAS);  2 females, Zhejiang, Longquan City, Fengyang Mountain[27 ◦55’N, 119 ◦11’E],  1400 m,  2007.VII.30, Zongyi Zhaocoll. (IZCAS);  1 male, Fujian,  29.5.2005, 2 km SE Xinqiao[27 ◦05’N, 117 ◦1’E], pitfall trap, leg. J. Turna(NMW);  1 male, 2 females, Fujian, 8.5–3.6. E’mei Feng,  1500–1600 m, 27 ◦01’N 117 ◦04’E, 2008, leg. J. Turna(NMW);  1 male, Sichuan, Str.v. PINGWU nach NANPING MAONIU SHAN, Sdseite  2420–2650 m, leg. A. PUCHNER,  29.V.–5.VI.2006(NMW);  1 male, Hubei, Mufu Shan Jiugongshan forestpark [29 ◦41’N, 114 ◦6’E], up to  1000 m, 3. /5. /  18.6.2002, leg. J. Turna(NMW);  2 males, 1 female, Fujian, Wuyi Mountain, [KUATUN] (  2300 m), 27.40 n. Br. 117.40 ö. L., J. Klapperich, 1938 ( Fukien) (NMW);  1 male, Fujian, Wuyi Mountain, [KUATUN, FUKIEN], 7.4.46 (TSCHUNG SEN.) (NMW);  3 males, Fujian, Wuyi Mountain, [KUATUN, FUKIEN], 14.5.46, leg. Tschung-Sen(NMW);  1 male, Fujian, Wuyi Mountain, [KUATUN, FUKIEN], 9.7.46 (TSCHUNG SEN.) (NMW);  1 female, Fujian,  7.5.2005, Fengshui Guan[27 ◦9’N, 117 ◦85’E],  1700 m, leg. J. Turna(NMW);  1 male, 1 female, Fujian, 8–9.5, E’mei Feng,  1200–1500 m, 27 ◦01’N 117 ◦04’E, 2008, leg. J. Turna(NMW);  1 male, 3 females, TAIWAN: Hualien, Pilu, alt.  2100 m, 24 ◦10’58”N 121 ◦23’16”E,  06.V.2006, pitfall trap, with carrions, leg. Y.F. Hsu(NMW).  Figure 7.  Thoracostrongylus birmanus, ( Fauvel, 1895), male and female from Hainan and Yunnan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) 10th tergite, female; (E) genital segment with second gonocoxite, female; (F) aedeagus, ventral view; (G) aedeagus, lateral view. Scale bar 1 mm.  Figure 8.  Thoracostrongylus formosanus Shibata, 1982, male and female from Sichuan, Zhejiang, Hunan, Fujian, Hubei and Taiwan, China. (A) 8th sternite, male; (B) 10th tergite, male; (C) 9th sternite, male; (D) genital segment with second gonocoxite, female; (E) 10th tergite, female; (F) aedeagus, ventral view; (G) aedeagus, lateral view. Scale bar 1 mm.  Description  Measurements. Body length: 9.2 –9.4 mm. ♂(mean, n = 5): HL: 1.35; HW: 1.83; CL: 0.88; PO: 0.25; PL: 1.73; PW: 1.50; EL: 2.05; EW: 2.20; ELS: 1.00. ♀(mean, n = 5):  HL: 1.33; HW: 1.83; CL: 0.83; PO: 0.28; PL: 1.80; PW: 1.55; EL: 2.30; EW: 2.45; ELS: 1.15.  Distribution   China( Sichuan, Zhejiang, Hunan, Fujian, Hubei, Taiwan).    Thoracostrongylus malaisei Scheerpeltz, 1965 ( Figure 9A–G)     Thoracostrongylus malaisei  Scheerpeltz, 1965: 245( typelocality: NE Burma, Kambaiti,  7000 ft.).  Smetana and Davies, 2000: 39(checklist); Herman, 2001: 3556(catalogue); Smetana, 2004: 686(catalogue of Palaearctic region; China, Yunnan).   Type material    Paratypes(labelled as “Cotypus”). BURMA: 1 male, N.E. Burma, Kambaiti,  2000 m,  27.5.1934, Malaise, Excoll. Scheerpeltz(NMW);  1 female, N.E. Burma, Kambaiti,  7000 m,  12.5.1934, R. MALAISE, Excoll. Scheerpeltz(NMW).  Additional materal.  1 female, CHINA: Yunnan, 14–21.6, 100 kmW Baoshan, 1993, Gaoligongshan Nat. Res., E. Jendek & O. Sausaleg. (NMW).  