Hemigymnochaeta laticeps Zumpt, 1953c: 490
The Calliphoridae of Namibia (Diptera: Oestroidea)
Kurahashi, Hiromu
Kirk-Spriggs, Ashley H.
Zootaxa
2006
2006-09-28
1322
1
1
131
4V69
1240433
Corti, 1895: 142
Corti
142
1895
[352,751,411,436]
Insecta
Calliphoridae
Hemigymnochaeta
Animalia
Diptera
66
67
Arthropoda
genus
TYPESPECIES: Hemigymnochaeta luteaCorti, 1895[= Ochromyia unicolorBigot, 1888], by monotypy. NOTES: A large genus restricted to the Afrotropical Region; species of which develop in various kinds of organic matter, such as mushrooms, dung and faeces. Some species drop their eggs in and between the openings of temporary nests of driver ants (Zumpt 1953c: 481). Engel and Cuthbertson (1937: 10, as Tricyclea resurgens) and Ferrar (1987: 85–86) note females of H. apiciferaCurran, 1931, collected at fungus material removed by termites ( Isoptera) from their nest in Zimbabwe. The Namibian species appear to be primarily associated with the fungus gardens of termites and the fruiting bodies of associated fungi.
Hemigymnochaeta bequaertiCurran, 1931a: 14. Figs. 41, 63.
TYPE LOCALITY: Liberia.
DISTRIBUTION: Widespread West Africa to southern Africa: Democratic Republic of Congo, Kenya, Liberia, Namibia*, Nigeriaand South Africa(Natal, Transvaal).
MATERIAL: 4♀, Salambalacampsite, 1–3.iii.2001, KirkSpriggs(1) ( HT)faeces; 2♀, same except: 3–4.iii.2001, KirkSpriggs(1) ( HT)fish; 2♂, 17♀, same except: 26–28.ii.2001, Marais& Kirk Spriggs(1) ( HT)fruit; 2♂, 4♀, same except: 8–10.ii.2004, KirkSpriggs(1) ( HT)fruit; 1♂, 5♀, same except: ( MT); 1♂, same except: 22–24.ii.2001, KirkSpriggs(1) & Marais( MT); 1♂, same except: 1–4.iii.2001, KirkSpriggs(1) ( PT)baited millipede; 2♀, Bum Hillcampsite ( Kwando River), 10–13.ii.2004, KirkSpriggs(1) ( HT)fruit; 1♀, same except: ( MT); 1♀, Toggenburg591 campsite, 18–20.ii.2004, KirkSpriggs(1) ( HT)fruit; 4♂, 2♀, Popa Fallsrestcamp, 13–16.ii.2004, KirkSpriggs(1) ( MT), riverine forest; 1♀, same except: ( HT)fruit. NOTES: Biology and immature stages unknown. Reared from mushrooms collected near Agatha, South Africaby Zumpt (1953c: 488). In Namibiacollected in hanging traps baited with fermenting fruit bait, rotting fish and faeces, in Malaise traps, and in a pitfall trap baited with a dead millipede. Northeastern Namibia; in the ‘arid’ and ‘mesic’ savanna biome ( Fig. 63). Most abundant in February ( vide Table 2).
Hemigymnochaeta laticepsZumpt, 1953c: 490. Figs. 42, 64.
TYPE LOCALITY: Rhodesia [= Zimbabwe].
DISTRIBUTION: Botswana, Namibia*, South Africa(Transvaal), Uganda, Upper Voltaand Zimbabwe.
MATERIAL: 3♀, Salambala campsite, 8–10.ii.2004, KirkSpriggs(1) ( HT) fruit; 2♀, same except: ( MT). NOTES: Biology, life history and immature stages unknown. In Namibiacollected in a Malaise trap and a hanging trap baited with fermenting fruit. Northeastern Namibia; in the ‘mesic’ savanna biome ( Fig. 64). Recorded in February ( vide Table 2).
Hemigymnochaeta maraisi sp. nov. Figs. 15–20, 39, 65.
TYPE LOCALITY: Namibia.
DISTRIBUTION: Katima Mulilo and Tsumkwe Districts of northeastern Namibia, in the ‘mesic’ savanna biome ( Fig. 65).
