Revision and cladistic analysis of the Afrotropical endemic genus Smeringopus Simon, 1890 (Araneae: Pholcidae) HUBER, BERNHARD A. Zootaxa 2012 2012-09-07 3461 1 1 138 4XVJ4 Huber, 2012 Huber 2012 [151,440,1719,1745] Arachnida Pholcidae Smeringopus Animalia Araneae 18 19 Arthropoda species mgahinga sp. nov.   Figs. 25, 32–33, 46–49, 73–93 Type. Male holotypefrom Uganda, Kisoro District, Mgahinga Gorilla National Park, Garama Cave, cave entrance ( 1°21.4’S, 29°37.9’E),  2500 ma.s.l.,  25.xi.2010( B.A. Huber), in ZFMK( Ar 8516). Etymology. The name is a noun in apposition, derived from the typelocality. Diagnosis. Distinguished from similar congeners (other species of the  rubrotinctusgroup) by shapes of procursus and embolus ( Figs. 73–75, 78, 79) and by strongly protruding posterior epigynal plate ( Fig. 47); from most other congeners also by long apophyses on male chelicerae ( Figs. 25, 76, 77).   FIGURES 73–80.  Smeringopus mgahinga. 73. Left cymbium and procursus, retrolateral view. 74–75. Left procursus, ventral and dorsal views. 76–77. Male chelicerae, lateral and frontal views. 78–79. Left embolus, prolateral and dorsal views. 80. Cleared female genitalia, dorsal view. Scale lines: 0.3 mm (73–75, 78–80), 0.5 mm (76–77). Male ( holotype). Total body length 6.2, carapace width 2.1. Leg 1: 56.2 (13.7 + 0.9 + 14.0 + 23.9 + 3.7), tibia 2: 10.3, tibia 3: 8.1, tibia 4: 10.9; tibia 1 L/d: 69. Habitus similar  S. ruhiza(cf. Fig. 20). Carapace with wide median and lateral brown bands, clypeus with pair of wide, indistinct brown bands, sternum brown, leg femora and tibiae with indistinct subdistal rings and light tips, abdomen dorsally with indistinct pattern, ventrally with three dark bands behind gonopore separated by narrow light lines. Distance PME-PME 175 µm, diameter PME 185 µm, distance PME-ALE 70 µm, distance AME-AME 60 µm, diameter AME 125 µm. Ocular area slightly elevated, secondary eyes with indistinct ‘pseudo-lenses’; deep but small thoracic pit ( Fig. 82). Chelicerae as in Figs. 76 and 77, with pair of strong apophyses; without modified hair at tip of apophysis ( Fig. 87). Palps as in Figs. 32 and 33, coxa with indistinct bulge, trochanter barely modified, femur with deep retrolateral furrow with distinct rim proximally ( Fig. 84), ventral end of rim with apophysis, cymbium without projection near tarsal organ, procursus with distinctive tip with bifid ventral apophysis and whitish prolateral process ( Figs. 73–75, 83, 84), bulb with relatively simple bifid embolus ( Figs. 78, 79, 85). Legs without spines, few vertical hairs, with curved hairs mainly on tibiae and metatarsi 1 and 2, but also some on femora and metatarsi and on legs 3 and 4, on tibiae and metatarsi also dorsally; retrolateral trichobothrium on tibia 1 at 2.5%; prolateral trichobothrium present on tibia 1. Gonopore with two epiandrous spigots ( Fig. 91).   FIGURES 81–93.  Smeringopus mgahinga. 81–82. Male prosoma, frontal and dorsal views. 