Matthews, 1974: 109 Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups Gunter, Nicole L. Weir, Thomas A. Zootaxa 2019 2019-11-08 4695 3 201 252 6P7WD Matthews, 1974 Matthews 1974 [151,572,1801,1828] Insecta Scarabaeidae Lepanus Animalia Coleoptera 3 204 Arthropoda species australis  ( Figs. 1C–D, F, K; 2A; 3A; 4A)     Lepanus australis  Matthews, 1974: 109.   Type material examined.   Holotype: ♂“ 5 mi. N. Batemans BayN.S.W.  16-VIII-67 E. Matthews” / “ Lepanus australis Det 1972M., E.G. Matthews HOLOTYPE” / “ ANICDatabase No. 25 057863” / “ HOLOTYPE”, in  ANIC[ 35°40’0”S, 150°15”0”E].  Paratypes: Macquarie Pass, [ 34°34’0”S, 150°38’0”E],  29.xii.1967, E. Matthews(25-057864 1 specimen ♂, 4 specimenssex undetermined in ANIC);  Clyde Mountain(general), [ 35°33’0”S, 149°57’0”E],  9.i.1968–  12.i.1968, E. Matthews(25-057865 1 specimensex undetermined in ANIC);  3–7 milesN of Eden, [ 37°0’0”S, 149°55’0”E],  13.i.1968, E. Matthews(25-057866 2 specimen ♂, 2 specimensex undetermined in ANIC);  5–17 milesS of Eden, [ 37°14’0”S, 149°55’0”E],  13.i.1968, E. Matthews(25-057867 1 specimensex undetermined in ANIC);  Mallacoota National Park, [ 37°30’0”S, 149°46’0”E],  13.i.1968, E. Matthews(25-057868 1 specimensex undetermined in ANIC).  Other material examined.Almost 600 other specimens were examined from around 150 localities.    Queensland: St Lawrence, 18.5 kmW, 22°21’4”S, 149°20’33”E,  29.vi–22.x.2000, (T107480–T 107482 in QM);  Fairlie’s Knob, 0.5 kmS, 25°30’32”S, 152°17’22”E,  20.x–20.xii.2000, (T 107490 in QM);  Fairlie’s Knob, 0.5 kmS,  20.xii.2000–  23.iii.2001, (T107491, T107493–T 107495 in QM);  Fairlie’s Knob, 0.5 kmS,  22–23.iii.2001, (T 107500 in QM);  Lake Bowaddy[Lake Bowarrady], Fraser Island, [ 25°9’0”S, 153°13’0”E],  3.xii.1975, G. Thompson& J.A. Slater(25-058488 1 specimenin QM);  Yidney Scrub, Fraser Island, [ 25°19’0”S, 153°11’0”E],  1.xii– 4.xii.1975, G. Thompson& J.A. Slater(25-058491 2 specimensin QM);  Booloumba Creek State Forest RiverTrail, 26°38’50”S, 152°38’45”E,  350m,  15.i.2000, R.A. Anderson(CMNEN-00030001 in CMNC);  Tinana Creek, 25°45’7”S, 152°41’38”E,  35 m,  22.xi–7.xii.2011, G.B. Monteith(25-0053884 1 specimenalso as COL 2345 in ANIC);  Tallegalla Weir Junction, 25°45’25”S, 152°41’49”E,  35m,  6–7.xii.2011, G.B. Monteith(25-053887 1 specimenin ANIC);  Nangur State Forest, 2nd site, 26°7’55”S, 151°58’43”E,  24.xi.1995–  3.ii.1996, (T 107570 in QM);  Jack Smith[Scrub Conservation] Park on Boat Mountain, 26°9’17”S, 151°54’34”E,  26.i–20.iv.1995, (T107462 – T107464, T107533–T 107552 in QM);  Jack Smith[Scrub Conservation] Park on Boat Mountain, 26°9’17”S, 151°54’34”E,  15.xii.1994–  26.i.1995, (T 107553 in QM);  Kin Kin Creek, lower, 26°14’15”S, 152°58’52”E,  18– 19.ii.2006, (T 135125 in QM);  Elanda Point, National Park Office, 26°15’28”S, 152°59’44”E,  17.ii.2006, (T135122– T 135124 in QM);  Cooran Plateauvia Gympie, [ 26°21’0”S, 152°47’0”E],  1300 m,  10–11.iii.1973, R.I. Storey(25- 058486  x 12in QM, 25-058581, 2 specimensin QDPC);  Cooran Plateau,  1300 feetvia Gympie, [ 26°21’0”S, 152°47’0”E],  396 m,  3.x–3.xi.1973, R.I. Storey(CMNEN-00030019 in CMNC);  Kingaroy, 6 kmSE, 26°34’48”S, 151°52’35”E,  23–26.xii.2005,(T132946–T 132949in QM);  Mapleton Falls National Park, 26°37’49”S, 152°50’21”E,  30.xi.1991-  8.i.1992, (T 107573 in QM);   4 kmW Buderim, [ 26°41’0”S, 153°0’0”E],  6.ix.1978, I.F.B. Common(25- 059200  x 1in ANIC);  Chevallum, via Nambour, 26°42’3”S, 153°0’33”E,  12.v–27.ix.1978, (T 107594 in QM);  Conondale Range, Site1, 26°42’46”S, 152°37’8”E,  20.iv-2.v.1995, (T107567–T 107568 in QM);  Mary Cairncross Park, 26°46’52”S, 152°52’56”E,  29.xi.1991-  8.i.1992, (T 107574 in QM);  Landsborough, 2.55 kmNE, 26°47’22”S, 152°58’41”E,  28.viii.2010, G.B. Monteith(25-  060783 1 specimenalso as COL 773 in ANIC);  Saddletree Creek, 26°51’50”S, 151°38’2”E,  29–30.xii.2010, G.B. Monteith& K. Will(25-  060832 x2also as COL1464-COL 1465 in ANIC);  Saddletree Creek, 