MATERIAL: videdescription, p. 28. Hemigymnochaeta trichaeta sp. nov. Figs. 21–26, 40, 66. TYPE LOCALITY: Namibia.
DISTRIBUTION: Bwabwata National Park of northeastern Namibia, in the ‘mesic’ savanna biome ( Fig. 66).
MATERIAL: videdescription, p. 32. Hemigymnochaeta unicolor(Bigot, 1888: 608) ( Ochromyia). Figs. 27–32, 67. TYPE LOCALITY: Sierra Leone.
DISTRIBUTION: Widespread in Afrotropical Region: Benin, Botswana, Democratic Republic of Congo, Ethiopia, Gambia, Ghana, Kenya, Liberia, Madagascar, Malawi, Mozambique, Namibia, Nigeria, Sierra Leone, Somalia, South Africa( Cape, Natal, Transvaal), Togo, Uganda, Zambia* and Zimbabwe. PREVIOUS RECORDS: Grootfontein [ 19°57'S, 18°12'E]; Okakarara [ 20°58'S, 17°43'E]; Omatjette [ 21°05'S, 15°05'E]; Epukiro [ 21°07'S, 19°12'E] (Zumpt 1953c: 488).
KNOWN RECORDS: Grootfontein [ 19°33'S, 18°05'E], 30.iv.1950; Omatjette [ 21°05'S, 15°05'E], 11.iv.1950(NMSA)
MATERIAL: 1♂, Ondundozonananandana Mts., 16.v–22.vi.1986, Griffin(1) ( PT); 1♂, 1♀, OkaukuejoDuikersdrinkrds. junction on, 15.v–16.vi.1986, Griffin(1) ( PT), H65447; 2♂, 1♀, Etosha Game Parkat: 20.v–18.vi.1986, Griffin(1) ( PT), H65568/H65569; 3♀, WolfnesOkondekards., W of junction of, 16.v–12.vi.1986, Griffin(1) ( PT) H65403; 2♂, 4♀, Ghaub47, 16–26.vii.1986, Irish(1) ( MT); 1♀, Sturmfeld252, iii.1986, Tölken, H65186; 1♂, Tobiroen, 15.v.1986, Curtis; 1♂, 1♀, Dorsland, 14.v–16.vi.1986, Griffin(1) ( PT); 1♀, HererolandOosat: 20.i.1984, Irish(1), H58092; 1♂, Okahandja6.xii.[19]72, Gaerdes, H25130; 1♀, Sprokieswoud, 7.viii–15.ix.1987, Griffin(1) ( PT); 1♀, Waterberg Plateau Park(2), 2.ii–1.iii.1993, Pusch( PT); 2♂, 11♀, Ruacana area, 14.iii.1997, Koch( MT); 3♂, 2♀, Helio(2), 27.iii–4.v.1988, Griffin(1) ( PT); 1♂, 2♀, Okaukuejo, SE1915 Bb, vi.1991, Versfeld; 1♀, B de la Bat Restcamp, 9–13.iv.1993, Green( MT); 1♂, 1♀, Persianer705, 25.ii.1994, Marais; 1♂, 1♀, Popa Falls(1), 26.ii–1.iii.1992, Marais& Pusch(LT); 1♀, Nakatwa, 8–13.iii.1992, Pusch& Marais( MT); 1♂, Mushare, Sof, 27.iii–28.iv.1988, Griffin(1) ( PT); 1♂, Wolfsnes, Wof, 24.iii–10.v.1988, Griffin(1) ( PT); 2♂, 4♀, Waterberg Plateau Park, Onjoka, iv.1992, Green( MT); 1♀, Grootberg191: Khoadi// HôasM24, 24–27.v.2001, Marais& KirkSpriggs(1) ( PT); 1♀, Shaka: Salamabala[ sic– Salambala] M8, 27.ii–2.iii.2001, KirkSpriggs(1) ( MT); 1♂, 1♀, mopane in SalambalaM4, 24–26.ii.2001, Marais& KirkSpriggs(1) ( YP); 1♀, Simataa: Salambala B6, 24–27.ii.2001, Marais& Kirk Spriggs(1) ( YP); 4♀, Shaka: Salambala M8, 27.ii–2.iii.2001, KirkSpriggs(1) ( MT); 11♂, 10♀, Salambala, 4 kmNE of, 25.ii–1.iii.2001, KirkSpriggs(1) & Marais( MT); 1♀, Salambala