Description  Measurements. Body length: 9.5–9.7 mm. ♂: HW: 1.85; HL: 1.40; CL: 0.85; PO: 0.30; PW: 1.45; PL: 1.52; EW: 2.05; EL: 2.00; ELS: 1.05. ♀(mean, n = 2): HW: 2.03; HL: 1.48; CL: 0.98; PO: 0.28; PW: 1.58; PL: 1.90; EW: 2.28; EL: 2.18; ELS: 1.20.  Distribution  China( Yunnan); Myanmar. [486,679,1360,1384] Taiwan 2200 Tsuifeng 19 424 1 Nantou 2001-07-19 2001-07-21 2001-07-19 Xiaodong Yu & Hongzhang Zhou China Danba County 2620 Wujiapeng village 19 424 15 6 9 Sichuan 2001-07-19 2001-07-21 2001-07-19 Xiaodong Yu & Hongzhang Zhou China Danba County 2410 22 427 Wujiapeng village 19 424 21 8 13 Sichuan 2001-07-19 2001-07-21 2001-07-19 Xiaodong Yu & Hongzhang Zhou China Danba County 2320 Wujiapeng village 22 427 9 1 8 Sichuan 2001-07-20 2001-07-22 2001-07-20 Xiaodong Yu & Hongzhang Zhou China Danba County 2620 Xigeshenzha town 22 427 2 2 Sichuan 2001-07-20 2001-07-22 2001-07-20 Xiaodong Yu & Hongzhang Zhou China Danba County 2640 Xigeshenzha town 22 427 1 1 Sichuan 2001-07-20 2001-07-22 2001-07-20 Xiaodong Yu & Hongzhang Zhou China Danba County 2610 Xigeshenzha town 22 427 7 5 2 Sichuan 2004-06-20 2004-06-23 2004-06-20 Xiaodong Yu China 1515 Gengda Wolong 22 427 1 1 Sichuan 2008-07-06 Zhuo Yang China 880 Jiaoshi Shennonggu 22 427 2 1 1 Hunan 2003-08-02 2003-08-05 2003-08-02 Xiaodong Yu & Jie Wu China 1740 Muping Baoxing 22 427 1 1 Sichuan 1997-06-09 Hongzhang Zhou China Fengtongzhai Baoxing 22 427 33 19 14 Sichuan 1997-06-08 Haisheng Zhou China Fengtongzhai Baoxing 22 427 14 6 8 Sichuan 2001-06-30 2001-07-03 2001-06-30 Xiaodong Yu & Hongzhang Zhou China 1795 Fengtongzhai Baoxing 22 427 1 1 Sichuan 2001-06-30 2001-07-03 2001-06-30 Xiaodong Yu & Hongzhang Zhou China 1680 Fengtongzhai Baoxing 22 427 3 3 Sichuan 2007-07-28 2007-07-31 2007-07-28 Zongyi Zhao & Zhuo Yang China 1400 Fengyang Mountain Longquan City 22 427 25 19 6 Zhejiang 2007-08-01 Zongyi Zhao China 1400 Fengyang Mountain Longquan City 22 427 7 2 5 Zhejiang 2007-07-31 Zongyi Zhao China 1400 Fengyang Mountain Longquan City 22 427 1 1 Zhejiang 2007-07-30 Zongyi Zhao China 1400 Fengyang Mountain Longquan City 22 427 2 2 Zhejiang 2005-05-29 SE Xinqiao & J. Turna China Fujian 22 427 1 1 Fujian J. Turna China 1550 Feng 22 427 3 2 1 Fujian 2006-05-29 2006-06-05 2006-05-29 China 2535 Sdseite Str. 22 427 1 1 Sichuan 2002-06-18 J. Turna China 1000 Mufu Shan Jiugongshan forest 22 427 1 1 Hubei Br. & L., J. Klapperich & Fukien China 2300 27.4 Wuyi Mountain 742 -117.4 22 427 3 1 2 Fujian China Wuyi Mountain 22 427 1 1 Fujian Tschung-Sen China Wuyi Mountain 22 427 3 3 Fujian China Wuyi Mountain 22 427 1 1 Fujian 2005-05-07 Fengshui Guan & J. Turna China 1700 Fujian 22 427 1 1 Fujian J. Turna China 1350 Feng 22 427 2 1 1 Fujian 2006-05-06 Y. F. Hsu Taiwan 2100 Pilu 22 427 4 3 1 Hualien [144,812,330,354] China Sichuan 23 428 1 Sichuan [477,752,513,537] Myanmar 2134 Kambaiti 23 428 1 1934-05-27 N. E. Burma & Malaise & Ex & Scheerpeltz Myanmar 2000 Kambaiti 23 428 1 1 paratype 1934-12-05 N. E. Burma & Ex & Scheerpeltz Myanmar 7000 Kambaiti 23 428 1 1 paratype Gaoligongshan Nat. Res., E & O. Sausa China Baoshan 23 428 1 1 Yunnan