26°51’50”S, 151°38’2”E,  22–29.xii.2010, Sota& Takami, (25-  060834 x10also as COL1467- COL 1468 in ANIC);  Saddletree Creek, 26°51’50”S, 151°38’2”E,  22–30.xii.2010, G.B. Monteith& K. Will(25-  060835 7 specimensalso as COL  1469–1470in ANIC);   5 kmE Yarraman, [ 26°52’0”S, 152°3’0”E],  19.vi.1982, L. Hill, (25-059203 1 specimenin ANIC);  Top of Blackbutt Range, 26°52’38”S, 152°11’32”E,  24.xi.1995-  3.ii.1996, (T107517–T 107532 in QM);  Bunya Mountains, Dandabah, 26°52’47”S, 151°35’44”E,  20.ix.2007, (T   151105in QM);  3 kmN Upper Yarraman State Forest, 26°52’47”S, 151°53’4”E,  30.iii.1983, (T107571–T 107572 in QM);  Bunya Mountain National Park, Westcott Campground, 26°51’51”S, 151°34’17”E,  1100 m,  18.i.2000, R.A. Anderson(CMNEN-00030002, CMNEN-00030004 in CMNC);  Bunya Mountain National Park, Dandabah Picnic Area, 26°52’54”S, 151°35’23”E,  1100 m,  19.i.2000, R.A. Anderson(CMNEN-00030003 in CMNC);  BlackbuttRange,  1 kmE Benarkin, [ 26°53’0”S, 152°10’0”E],  19.vi.1982, L. Hill, (25-059202 1 specimenin ANIC);  Upper Yarraman State Forestvia Maidenwell, [ 26°54’0”S, 151°54’0”E],  11.i.1975, G.B. Monteith(25- 058493 3 specimensin QM);  Wild Horse Mountain, 1.2 kmSW, 26°55’26”S, 152°59’24”E,  15–16.xi.2007, (T   151108in QM);  Wild Horse Mountain, 1.2 kmSW, 26°55’26”S, 152°59’24”E,  23.i–13.ii.2008, (T154607– T 154614 in QM);  Bribie Island, 1.3 kmNE of national park headquarters, 27°0’54”S, 153°7’30”E,  10 m,  8–9.x.2010, G.B. Monteith(25-  060876 x1also as COL 1141in ANIC);  Bribie Island, 0.85 kmNE of national park headquarters, 27°1’5”S, 153°7’23”E,  8–9.x.2010, G.B. Monteith(25-  060877 1 specimenalso as COL 1142 in ANIC);  Brisbane, Forest Park, Scrub CreekRoad, [ 27°1’0”S, 152°26’0”E],  18.ix–29.x.1998, P. Bouchard(25-058494 1 specimenin QM);  Brisbane ForestPark, 27°25’41”S, 152°50’18”E,  1–15.x.2002, J. Skevington& J. Cumming(CMNEN- 00030008 in CMNC);  Blackbutt Range( Bottom),  8 milesE Blackbutt, [ 27°3’0”S, 151°55’0”E],  22–23.xii.1973, R.I. Storey(25-058580 2 specimensin QDPC);  Mount Deongwar, 3 kmS, Site3, 27°13’41”S, 152°15’36”E,  30.xii.1998–  26.iii.1999, (T107285–T 107292 in QM);  Mount Deongwar, 3 kmS, Site3, 27°13’41”S, 152°15’36”E,  14.x–30.xii.1998, (T 107391 in QM);  Mount Deongwar, 3 kmS, Site2, 27°13’49”S, 152°15’25”E,  30.xii.1998–  26.iii.1999, (T107282–T 107284 in QM);  Mount Deongwar, 3 kmS, Site2, 27°13’49”S, 152°15’25”E,  14.x– 31.xii.1998, (T 107390 in QM);  Mount Glorious, barracks, 27°19’54”S, 152°45’12”E,  16–28.iv.2007, (T 148366 in QM);  Mount Glorious, barracks, 27°19’54”S, 152°45’12”E,  12–15.iii.2008, (T 156700 in QM);  Crows Nest National Park, Perseverance Section, Site1, 27°19’8”S, 152°6’’27”E,  4–5.xii.2003, (T  120685–T   120687in QM);  Mount Glorious, [ 27°20’0”S, 152°46’0”E],  630 m,  14.xi.1986–  30.i.1987, T. Hiller, (25-059197 3 specimens, 25- 059198, 2 specimensin ANIC);  Mount Glorious, [ 27°20’0”S, 152°46’0”E],  635 m,  ii.1989, H.F. & A.T. Howden(CMNEN-00030007 in CMNC);  Stony Creek, via Samford, 27°20’20”S, 152°47’52”E,  22.x.1994-  2.ii.1995, (T107554–T 107558 in QM);  Stony Creek, via Samford, 27°20’20”S, 152°47’52”E, 2.ii–iv.1995, (T107559– T 107566 in QM);  Mount Glorious, 2 kmS, 27°21’9”S, 152°46’7”E,  28–29.i.2006, (T133758–T 133761 in QM);  Boombana National Park, Site1, 27°24’8”S, 152°47’22”E,  1.x.2003–  30.x.2003, (T   122130in QM);  Boombana National Park, Site1, 27°24’8”S, 152°47’22”E,  15–16.ii.2004, (T   122132in QM);  Boombana National Park, Site1, 27°24’8”S, 152°47’22”E,  2.i.2004–  30.i.2004, (T  122794–T   122795in QM);  Boombana National Park, 27°25’55”S, 152°53’49”E,  230 m,  18.ii.2012–  19.ii.2012, G.B. Monteith(25-053911  x 1in ANIC);  McAfee’s Lookout, 0.5 kmSW, 27°25’58”S, 152°52’14”E,  18.x.1999-  6.ii.2000, (T 107475 in QM);  McAfee’s Lookout, 1 kmSW, 27°26’15”S, 152°52’14”E,  18.x.1999–  6.ii.2000, (T 107476 in QM);  Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E,  21.xii.1999–  27.i.2000, (T107471–T 