forest, 23–29.xii.2002, KirkSpriggs(2) ( MT); 2♂, 3♀, Salambalacampsite, 28–29.xii.2002, Kirk Spriggs(2) ( HT)fish; 1♂, same except: 3–4.iii.2001, KirkSpriggs(1) ( HT)fish; 1♂, 3♀, same except: 23–28.xii.2002, KirkSpriggs(2) ( HT)mango; 18♂, 24♀, same except: 26–28.ii.2001, Marais& KirkSpriggs(1) ( HT)fruit; 4♂, 7♀, same except: 22–24.ii.2001, KirkSpriggs(1) & Marais( MT); 6♀, Salambalapan, 1–4.iii.2001, KirkSpriggs(1) ( MT); 1♀, same except: 23–27.xii.2002; 7♀, Simanya: Okavango River, 23–24.i.1998, KirkSpriggs(1) & Marais( MT)primary woodland; 3♀, Dussi, 10 kmW of, 29–30.xii.1998, KirkSpriggs(1), Marais& Mann( MT); 2♂, 5♀, Divuju: Okavango River, 1.i.1999, KirkSpriggs(1), Marais& Mann( MT) & (S); 1♂, Xawashapan, 2 kmW, 26–27.xii.1998, KirkSpriggs(1), Marais& Mann( MT); 1♂, Xawashapan, 26.xii.1998, Marais& KirkSpriggs(1) (S); 3♀, Namib Desert Res, Stn., Kuiseb Riv., 14–26.ii.1997, Kapofi& Irwin( MT)riparian vegetation; 1♀, Varianto771/2, 30.iii–1.iv.2003, KirkSpriggs(1) ( MT); 1♀, same except: KirkSpriggs(1) & Mey (LT); 3♂, 2♀, same except: ( HT)fungi; 5♂, 9♀, same except: 29–30.iii.2003, KirkSpriggs(1) ( HT)fruit; 2♂, Mile46, 25–27.iii.2003, KirkSpriggs(1) ( MT); 1♂, 1♀, same except: ( HT)fruit; 1♂, 1♀, Rundu( Kavango Lodge), 28.iii.2003, KirkSpriggs(1) ( HT)fruit; 4♂, 2♀, same except: 27–29.iii.2003, KirkSpriggs(1) & Mey (LT); 4♀, Somerkoms521, 6–8.ii.2001, Kirk Spriggs(1), Marais& Wheeler( MT); 3♀, Luchando village, 26–27.ii.2001, KirkSpriggs(1) & Marais( MT); 1♂, 4♀, Xawashapan, 27.xii.1998, KirkSpriggs(1) ( MT); 1♂, 3♀, Mannheim Research Station, 31.i–7.ii.2000, KirkSpriggs(1) ( MT)mango plantation; 1♀, Salambalacampsite, 1–4.iii.2001, KirkSpriggs(1) ( PT)baited millipede; 3♀, Hippo Lodge( Zambezi River), 6–7.ii.2004, KirkSpriggs(1) ( HT)fruit; 2♂, 5♀, Toggenburg591 campsite, 18–20.ii.2004, KirkSpriggs(1) ( HT)fruit; 1♀, same except: ( MT); 6♂, 18♀, Popa Fallsrestcamp, 13–16.ii.2004, KirkSpriggs(1) ( MT), riverine forest; 3♀, Bum Hillcampsite ( Kwando River), 10–13.ii.2004, KirkSpriggs(1) ( MT); 14♂, 17♀, same except: ( HT)fruit; 1♂, 4♀, Salambalacampsite, 8–10.ii.2004, KirkSpriggs(1) ( MT); 3♂, 6♀, same except: ( HT)fruit; 6♂, 5♀, Omatjenne Agricultural Research Station, 7.ii.2004, Shihepo, fruiting Termitomyces schimperi[all with associated puparia, 3♂, larva coll: 7.ii.2004, pupated: 15.ii.2004, emerged: 22.ii.2004; 3♂, 2♀, larva coll: 7.ii.2004, pupated: 14.ii.2004, emerged: 21.ii.2004; 3♀, larva coll: 7.ii.2004, pupated: 15.ii.2004, emerged: 21.ii.2004]; 2♂, 7♀, same except: fruiting Termitomyces schimperi[all with associated puparia, larva coll: 5.ii.2004, pupated: 14.ii.2004, emerged: 20.ii.2004]; 1♂, Veterinary ServicesHQ ( KM1)(2), 5–12.iii.2004, Mudenga( NUL); 1♂, Hungorobravine at(1): 25.iv.2000, Meakin/ Raleigh Int.