83–84. Right palp, retrolateral and retrolatero-dorsal views. 85. Left procursus and embolus, prolateral view. 86. Tip of right procursus. 87. Tip of male cheliceral apophysis. 88. Trichobothria on male tibia 1. 89. Male tarsus 1 pseudosegmentation. 90. Male tarsus 1 claws. 91. Male gonopore. 92. Epigynum. 93. Female ALS. Scale lines: 20 µm (86), 30 µm (87, 93), 40 µm (89), 50 µm (90), 100 µm (85, 88, 91), 200 µm (83, 84, 92), 500 µm (82), 600 µm (81). Variation. Tibia 1 in11 other males: 8.8–15.1 (mean 12.4). The cheliceral apophyses vary slightly in width and length. Female. In general similar to male; tibia 1 in 14 females: 8.1–13.9 (mean 11.4). Epigynum anterior plate with indentation, without pockets ( Figs. 46, 48); posterior plate strongly projecting ( Fig. 47); internal genitalia as in Figs. 49and 80. ALS with eight spigots each ( Fig. 93). Distribution. Known from southwestern Ugandaand from a female from CongoD.R. ( Nord-Kivu) assigned tentatively ( Fig. 58).   Materialexamined. UGANDA:  Western Region: Kisoro District, Mgahinga Gorilla N.P., Garama Cave: 1♂ holotypeabove; same data, 4♂ 4♀in ZFMK( Ar8517); same data, 2 juvs in pure ethanol, in ZFMK( Uga105).  Mgahinga Gorilla N.P., Gorge(1°22.9’S, 29°35.8–36.0’E),  2900–3000 ma.s.l.,  25.xi.2010( B.A. Huber), 3♂ 3♀in ZFMK( Ar8518-19); same data, 1♀2 juvs in pure ethanol, in ZFMK( Uga146).  Mgahinga Gorilla N.P., bamboo forest (~1°22.0–22.3’S, 29°36.3–36.9’E),  2500–2600 ma.s.l.,  25.xi.2010( B.A. Huber), 1♂ 1♀in ZFMK( Ar8520).  Kabale District, Ruhiza, Bwindi ImpenetrableN.P. (~ 1°02’S, 29°46’E), ~  2300 ma.s.l., roadcut,  25.ix.1996( C.E. Griswold), 2♂ 1♀in CAS; same data, on buildings and roadcut,  12.–16.ix.1996(C.E. Griswold), 1♂ 4♀2 juvs in CAS. Assigned tentatively:  CONGOD.R.:  Nord-Kivu Prov.: Bikara,  18 kmS Lubéroon route Lubéro-Goma[ 0°15’S, 29°12’E],  1200 ma.s.l.,  xii.1976( M. Lejeune), 1♀in MRAC(159843). 3747820727 2010-11-25 ZFMK B. A. Huber Uganda 2500 -1.3566667 Mgahinga Gorilla National Park 130 29.631666 Mgahinga Gorilla National Park 18 19 Ar 8516 1 Kisoro District holotype 3747820470 ZFMK Mgahinga Gorilla N. P. & Ar Uganda Material Garama Cave Western Region 21 22 9 4 5 Kisoro holotype 3747820948 [798,1432,439,463] ZFMK Uga Uganda Material Garama Cave Western Region 21 22 1 Kisoro holotype 3747820523 2010-11-25 ZFMK B. A. Huber Uganda 2950 -1.3816667 Gorge 21 22 6 3 3 Kisoro holotype 3747820443 [401,1065,511,535] 2010-11-25 ZFMK Uga Uganda 2950 -1.3816667 Gorge 21 22 1 1 Kisoro holotype 3747820563 2010-11-25 ZFMK B. A. Huber Uganda 2550 21 22 2 1 1 Kisoro holotype 3747820314 1996-09-25 CAS C. E. Griswold Uganda 2300 -1.0333333 Bwindi Impenetrable 1308 29.766666 Ruhiza 21 22 3 1 2 Kabale holotype 3747820509 1996-09-12 1996-09-16 1996-09-12 CAS Griswold Uganda 2300 -1.0333333 Bwindi Impenetrable 1308 29.766666 Ruhiza 21 22 5 4 1 Kabale holotype 3747820761 1976-12 R M. Lejeune Democratic Republic of the Congo 1200 -0.25 CONGO 1308 29.2 Bikara 21 22 1 1 Nord-Kivu