107473 in QM);  Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E,  27.i– 15.iii.2000, (T 107474 in QM);  Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E,  15.iii–18.v.2000, (T107478– T 107479 in QM);  Enoggera Reservoir, Site5, 27°26’44”S, 152°55’19”E,  16.x–21.xii.1999, (T107467–T 107469 in QM);  Enoggera Reservoir, site 5, 27°26’44”S, 152°55’19”E,  21.xii.1999–  27.i.2000, (T 107470 in QM);  Enoggera Reservoir, site 3, 27°26’44”S, 152°55’19”E,  15.iii–18.v.2000, (T 107477 in QM);  McAfee’s Lookout, 0.7 kmSW, 27°26’8”S, 152°52’17”E,  6.vii–16.x.1999, (T107393–T 107394 in QM);  Gold Creek Reservoir, Site4, 27°27’43”S, 152°52’21”E,  12–14.x.2007, (T   151106in QM);  Gold Creek Reservoir, 27°27’46”S, 152°52’54”E,  12–14.x.2007, (T   151107in QM);  Slaughter Falls, 27°28’44”S, 152°57’34”E,  6–7.ii.2001, (T 107483 in QM);  Boscombe Road, Brookfield, 27°28’57”S, 152°55’1”E,  12–13.ii.2011, G.B. Monteith(25-053877 1 specimenalso as COL 1678 in ANIC);  Gold Creek Reservoir, Site2, 27°28’5’’S, 152°52’20”E,  4.xi.2003, (T   122131in QM);  Upper Brookfield Road, Brisbane, [ 27°29’0”S, 152°52’0”E],  24.viii–7.ix.1975, R.I. Storey(25-058489 1 specimenin QM);  Upper Brookfield Road, Brisbane, [ 27°29’0”S, 152°52’0”E],  22.ix–4.x.1975, R.I. Storey(25-058490 5 specimensin QM);  Kenmore Kookaburra Street, 27°29’28”S, 152°55’23”E,  16–17.xi.2010, <collector unknown>, (25-  060906 1 specimenalso as COL 1197 in ANIC);  Kenmore Hills, 27°29’28”S, 152°55’24”E,  20–21.i.2011, G.B. Monteith(25- 065490 2 specimensin ANIC);  Kenmore, 27°30’24”S, 152°55’52”E,  20–21.i.2011, G.B. Monteith(25-065489 2 specimensin ANIC);  Mabb Street, Kenmore, 27°30’25”S, 152°55’52”E,  16–17.xi.2010, G.B. Monteith(25-  060889 1 specimenalso as COL 1196 in ANIC);  The Knobby,  4 kmW Glamorgan Vale, 27°30’27”S, 152°35’18”E,  13.i– 16.v.1999, (T 107279 inQM–T 107281 in QM);  Cameron’s Scrub, top ridge, 27°30’29”S, 152°43’54”E,  13.i– 16.v.1999, (T107303–T 107310 in QM);  Cameron’s Scrub, 27°30’32”S, 152°43’48”E,  75 m,  30.iv–1.v.2012, G.B. Monteith(25-057574 2 specimens, 25-075575 5 specimensin ANIC);  Long Pocket, river bank, 27°30’34”S, 152°59’53”E,  1.xi–4.xi.1999, (T 107395 in QM);  Cameron’s Scrub, gully, 27°30’35”S, 152°43’54”E,  13.i–16.v.1999, (T107311–T 107347 in QM);  Cameron’s Scrub, gully, 27°30’35”S, 152°43’54”E,  11.xi.1998–  13.i.1999, (T107349– T 107369 in QM);  Cameron’s Scrub, below road, 27°30’45”S, 152°43’39”E,  13.i–16.v.1999, (T107293–T 107295 in QM);  Belmont Hill, top, 27°30’48”S, 153°6’’59”E,  12.xii.1997–  7.v.1998, (T107417–T 107433 in QM); Cameron’s Scrub, Snigtrack, 27°30’49”S, 152°43’43”E,  13.i–16.v.1999, (T107296–T 107302 in QM); Cameron’s Scrub, Snigtrack, 27°30’49”S, 152°43’43”E,  11.xi.1998–  13.i.1999, (T 107389 in QM);  Whites Hill Reserve, Carnelian Street, 27°30’58”S, 153°4’41”E,  20–22.iv.2001, (T 107499 in QM);  Pullenvale State Forest, 27°30’59”S, 152°51’44”E,  9–11.iv.2001, (T107465–T 107466 in QM);  Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E,  28.vii–1.ix.2003, (T  120590–T   120591in QM);  Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E,  27.v–30.vi.2003, (T   120592in QM); Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E,  1.xii.2003-  2.i.2004, (T   122796in QM);  Sankey’s Scrub, Camp Hill, 27°31’1”S, 153°5’6”E,  20–22.iv.2001, (T 107497 in QM);  Rainbow Forest, Cubberla Creek, 27°31’6”S, 152°57’40”E,  24.xi–21.xii.2005, (T132653–T 132654 in QM);  Rainbow Forest, Cubberla Creek, 27°31’6”S, 152°57’40”E,  24–25.xi.2005, (T132655–T 132657 in QM);  Perry’s Knob,  4 kmNNE Rosewood, 27°36’11”S, 152°36’17”E,  13.i.1999–  16.v.1999, (T107252–T 107278 in QM);  Perry’s Knob,  4 kmNNE Rosewood, 27°36’11”S, 152°36’17”E,  11.xi.1998–  13.i.1999, (T107348, T107370–T 107388 in QM);  Mount Cotton, Scott’s Dam, 27°36’58”S, 153°12’21”E,  12.xii.1997–  7.v.1998, (T107396–T 107402 in QM);  Mount Cotton, upper gully, 27°36’58”S, 153°12’32”E,  12.xii.1997–  7.v.1998, (T107403–T 