(LT) Bberg UV 31; 1♀, Upper Hungorobravine at: 7.iv.2000, Meakin/ Raleigh Int( MT) Bberg Mal36; 1♀, Fallsrock ravine at: 17.iv.2000, Meakin/ Raleigh Int.( MT) Bberg Mal37. NOTES: The commonest species of the genus in Africa (Zumpt 1953c: 486) bred from mushrooms in South Africa. Cuthbertson (1935: 17, as H. clara) notes that flies are attracted to the honeydew of scale insects ( Hemiptera: Coccidae) on fruit trees and were occasionally observed on fresh cattle dung. He briefly discusses the breeding habits and notes a large number of flies as being attracted to a mushroom growing out of a large termite hill in Zimbabwe. Later (Cuthbertson 1939: 145, as Tricyclea clara) also notes the occurrence of larvae in wild mushrooms in December and January in Zimbabwe. In Namibiacollected in pitfall, yellow pan and Malaise traps; by sweeping, and at UVlight. Further collected in hanging traps baited with fermenting mango, fruit bait, rotting fish and rotting fungi, and in a pitfall trap baited with a dead millipede. Most significantly, the species has been reared in numbers from the fruiting bodies of the fungus Termitomyces schimperi(Pat.), which grows from termite hills in the rainy season. Occurring at low and high elevations on the Brandberg ( 700 m, 1170 m, 1920 m). Widely distributed in central, north and northeastern Namibia; in the ‘arid’ and ‘mesic’ savanna, namakaroo and desert biomes ( Fig. 67). Recorded throughout the year, except the colder months of September, October and November; most abundantly in February ( vide Table 2). Cuthbertson (1935: 17, as H. clara) briefly describes the egg, larvae and puparium and illustrates: the larval anal compartment, lateral aspect (Plate IV, 15a); posterior spiracles, from behind ( ibid, 15b); cephaloskeleton, lateral aspect and detail of the mouthooks ( ibid, 15d and 15f); anterior spiracles ( ibid, 15e), and the pupal posterior spiracles, from behind ( ibid, 15c).
Hemigymnochaeta varia(Hough, 1898: 178) ( Parochromyia). Fig. 68. TYPE LOCALITY: Somalia. DISTRIBUTION: Widespread in East Africato southern Africa: Angola, Democratic Republicof Congo, Kenya, Namibia, Rwanda, South Africa( Cape, Natal, Transvaal), Somalia, Tanzania, Ugandaand Zimbabwe. PREVIOUS RECORD: Omatjette [ 21°05'S, 15°05'E] (Zumpt 1953c: 490). KNOWN RECORDS: Gobabis [ 22°45'S, 18°97'E], 11.iii.1971; Okakuejo [=Okaukuejo] [ 19°18'S, 15°93'E], 13.iii.1971; Omatjette [ 21°05'S, 15°05'E], 11.iv.1950(NMSA).
