Revision of Diplosynapsis Enderlein, 1914 (Diptera, Asilidae, Asilinae) with description of four new species Vieira, Rodrigo Rafael, José Albertino Zootaxa 2016 4208 2 127 148 9HRK7 [151,404,1609,1635] Insecta Asilidae Diplosynapsis GBIF Animalia Diptera 10 137 Arthropoda species fisheri sp. nov.   Diagnosis.Sternite 8 with medioapical margin produced ( Fig. 45); epandrium with ventral digitiform projection ( Figs. 38, 40); gonocoxite apex bifid ( Figs. 40, 41); female: hind femur with 3 black setae anteriorly; 4–5 black anteroventrally.   Description. Male. Holotype.Head. Scape and pedicel black, postpedicel brown; 2 long, black ocellar macrosetae and 5 setae; vertex black; frons and face black, golden and grey tomentose on lateral margins; mystax with black setae; palpus with black setae; proboscis black with black and yellow setae; labial setae yellowish; occiput grey tomentose; occipital with black and yellow setae; 3–4 black postocular macrosetae. Thorax. Antepronotum and postpronotum black, sparsely grey tomentose; mesonotum black; lateral margin of mesonotum, postpronotal, sutural and prescutellar spot and scutellum grey tomentose; mesopleuron black, grey tomentose. Chaetotaxy: 2 black notopleural macrosetae; 1 black supra-alar setae; 1 black postalar setae; black dorsocentral setae; no anatergal setae; 7 black, apical scutellar macrosetae; black discal scutellar setae; black katatergal macrosetae; setae on meron+metanepisternal yellow and blackish. Wing ( Fig. 39). Light brown, with costal dilation; crossvein r-m beyond middle of discal cell; microtrichia on posterior wing margin arranged in a single plane; halter yellow-brown. Legs ( Fig. 37). Black. Chaetotaxy: fore femur with black and yellow setae; mid femur with 2 black macrosetae anteriorly and 3 anteroventrally; hind femur with 1 black macrosetae anteriorly, 4 anteroventrally, 1 anterior preapical dorsal macroseta, and another posteriorly; fore tibia apical 3/4, hind tibia 4/5, all tarsomeres ventrally, and fore and hind tarsomeres with brown, short, spiniform setae anteriorly; fore tibia with 4 black macrosetae posteriorly; mid tibia with 3 black setae anteroventrally; hind tibia with 1 black apical and 1 black posterior macroseta; tarsomeres with black setae. Abdomen. Tergites 2–5 black, except apex brown and basal grey spot laterally; tergites and sternites 6–7 brown with dull grey tomentum ( Figs. 37, 38); sternite 8 with medioapical margin produced ( Fig. 45); sternites black with brown apex, grey tomentose. Terminalia ( Figs. 38, 40–43). Black; terminalia oblique to body axis ( Fig. 37). Epandrium with ventral digitiform projection ( Figs. 38, 40); hypandrium developed laterally, with rounded ventral margin; gonocoxite apex bifid ( Figs. 40, 41); gonostylus apex rounded with short spines ( Figs. 41, 42); ejaculatory apodeme long and wide proximally, in lateral view ( Fig. 43); aedeagus apex without ventral projection ( Figs. 43, 44).  Lengths.Body 23.8 mm; wing 15.6 mm.   Holotype condition.Wing mounted in microslide with Canada balsam and terminalia in microtube with glycerin. Microtube and microslides pinned with specimen.   Variation.Length: Body 18.2 mm– 28 mm; wing: 11.2 mm– 16 mm; antenna wholly black; 3 long, black ocellar macrosetae; halter brown; mid femur with 3 black macrosetae anteriorly; hind femur with 5 black macrosetae anteroventrally.  Female.( Figs. 46–48): Similar to male, except for body length: 16 mm–27.8 mm, wing 14.5 mm–17.1 mm; 5 long black ocellar macrosetae; mid femur with 3 black setae anteriorly; hind femur with 3 black setae anteriorly; 4– 5 black anteroventrally; tergites I–VI lateral margin, grey tomentose; tergite 8 long, slender and black ( Figs. 46, 48); tergite 9 + 10 membranous medially; sternites 1–7 black, grey tomentose; 3 spermathecae capsule small, rounded and sclerotized ( Fig. 47).   