107416 in QM);  Mount Cotton, 27°36’58”S, 153°12’21”E,  3.ix–12.xii.1997, (T107450–T107461, T 107496 in QM);  Burtonsfirebreak RF (B6), 27°38’6”S, 152°1’41”E,  410m,  2–20.xi.2015, G.B. Monteith& K. Ebert, (25-065488 6 specimensin ANIC);  Mount Tamborine National Park, [ 27°53’11”S, 153°10’51”E],  700 m,  3.iii.1984, L. Masner(CMNEN-00030000 in CMNC);  Cunninghams Gap, 28°3’0”S, 152°24’0”E],  1–2.iii.1973, R.I. Storey(25-058487 1 specimenin QM, 25-058582 1 specimenin QDPC);  Moss’ Wellat Spicer’s Gap, 28°4’28”S, 152°25’23”E,  20.ii–3.iv.1994, (T 107569 in QM);  Black Rock Scrub, 28°7’32”S, 152°39’48”E,  6.x–3.xii.2000, (T107484–T 107489 in QM);  Black Rock Scrub, 28°7’32”S, 152°39’48”E,  2.xii.2000–  13.v.2001, (T107503–T107510, T107513, T107515–T 107516 in QM);  Lamington National Park, IBISCA SiteIQ–700-OF, 28°8’2.4”S, 153°6’’46.8”E,–  30.iii.2007, (T108395–T 108396 in QM);  Lamington National Park, IBISCA SiteIQ–300-D, 28°8’31.2”S, 153°7’58.8”E,  18–23.i.2007, (T 145450 in QM);  Lamington National Park, IBISCA SiteIQ–300-B, 28°9’18”S, 153°8’20.4”E,  16–21.x.2006, (T145447– T 145449 in QM);  Binna Burra, below, 28°11’20”S, 153°11’10”E,  610 m,  29.ii–1.iii.2012, G.B. Monteith(25- 053908 1 specimenalso as COL 2391 in ANIC);  Teviot Falls, via Boonah, 28°13’25”S, 152°28’45”E,  5.x.1975–  22.ii.1976, (T107592–T 107593 in QM);  Springbrook, Mount Wunburra, 28°14’24”S, 153°16’3”E,  1–2.iv.2006, (T 135121 in QM);  Mount French, via Boonah, 28°18’57”S, 152°50’53”E,  5.x.1975–  22.ii.1976, (T107575–T 107591 in QM);   New South Wales: Unumgar State Forestnear Woodenbong, Pole Bridgeroad 788, [ 28°24’0”S, 152°40’0”E],  430 m,  2–11.i.1987, A. Newton& M. Thayer(25-059195 3 specimens, 25-059196 1 specimenin ANIC);  Mount Lindsay Hwy,  13 mi.S of Rathdowney, [ 28°24’0”S, 152°52’0”E],  24–25.ii.1973, R.I. Storey(25- 058492 1 specimenin QM);  Border Ranges National Park  1 kmSE Richmond Gap, [ 28°27’0”S, 152°53’0”E],  350 m,  13.iv.1993, D.S. Chandler(25-059190 1 specimenin ANIC);  Wilson Park,  3 kmSE of Lismore, [ 28°49’0”S, 153°16’0”E],  50 m,  25.viii.1982, S. & J. Peck(25-059188 2 specimensin ANIC);  Iluka Nature Reserve, [ 29°24’0”S, 153°21’0”E],  20–24.xi.1998, G. Williams(25-058088 1 specimenin GWC);  Kangaroo River State Forestalong Burns Roadfrom Lowana, 30°4’30”S, 152°52’5”E,  400m,  21.iii–8.v.1999, S. Lassau& C. Lemann(25-  060779 1 specimenalso as COL 760 in AM); vicinity of Toormina Park, Toormina, [ 30°22’0”S 153°6’0”E],  2.xii.1997, G. Williams(25-058095 1 specimenin GWC);  Scotchman State Forest, Grahamstrail off Horseshoe Road, 30°25’23”S, 152°49’45”E,  23–26.ix.2011, N. Gunter(25-057582 1 specimenalso as COL2006, 25-057583 1 specimenalso as COL 2007 in ANIC);  Newry State Forest, Hungry Head Road, 30°31’41”S, 153°0’22”E,  23–26.ix.2011, N. Gunter(25-057580 5 specimensalso as COL2004, 25-057581 1 specimenalso as COL 2005 in ANIC);  Walcha Survey Big Hill, 30°37’46”S, 152°14’6”E,  i.1993, I. Oliver, (25-  060781 2 specimensin AM);  Racecourse Headland S of Crescent Head[Racecourse Head], [ 31°15’0”S, 152°58’0”E],  20.i.1999, G. & T. Williams(25-058089 1 specimenin GWC);  Point Plomer, Limeburners Creek National Park N of Port Macquarie, [ 31°19’0”S, 152°58’0”E],  23– 24.x.2010, G. & T. Williams(25-  060888 1 specimenalso as COL 1195 in ANIC);  Winmurra, approximately  50 kmNW Taree, Doyles River, [ 31°34’0”S, 152°6’0”E],  530 m, 2002, 2008, D. Bray(25-053894 1 specimenin GWC); N end of Easternfire trail, Yarratt State Forest, NW of Wingham, [ 31°44’0”S, 152°26’0”E],  8.x.1981, G. & T. Williams(25-057875 1 specimenin ANIC, 25-058090 2 specimensin GWC);  Lorien Wildlife Refuge  3 kmN Lansdowne[ 31°45’0”S, 152°32’0”E],  17.x.2010, G. & T. Williams(25-  060887 1 specimenalso as COL 1194 in ANIC);   4 kmW Lansdowne, Lorein, 152°32’0”E, 31°45’0”S,  12.v–12.vi.1986, H.F. & A.T. Howden, (CMNEN-00030009- 00030010 in CMNC);  4kmW. Lansdowne, Lorien, 152°32’0”E, 31°45’0”S,  