MATERIAL: 1♂, 1♀, Simanya: Okavango River, 23–24.i.1998, KirkSpriggs(1) & Marais( MT)primary woodland; 1♀, Ruacana area, 14.iii.1997, Koch( MT); 1♀, Nama, 20–22.xii.1998, Kirk Spriggs(1), Marais& Mann( MT); 3♀, Persianer705, 25.ii.1994, Marais; 3♀, Toggenburg591 campsite, 18–20.ii.2004, KirkSpriggs(1) ( HT)fruit. NOTES: Cuthbertson (1935: 17, as H. pallens) reared larvae from the broken down fungus beds of termites and adults from Aardvark burrows in termite nests. He also observed adults of both sexes on fresh cattle dung in shade. Engel and Cuthbertson (1937: 10, as Tricyclea angola) note larvae from mushrooms in Ugandaand provide details of the immature stages. Later (Cuthbertson 1939: 17, as Tricyclea tridentata) noted both sexes encountered on faeces, pig dung and fresh cattle dung, plus males in blossoms. In Namibiaonly collected in Malaise traps and in a hanging trap baited with fermenting fruit. Widely distributed over central and northern Namibia; apparently restricted to the ‘arid’ savanna biome ( Fig. 68). Recorded from December to March ( vide Table 2). Cuthbertson (1935: 17–18) provides a brief description of the larva and puparium and illustrates the following larval parts: anal compartment, lateral aspect (Plate IV, 16a); posterior spiracles, from behind ( ibid, 16b); anterior spiracle ( ibid, 16c); and cephaloskeleton, lateral aspect ( ibid, 16d).
[264,526,828,851]
LOCALITY
Liberia
Liberia
66
67
1
holotype
[390,1044,923,947]
2001-03-01
2001-03-03
2001-03-01
HT
Kirk-Spriggs
Salambala
66
67
4
4
2001-03-03
2001-03-04
2001-03-03
HT
Kirk-Spriggs
66
67
2
2
2001-02-26
2001-02-28
2001-02-26
HT
Marais & Spriggs
66
67
19
17
2
[554,1171,987,1011]
2004-02-08
2004-02-10
2004-02-08
HT
Kirk-Spriggs
66
67
6
4
2
MT
66
67
6
5
1
[542,1269,1019,1043]
2001-02-22
2001-02-24
2001-02-22
MT
Kirk-Spriggs & Marais
66
67
1
1
2001-03-01
2001-03-04
2001-03-01
PT
Kirk-Spriggs
66
67
1
1
2004-02-10
2004-02-13
2004-02-10
HT
Kirk-Spriggs
Kwando
Bum Hill
66
67
2
2
Rivers
[997,1267,1083,1107]
MT
66
67
1
1
Rivers
2004-02-18
2004-02-20
2004-02-18
HT
Kirk-Spriggs
Toggenburg
66
67
1
1
Rivers
2004-02-13
2004-02-16
2004-02-13
MT
Kirk-Spriggs
Popa Falls
66
67
6
2
4
Rivers
[956,1200,1147,1171]
HT
66
67
1
1
Rivers
[264,707,1436,1459]
LOCALITY
Zimbabwe
Zimbabwe
66
67
1
holotype
[264,540,1788,1811]
LOCALITY
Namibia
Namibia
66
67
1
holotype
[264,540,412,435]
LOCALITY
Namibia
Namibia
67
68
1
holotype
[264,585,636,659]
LOCALITY
Sierra Leone
Sierra Leone
67
68
1
holotype
RECORDS
-21.116667
1265
19.2
67
68
1
[264,972,860,883]
1950-04-30
KNOWN, RECORDS
-19.55
1271
18.083334
67
68
1
[390,1124,923,947]
1986-05-16
1986-06-22
1986-05-16
PT
Griffin
Ondundozonananandana Mts.