Etymology.A patronym to the researcher Eric Fisher, PhD.   Distribution. Costa Rica(Guapiles, Heredia, Limon, Puntarenas) ( Fig. 75).     Typematerial examined. Holotype:C.[ OSTA] RICA:  Heredia,  Finca La Selva,  4.Apr.[il]. [19]80 (1146). H. Hespenheide/ Holotype  Diplosynapsis fisheri( ♂EFISHER).    Paratypes: COSTA RICA: Prov.[ince]  Heredia: 11 kmSE. La. Virgen,  450-550m, 10°20’N 84°04’W  8 Abril 2003, INBio-OET-ALAS transect / 0  8 Abril 2003 Transect05/M/01/061 / Project ALAS,  INB0003239606/ Paratype  Diplosynapsis fisheri( ♀EFISHER);  COSTA RICA,  Heredia:Est.[ácion]. Biol.[ógica]  La Selva,  50- 150m, 10°26’N, 84°01’ W,  May 2000, INBio-OET /  29 May/ 2000 Bosquesecundário / M/19/749 / Costa Rica INBIO  CRI0022730022 / Paratype  Diplosynapsis fisheri( ♀EFISHER);  COSTA RICA:  Heredia, Pr[ovince]:  La Selva Biol[ogical] Sta.[tion],  3 kmS Pto.[Puerto] Viejo, 10°26’N, 84°01’ W/  23.vii.1993, H.A. Hespenheide/ Paratype  Diplosynapsis fisheri( ♂EFISHER); same data, except:  20.vii.1982/ Paratype  Diplosynapsis fisheri( ♂EFISHER); same data, except:  15.iv.1982/ Paratype  Diplosynapsis fisheri( ♂EFISHER); same data, no date /  Efferiasp. A, det. E.M. Fisher/ Paratype  Diplosynapsis fisheri( ♂EFISHER);  COSTA RICA: Prov.[ince]  Heredia: 11kmSE. La. Virgen,  250-350m, 10°21’N 84°03’W,  18 Abril 2004, 03/ELP/01 Transect/  18 Abril 2004 Transect05/M/01/061 / Project ALAS,  INB0003622914/ Paratype  Diplosynapsis fisheri( ♂EFISHER);  COSTA RICA, LimonProv.[ince]  Guapiles,  vi.24-[19]65, Coll. R.J.Hamton / Paratype  Diplosynapsis fisheri( ♀EFISHER);  COSTA RICA, Cartago, 20 km, ne, Turrialba,   820m.  July 23, 1975, E.M. Fisher, collr. / Paratype  Diplosynapsis fisheri( ♀EFISHER); same data / =  DiplosynapsisEnderlein Enderl.[ein] det. E.M. Fisher[19]92 / Paratype  Diplosynapsis fisheri( ♂EFISHER);  COSTA RICA,  PuntarenasProv.[ince], Parque Nacional Corcovado, Est.[ácion] Sirena, 8° 28’ 31” N, 83°36’ W/  25.iii.1981 H.A. Hespenheide/ Paratype  Diplosynapsis fisheri( ♀EFISHER);  COSTA RICA:  LimonProv.[ince] 17 km, W. Guapiles, 13, Jul.[y], 1986, F.T. Hovore, P.H. Sullivan/ Paratype  Diplosynapsis fisheri( ♀EFISHER). 1416638196 4. Apr. OSTA Finca La Selva & H. Hespenheide Finca La Selva 11 138 1 1 Heredia holotype 1416638209 2003-04-08 ALAS, INB Transect & Project Costa Rica 500 10.333333 Heredia 1298 -84.066666 11 138 INB0003239606 1 1 Heredia paratype 1416638198 2000-05 2000-05-29 2000-05 INBIO, CRI La Selva & Bosque & Costa Rica Costa Rica 100 10.433333 La Selva 1298 -84.01667 11 138 CRI0022 1 1 Heredia paratype 1416638204 1993-07-23 La Selva Biol & H. A. Hespenheide Costa Rica Pr 10.433333 Viejo 1298 -84.01667 La Selva Biol 11 138 1 1 Heredia paratype 1416638210 1982-07-20 La Selva Biol & H. A. Hespenheide Costa Rica Pr 10.433333 Viejo 1298 -84.01667 La Selva Biol 11 138 1 1 Heredia paratype 1416638212 [292,1187,1591,1617] 1982-04-15 La Selva Biol & H. A. Hespenheide Costa Rica Pr 10.433333 Viejo 1298 -84.01667 La Selva Biol 11 138 1 1 Heredia paratype 1416638195 1982-04-15 La Selva Biol & H. A. Hespenheide Costa Rica Pr E. M. Fisher 10.433333 Viejo 1298 -84.01667 La Selva Biol 11 138 1 1 Heredia paratype 1416638208 2004-04-18 ALAS, INB Transect & Project Costa Rica 300 10.35 Heredia 1298 -84.05 11 138 INB0003622914 1 1 Heredia paratype 1416638200 1924-06 Coll. R. J. Costa Rica Guapiles 11 138 1 1 Limon paratype 1416638206 1975-07-23 E. M. Fisher Costa Rica 820 Turrialba 11 138 821 1 Cartago paratype 1416638218 1975-07-23 E. M. Fisher Costa Rica E. M. Fisher 820 Turrialba 11 138 1 1 Cartago paratype 1416638215 1981-03-25 H. A. Hespenheide Costa Rica 8.475277 Sirena 915 -83.6 Parque Nacional Corcovado 11 138 1 1 Puntarenas paratype 1416638202 W. Guapiles & Jul. & F. T. Hovore & P. H. Sullivan Costa Rica Limon 11 138 1 1 Limon paratype