1.iv.1987, H.F. & A.T. Howden, (CMNEN-00030011-00030018 in CMNC);  Newbys Lane, Lansdowne, [ 31°47’0”S, 152°32’0”E],  12–14.iv.1981, G. & T. Williams(25-057874 1 specimenin ANIC);  Lorien, Lansdowne, [ 31°52’0”S, 152°29’0”E],  10–12.iv.1981, G. & T. Williams(25-058097 2 specimensin GWC);  Lansdowne Recreational ReserveSE Lansdownevia Taree, [ 31°47’0”S, 152°32’0”E],  24–25.x.1984, G. Williams(25-058092 1 specimenin GWC); approximately  24 kmNNW Gloucester, [ 31°49’0”S 151°52’0”E],  19–20.xii.1991, G. & T. Williams(25-058094 1 specimenin GWC);   3.5 kmNE of Harrington, [ 31°50’0”S, 152°43’0”E],  18.viii.1990, G. & T. Williams(25-  060778 1 specimenalso as COL 759 in AM);  Crowdyhead approximately  7 kmN of Harrington, [ 31°51’0”S, 152°45’0”E],  28.ix.1982, G. & T. Williams(25-058100 2 specimensin GWC);  Harrington, [ 31°52’0”S, 152°42’0”E],  14.xi.1976, G. WilliamsSAMA25-027647 2 specimensin SAM);  Harrington, [ 31°52’0”S, 152°42’0”E],  25.xii.1976, G. Williams(25- 058104 1 specimenin GWC);  Harrington, [ 31°52’0”S, 152°42’0”E],  20.xii.1976, G. Williams(25-059193 1 specimenin ANIC);  Harrington, [ 31°52’0”S, 152°42’0”E],  10.x.1976, G. Williams(25-059194 1 specimenin ANIC);  Kiwarrak State Forest, [ 31°57’0”S, 152°27’0”E],  25.v.1992, T. Gush(25-059191 1 specimenin ANIC);  Kiwarrak State Forestnear Taree, [ 31°58’0”S, 152°26’0”E],  19.ix.1979, W. King(25-  060786 1 specimenalso as COL777 and K 266951 in AM);  Kiwarrak State Forest, [ 31°58’0”S, 152°28’0”E],  29.v–1.vi.1993, V. Lorimer(25-058712 2 spec- imens in ANIC);  Kiwarrak State Forest, via Taree, approximately  1 kmN of Breakneck Lookout, [ 31°59’0”S, 152°25’0”E],  17.ii.1982, G. & T. Williams(25-057873 1 specimenin ANIC, 25-058102 2 specimensin GWC);  Saltwatervicinity of Old Bar, [ 32°0’0”S, 152°34’0”E],  22.x.1981, G. & T. Williams(25-058096 1 specimenin GWC);  Black Head( Hallidays Point), [ 32°4’0”S, 152°33’0”E],  13.x.1981, G. & T. Williams(25-058101 2 specimensin GWC);  Gloucester River, Barrington Tops National Park, [ 32°4’0”S, 151°41’0”E],  12–14.xi.1981, T.A. Weir(25-059189  x 1in ANIC); on Mountain Road 0.2 kmoff junction with Kunungra Road, 32°8’22”S, 151°44’4”E,  4.ii–9.iv.1993, G. Cassis& M. Gray(25-  060775 1 specimenalso as COL756 and K 266946 in AM);  Southbase of Cape Hawkevia Forster, [ 32°13’0”S, 152°34’0”E],  30.ix.1981, G. & T. Williams(25-058099 1 specimenin GWC);  O’Sullivan Gap Reserve,  11 kmNE Bulahdelah, [ 32°20’0”S, 152°17’0”E],  50 m,  11.vi–27.viii.1982, S. & J. Peck(25-059187 2 specimensin ANIC);  Bombah Point Swamp[ 32°30’0”S, 152°17’0”E],  1.v.1993, V. Lorimer(25- 058713 1 specimenin ANIC);  Mungo Brush, Myall Lakes National Park, [ 32°32’0”S, 152°19’0”E],  26.x.1981, G. & T. Williams(25-057872 2 specimensin ANIC, 25-058103 2 specimensin GWC);  Mungo Brush, [ 32°36’0”S, 152°16’0”E],  1.v.1993, V. Lorimer(25-058711 2 specimensin ANIC);  Mount Coricudgynear Olinda Station, [ 32°50’0”S, 150°21’0”E],  5–9.iii.2002, G. Hangay(25-057878 2 specimens, 25-057879 1 specimenin ANIC);  Gowan, [ 33°12’0”S, 149°22’0”E],  25.vii.1976, B.B. Lowery(25-057871 4 specimensin ANIC);  Mount Irvine, [ 33°29’0”S, 150°28’0”E],  5.i.1980, G. & T. Williams(25-058098 1 specimenin GWC);  Southslope, Hat Hill, Hat Head, [ 33°36’0”S, 150°20’0”E],  4–5.ii.1985, G. Williams(25-058091 2 specimensin GWC);  Ingleside, Katandra Bushland Sanct., [ 33°41’0”S, 151°16’0”E],  xii.1998, G. Hangay(25-059192 1 specimenin ANIC);  Sassafras Gullyvia Springwood, [ 33°42’0”S, 150°32’0”E],  26.xii.1979, G. & T. Williams(25-058093 2 specimensin GWC);  West Pennant Hills, [ 33°45’0”S, 151°2’0”E],  xii.1995, G. O’Reilly(25-  060780 1 specimenalso as COL761 and K 266947 in AM);  West Killara, [ 33°46’0”S, 151°7’0”E],  7.vii.1991, T. Gush(25-057880 2 specimensin ANIC);   5 mi.N Batemans Bay, [ 35°40’0”S, 150°15’0”E],  15.xii.1967, E. Matthews(25-027645 1 specimen, 25-027646 1 specimenin SAMA);   5 mi.N Batemans Bay, [ 35°40’0”S, 150°15’0”E],  6.iii.1978, S. & J. Peck(CMNEN-00030005 