Taxonomic assessment of sharks, rays and guitarfishes (Chondrichthyes: Elasmobranchii) from south-eastern Arabia, using the NADH dehydrogenase subunit 2 (NADH 2) gene Henderson, Aaron C. Reeve, Alan J. Jabado, Rima W. Naylor, Gavin J. P. Zoological Journal of the Linnean Society 2016 2015-10-05 176 2 399 442 D2V 1158979 (EAGLE RAYS) [308,634,1505,1526] Elasmobranchii Myliobatidae Animalia Myliobatiformes 31 430 Chordata family  Two species of  Aetomylaeuswere also included in the analysis, namely  A. nichofiiand  A. milvus, and they exhibited a between-group mean distance of 18.43 ± 1.33%. Naylor et al. (2012)noted a division within their  A. nichofiispecimens, with specimens from the Persian Gulf and Australiaclustering beside the bulk of their specimens, which came from Borneo. The clusters were therefore designated  A. nichofii(Borneo),  A. cf. nichofii1 (Persian Gulf) and A. cf. 1,  Acroteriobatus salalah; 2,  Acroteriobatussp.; 3,  Glaucostegus halavi; 4,  Rhinobatossp.(cluster A); 5,  Rhinobatossp.(cluster B); 6,  Rhinobatos punctifer.   Table 4.Between-group mean p-distances (%) for rhinobatid taxa from south-eastern Arabia (below the diagonal) with their standard error values (%) (above the diagonal)    1 2 3 4 5 6  1 1.02 1.35 0.96 1.03 1.13  2 9.89 1.46 1.04 1.06 1.09  3 18.91 20.55 1.34 1.38 1.43  4 9.94 10.49 18.59 0.42 1.06  5 9.94 10.94 19.09 2.12 1.08  6 11.89 11.99 21.50 11.39 11.53 this contained  A. salalahand the other undescribed species, which was designated  Acroteriobatussp.( Fig. 8). These congeners also exhibit a high between-group mean distance. The final rhinobatid species was  Glaucostegus halavi. None of these species was reported by Naylor et al. (2012); however, when viewed alongside closely related taxa all the clusters remained distinct ( Fig. 9).   Figure 8.  Rhinobatossp.(top) is morphologically and genetically distinct from  Rhinobatos punctifer(middle). The bottom image shows  Acroteriobatussp.   nichofii2 ( Australia). All the current specimens clustered with  A. cf. nichofii1. The current  A. milvusspecimens clustered with two Persian Gulf specimens reported by Naylor et al. (2012).