67
68
1
1
H65447
1986-05-15
1986-06-16
1986-05-15
PT
Griffin
Okaukuejo-Duikersdrink
67
68
2
1
1
H65568, H65569
1986-05-20
1986-06-18
1986-05-20
PT
Griffin
Etosha Game Park
67
68
3
1
2
H65403
1986-05-16
1986-06-12
1986-05-16
PT
Griffin
Wolfnes-Okondeka
67
68
3
3
1986-07-16
1986-07-26
1986-07-16
MT
Irish
Ghaub
67
68
6
4
2
H65186
[473,963,1051,1075]
1986-03
Tolken
Sturmfeld
67
68
1
1
[975,1319,1051,1075]
1986-05-15
Curtis
Tobiroen
67
68
1
1
[312,858,1083,1107]
1986-05-14
1986-06-16
1986-05-14
PT
Griffin
Dorsland
67
68
2
1
1
H58092
1984-01-20
Irish
Hereroland-Oos
67
68
1
1
H25130
[413,911,1115,1139]
Gaerdes
Okahandja
67
68
1
1
1987-08-07
1987-09-15
1987-08-07
PT
Griffin
Sprokieswoud
67
68
1
1
[486,1116,1147,1171]
1993-02-02
1993-03-01
1993-02-02
PT
Pusch
Park
Waterberg
67
68
1
1
Plateau
1997-03-14
MT
Koch
Ruacana area
67
68
13
11
2
Plateau
[659,1217,1179,1203]
1988-03-27
1988-05-04
1988-03-27
PT
Griffin
Helio
67
68
5
2
3
Plateau
1991-06
Okaukuejo, SE & Versfeld
Bb
67
68
3
2
1
Plateau
1993-04-09
1993-04-13
1993-04-09
MT
B de la Bat Restcamp & Green
67
68
1
1
Plateau
[390,867,1243,1267]
1994-02-25
Marais
Persianer
67
68
2
1
1
Plateau
1992-02-26
1992-03-01
1992-02-26
Marais & Pusch
Popa Falls
67
68
2
1
1
Plateau
[555,1107,1275,1299]
1992-03-08
1992-03-13
1992-03-08
MT
Pusch & Marais
Nakatwa
67
68
1
1
Plateau
1988-03-27
1988-04-28
1988-03-27
PT
Mushare, S & Griffin
67
68
1
1
Plateau
[685,1271,1307,1331]
1988-03-24
1988-05-10
1988-03-24
PT
Wolfsnes, W & Griffin
67
68
1
1
Plateau
1992-04
MT
Green
Waterberg
Onjoka
Park
67
68
6
4
2
Plateau
2001-05-24
2001-05-27
2001-05-24
PT
Marais & Kirk-Spriggs
Grootberg
Hoas
Khoadi
67
68
1
1
Plateau
2001-02-27
2001-03-02
2001-02-27
MT
Kirk-Spriggs
Salamabala
Shaka
67
68
1
1
Plateau
2001-02-24
2001-02-26
2001-02-24
YP
Marais & Kirk-Spriggs
Salambala
67
68
2
1
1
Plateau
2001-02-24
2001-02-27
2001-02-24
YP
Salambala B & Marais & Spriggs
Simataa
67
68
1
1
Plateau
[497,1204,1467,1491]
2001-02-27
2001-03-02
2001-02-27
MT
Salambala M & Kirk-Spriggs
Shaka
67
68
4
4
Plateau
2001-02-25
2001-03-01
2001-02-25
MT
Kirk-Spriggs & Marais
Salambala
67
68
21
10
11
Plateau
2002-12-23
2002-12-29
2002-12-23
MT
Kirk-Spriggs
Salambala forest
67
68
1
1
Plateau
2002-12-28
2002-12-29
2002-12-28
HT
Spriggs
Salambala
67
68
5
3
2
Plateau
[548,1115,1563,1587]
2001-03-03
2001-03-04
2001-03-03
HT
Kirk-Spriggs
67
68
1
1
Plateau
2002-12-23
2002-12-28
2002-12-23
HT
Kirk-Spriggs
67
68
4
3
1
Plateau
2001-02-26
2001-02-28
2001-02-26
HT
Marais & Kirk-Spriggs
67
68
42
24
18
Plateau
2001-02-22
2001-02-24
2001-02-22
MT
Kirk-Spriggs & Marais
67
68
11
7
4
Plateau
[478,1108,1659,1683]
2001-03-01
2001-03-04
2001-03-01
MT
Kirk-Spriggs
Salambala
67
68
6
6
Plateau
[487,1323,1691,1715]
1998-01-23
1998-01-24
1998-01-23
MT
Kirk-Spriggs & Marais
Simanya
67
68
7
7
Okavango River
1998-12-29
1998-12-30
1998-12-29
MT
Kirk-Spriggs & Marais & Mann
Dussi
67
68
3
3
Okavango River
1999-01-01
MT
Kirk-Spriggs & Marais & Mann
Divuju
67
68
7
5
2
Okavango River
[360,1272,1787,1811]
1998-12-26
1998-12-27
1998-12-26
MT
Kirk-Spriggs & Marais & Mann
Xawasha
67
68
1
1
Okavango River
1998-12-26
Marais & Kirk-Spriggs
Xawasha
67
68
1
1
Okavango River
1997-02-14
1997-02-26
1997-02-14
MT
Kapofi & Irwin
Namib Desert Res
Kuiseb Riv.