in CMNC);  Gerroa, [ 34°46’33”S, 150°47’49”E],  22.xii.1974, H.F. & A.T. Howden(CMNEN-00030020-00030024 in CMNC);  Princes Highway Corunna State Forest, 36°16’48”S, 150°7’33”E,  12.iii.1999, L. Wilkie, R. Harris, & H. Smith(25-  060787 50 specimensand K266949 also as COL778-COL 780 in ANIC);   Victoria: Mallacoota National Park, [ 37°30’0”S, 149°46’0”E],  13.i.1968, E. Matthews(25-027649 1 specimenin SAMA);  Mallacoota National Park, [ 37°30’0”S, 149°46’0”E],  21–26.v.1978, S. & J. Peck(25-057869 1 specimenin ANIC);  Mallacoota National Park, [ 37°30’0”S, 149°46’0”E],  26.v.1978, S. & J. Peck(25-057870 1 specimenin ANIC);  Croajingolong National Park, South West Arm Road,  11 kmNW Mallacoota, [ 37°31’0”S, 149°41’0”E],  20 m,  10–23.ii.1993, A. Newton& M. Thayer(25-057876 2 specimens, 25-057877 1 specimenin ANIC);  Mallacoola National Park, Double Creek, [ 37°31’47”S, 149°41’13”E],  3 m,  22.v.1978–  26.v.1978, S. & J. Peck(CMNEN-00030006 in CMNC).   Description. Holotypeuniformly dark brown almost black in colour. Rest of specimens range from light brown to black. Antennal clubs grey. Total length: 2.7–3.5 mm. Holotypemeasurements: Total length 3.1 mm, elytral width 2.0 mm.  Male. Head: Surface with fine punctation. Basal carina absent. Dorsal part of the eyes narrow, separated by an interocular space approximately 16 times the eye width (40: 2.5); eye canthus dividing eye. Head 1.4 x wider than long (49:35). Prothorax: Pronotal surface with very fine punctation anteriorly to fine punctation posteriorly. Anterior angles sharp (less than 90°). Basal margin slightly angulate. Hypomeral surface reticulate. Hypomeral striae over half the length of hypomeron and very close to lateral margin, almost appearing double. Pronotum 1.7 x wider than long (40:24). Elytra: Surface smooth with impressed, impunctate striae, striae 1–3 or 1–4 appearing double, 5–7 appearing single; intervals very finely punctate. Stria 6 extending to elytral base and 7 not extending to elytral base. Striae 8 slightly curving inwards posteriorly. Elytra almost equal in width and length (48:45). Legs: Front edge of protibiae deeply angulate. Abdomen: Pygidium simple, smooth, finely punctured and slightly convex. Abdominal ventrites reticulate right across. Ventrite 6 very finely punctate. Pterothorax: Medial lobe of metaventrite punctate and broadly margined between the mesocoxae. Lateral lobe of metaventrite punctate, inner border next to mesocoxae anteriorly straight and lateral border virtually absent. Mesoventrite notched anteriorly. Mesoventrite smooth and nitid with very fine punctation. Mesepimeron very finely reticulate. Metanepisternum reticulate.   FIGURE 1. Some diagnostic characters of the  Lepanus australisspecies group. A, Male protibia of  Lepanus eungelladotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of  Lepanus eungelladotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of  Lepanus australisdotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of  Lepanus australisdotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of  Lepanus dukungarriwhite arrow points to row of punctures on metanepisternum; F, lateral view of  Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of  Lepanus oxleyiwhite arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of  Lepanus dukungarrishowing medial expansion; I, Head of  Lepanus oxleyiwhite arrow points to interrupted basal carina; J, lateral view of  Lepanus latheticusdotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K,  Lepanus australisat bird droppings (image: Geoff Williams).  Female. As per  L. australisspecies group.   Distribution.  Lepanus australishas the one of the broadest distributions of the genus ( Fig. 5A) and has been recorded in eastern Australiafrom as far north as St Lawrence, Queenslandto Mallacoota, Victoria( 2000 kmsouth). It is known from a variety of habitats (rainforest, palm forest, wet and dry sclerophyll forests) in the following Interim Biogeographic Regionalisation for Australia(IBRA) bioregions: South Eastern Queensland(SEQ), NSW North Coast (NNC), New EnglandTablelands (NET), Sydney Basin (SYB), South East Corner (SEC), South Eastern Highlands (SEH), the most easterly extents of the Brigalow Belt North (BBN), and Brigalow Belt South (BBS). BIOCLIM modelling suggests appropriate climatic conditions extend further inland than currently recorded, however despite extensive sampling in some of the adjacent IBRA regions,  L. australishas not been collected.   Comments.Most specimens have been collected in baited pitfall traps (mushroom, dung, or carrion), but it has also been collected in berlesed litter, at faeces, and in flight intercept traps. Molecular sequences have been generated from numerous locations across the range. The phylogeny of Gunter & Weir (2017)includes representatives from three Queenslandpopulations: Kenmore, Brisbane (25- 060906also as COL1197), Saddletree Creek, Bunya Mountains (25- 060835also as COL1470), and Tinana Creek near Maryborough (25-0053884 also as COL2345) but other specimens from New South have also been sequenced and confirm the wide distribution of the species. The specimen from Kenmore was included in the phylogeny of Gunter et al.(2019) and it formed a relationship with an undescribed species in the  L. pisoniaespecies group informally referred to as  Lepanusspecies NQ19.  Lepanus australisis one of several species of  Lepanusthat have been observed at bird droppings ( Fig. 1K), previously thought to be a food source underutilised by dung beetles. 2448758245 1967-08-16 ANIC N. Batemans Bay & E. Matthews Batemans Bay 3 204 1 1 holotype 2448757220 1967-12-29 ANIC E. Matthews -34.566666 Macquarie Pass 19 150.63333 4 205 5 1 paratype 2448759213 1968-01-09 1968-01-12 1968-01-09 ANIC E. Matthews -35.55 Clyde Mountain 19 149.95 4 205 1 paratype 2448756216 1968-01-13 ANIC E. Matthews -37.0 Eden 19 149.91667 4 205 4 2 paratype 2448756223 1968-01-13 ANIC E. Matthews -37.233334 Eden 19 149.91667 4 205 1 paratype 2448756221 1968-01-13 ANIC E. Matthews -37.5 Mallacoota National Park 19 149.76666 4 205 1 paratype 2448758228 2000-06-29 2000-10-22 2000-06-29 QM -22.351112 St Lawrence 21 149.3425 4 205 1 Queensland 2448757221 [220,1217,476,503] 2000-10-20 2000-12-20 2000-10-20 QM -25.508888 Fairlie's Knob 21 152.28944 4 205 1 Queensland 2448756217 2000-12-20 2001-03-23 2000-12-20 QM Fairlie's Knob 4 205 1 Queensland 2448759212 2001-03-22 2001-03-23 2001-03-22 QM Fairlie's Knob 4 205 1 Queensland 2448757217 1975-12-03 QM G. Thompson & J. A. Slater -25.15 Fraser Island 21 153.21666 Lake Bowaddy 4 205 1 Queensland 2448759216 1975-12-01 1975-12-04 1975-12-01 QM G. Thompson & J. A. Slater -25.316668 Fraser Island 21 153.18333 Yidney Scrub 4 205 2 Queensland 2448756219 2000-01-15 CMNC R. A. Anderson 350 -26.647223 Booloumba Creek State Forest River 20 152.64583 4 205 1 Queensland 2448756218 2011-11-22 2011-12-07 2011-11-22 ANIC G. B. Monteith 35 -25.751944 Tinana Creek 21 152.6939 4 205 1 Queensland 2448759223 2011-12-06 2011-12-07 2011-12-06 ANIC G. B. Monteith 35 -25.756945 Tallegalla Weir Junction 21 152.69695 4 205 1 Queensland 2448758226 [337,1432,764,791] 1995-11-24 1996-02-03 1995-11-24 QM -26.131945 Nangur State Forest 20 151.9786 4 205 1 Queensland 2448759222 1995-01-26 1995-04-20 1995-01-26 QM -26.154722 Park on Boat Mountain 20 151.90944 Jack Smith 4 205 1 Queensland 2448758223 1994-12-15 1995-01-26 1994-12-15 QM -26.154722 Park on Boat Mountain 20 151.90944 Jack Smith 4 205 1 Queensland 2448758219 2006-02-18 2006-02-19 2006-02-18 QM -26.237501 Kin Kin Creek 20 152.98111 4 205 1 Queensland 2448758224 2006-02-17 QM -26.257778 National Park Office 20 152.99556 Elanda Point 4 205 1 Queensland 2448757218 1973-03-10 2012-10 1973-03-10 QM, QDPC R. I. Storey 1300 -26.35 Gympie 20 152.78334 Cooran Plateau 4 205 2 Queensland 2448758221 1973-10-03 1973-11-03 1973-10-03 CMNC R. I. Storey 396 -26.35 Gympie 