Stn.
67
68
3
3
Okavango River
2003-03-30
2003-04-01
2003-03-30
MT
Kirk-Spriggs
68
69
Varianto
67
68
1
1
Okavango River
HT
68
69
5
2
3
Okavango River
[631,1263,315,339]
2003-03-29
2003-03-30
2003-03-29
HT
Kirk-Spriggs
68
69
14
9
5
Okavango River
[312,866,347,371]
2003-03-25
2003-03-27
2003-03-25
MT
Kirk-Spriggs
Mile
68
69
2
2
Okavango River
[878,1171,347,371]
HT
68
69
2
1
1
Okavango River
2003-03-28
HT
Kirk-Spriggs
Lodge
Rundu
68
69
2
1
1
Kavango West
2001-02-06
2001-02-08
2001-02-06
MT
Spriggs & Marais & Wheeler
Somerkoms
68
69
4
4
Kavango West
2001-02-26
2001-02-27
2001-02-26
MT
Kirk-Spriggs & Marais
Luchando village
68
69
3
3
Kavango West
[469,1100,475,499]
1998-12-27
MT
Kirk-Spriggs
Xawasha
68
69
5
4
1
Kavango West
2000-01-31
2000-02-07
2000-01-31
MT
Kirk-Spriggs
Mannheim Research Station
68
69
4
3
1
Kavango West
2001-03-01
2001-03-04
2001-03-01
PT
Kirk-Spriggs
Salambala
68
69
1
1
Kavango West
2004-02-06
2004-02-07
2004-02-06
HT
Kirk-Spriggs
Hippo Lodge
68
69
3
3
Zambezi
2004-02-18
2004-02-20
2004-02-18
HT
Kirk-Spriggs
Toggenburg
68
69
7
5
2
Zambezi
[777,1034,603,627]
MT
68
69
1
1
Zambezi
2004-02-13
2004-02-16
2004-02-13
MT
Kirk-Spriggs
Popa Falls
68
69
24
18
6
Zambezi
2004-02-10
2004-02-13
2004-02-10
MT
Kirk-Spriggs
Kwando
Bum Hill
68
69
3
3
Rivers
[824,1158,667,691]
HT
68
69
31
17
14
Rivers
2004-02-08
2004-02-10
2004-02-08
MT
Kirk-Spriggs
Salambala
68
69
5
4
1
Rivers
[933,1219,699,723]
HT
68
69
9
6
3
Rivers
2004-02-07
2004-02-22
2004-02-07
Shihepo
Omatjenne Agricultural Research Station
68
69
22
10
12
Rivers
[548,960,891,915]
KM
Veterinary Services
68
69
KM1
1
1
Rivers
[971,1319,892,915]
2004-03-05
2004-03-12
2004-03-05
NU-L
Mudenga
68
69
1
Rivers
[312,1192,923,947]
2000-04-25
UV
Meakin & Raleigh Int. & Bberg
Hungorob
68
69
UV 31
1
1
Rivers
2000-04-07
MT
Meakin & Raleigh Int
Upper Hungorob
68
69
1
1
Rivers
[931,1083,956,979]
Bberg Mal
68
69
1
Rivers
2000-04-17
MT
Meakin & Raleigh Int.
Falls
68
69
1
1
Rivers
[757,908,988,1011]
Bberg Mal
68
69
1
Rivers
LOCALITY
Somalia
Democratic Republic
Widespread in East Africa
southern Africa
68
69
1
RECORD
Uganda
-21.083334
Uganda
1265
15.083333
68
69
1
[389,1275,347,371]
1998-01-23
1998-01-24
1998-01-23
MT
Kirk-Spriggs & Marais
Simanya
69
70
2
1
1
Okavango River
[494,954,379,403]
1997-03-14
MT
Koch
Ruacana area
69
70
1
1
Okavango River
1998-12-20
1998-12-22
1998-12-20
MT
Spriggs & Marais & Mann
Nama
69
70
1
1
Okavango River
[688,1092,411,435]
1994-02-25
Marais
Persianer
69
70
3
3
Okavango River
2004-02-18
2004-02-20
2004-02-18
HT
Kirk-Spriggs
Toggenburg
69
70
3
3
Okavango River