20 152.78334 Cooran Plateau 4 205 1 Queensland 2448758218 2005-12-23 2005-12-26 2005-12-23 QM -26.580002 Kingaroy 20 151.87639 4 205 1 Queensland 2448758220 1991-11-30 1992-01-08 1991-11-30 QM -26.630278 Mapleton Falls National Park 20 152.83916 4 205 1 Queensland 2448759219 1978-09-06 ANIC Common -26.683332 4 km W Buderim 20 153.0 4 205 1 Queensland 2448759221 [416,1432,1124,1151] 1978-05-12 1978-09-27 1978-05-12 QM -26.700834 Nambour 20 153.00917 Chevallum 4 205 1 Queensland 2448758227 [151,1236,1160,1187] 1995-04-20 1995-05-02 1995-04-20 QM -26.712778 Site 20 152.6189 Conondale Range 4 205 1 Queensland 2448758217 1991-11-29 1992-01-08 1991-11-29 QM -26.78111 Mary Cairncross Park 20 152.88222 4 205 1 Queensland 2448759211 2010-08-28 83-06-07 2010-08-28 ANIC G. B. Monteith -26.789444 Landsborough 20 152.97804 4 205 1 Queensland 2448758216 2010-12-29 2010-12-30 2010-12-29 ANIC G. B. Monteith & K. Will -26.86389 Saddletree Creek 20 151.63388 4 205 1 Queensland 2448757222 2010-12-22 2010-12-29 2010-12-22 ANIC Sota & Takami -26.86389 Saddletree Creek 20 151.63388 4 205 1 Queensland 2448759217 2010-12-22 35-06-08 2010-12-22 ANIC G. B. Monteith & K. Will -26.86389 Saddletree Creek 20 151.63388 4 205 7 Queensland 2448757214 1982-06-19 ANIC Hill -26.866667 5 km E Yarraman 20 152.05 4 205 1 Queensland 2448759215 1995-11-24 1996-02-03 1995-11-24 QM -26.877222 Top of Blackbutt Range 20 152.19223 4 205 1 Queensland 2448756220 05-11-15 2007-09-20 05-11-15 QM -26.879723 Dandabah 20 151.59555 Bunya Mountains 4 205 1 Queensland 2448758222 1983-03-30 QM -26.879723 Upper Yarraman State Forest 20 151.88445 4 205 1 Queensland 2448757215 2000-01-18 CMNC R. A. Anderson 1100 -26.864166 Westcott Campground 20 151.57138 Bunya Mountain National Park 4 205 1 Queensland 2448757213 2000-01-19 CMNC R. A. Anderson 1100 -26.881666 Dandabah Picnic Area 20 151.58972 Bunya Mountain National Park 4 205 1 Queensland 2448756222 1982-06-19 ANIC L. Hill -26.883333 Blackbutt 20 152.16667 4 205 1 Queensland 2448757216 1975-01-11 QM G. B. Monteith -26.9 Maidenwell 20 151.9 Upper Yarraman State Forest 4 205 3 Queensland 2448759218 08-11-15 2007-11-16 08-11-15 QM -26.92389 Wild Horse Mountain 20 152.99 4 205 1 Queensland 2448757219 2008-01-23 2008-02-13 2008-01-23 QM -26.92389 Wild Horse Mountain 20 152.99 4 205 1 Queensland 2448758225 2010-10-08 2010-10-09 2010-10-08 ANIC G. B. Monteith 10 -27.015 Bribie Island 20 153.125 4 205 1 Queensland 2448759214 2010-10-08 77-06-08 2010-10-08 ANIC G. B. Monteith -27.018055 Bribie Island 20 153.12306 4 205 1 Queensland 2448757212 1998-09-18 1998-10-29 1998-09-18 QM P. Bouchard Brisbane -27.016666 Scrub Creek 20 152.43333 Forest Park 4 205 1 Queensland 2448758229 2002-10-01 2002-10-15 2002-10-01 CMNC J. Skevington & J. Cumming -27.428055 Brisbane Forest 20 152.83833 4 205 1 Queensland 2448759220 1973-12-22 1973-12-23 1973-12-22 QDPC R. I. Storey -27.05 Bottom 20 151.91667 Blackbutt Range 4 205 2 Queensland 2448758215 1998-12-30 1999-03-26 1998-12-30 QM 5 206 -27.228056 Site 20 152.26 Mount Deongwar 4 205 1 Queensland 2448756207 1998-10-14 1998-12-30 1998-10-14 QM -27.228056 Site 20 152.26 Mount Deongwar 5 206 1 Queensland 2448759205 1998-12-30 1999-03-26 1998-12-30 QM -27.230278 Site 20 152.25694 Mount Deongwar 5 206 1 Queensland 2448759208 1998-10-14 1998-12-31 1998-10-14 QM -27.230278 Site 20 152.25694 Mount Deongwar 5 206 1 Queensland 2448758210 2007-04-16 2007-04-28 2007-04-16 QM -27.331667 Mount Glorious 20 152.75333 5 206 1 Queensland 2448757210 [223,1236,296,323] 2008-03-12 2008-03-15 2008-03-12 QM -27.331667 Mount Glorious 20 152.75333 5 206 1 Queensland 2448757202 2003-12-04 87-12-06 2003-12-04 QM Crows Nest National Park -27.31889 Site Perseverance Section 5 206 1 Queensland 2448758212 1986-11-14 1987-01-30 1986-11-14 ANIC T. Hiller 630 -27.333334 Mount Glorious 20 152.76666 5 206 5 Queensland 2448757206 1989-02 CMNC H. F. & A. T. Howden 635 -27.333334 Mount Glorious 20 152.76666 5 206 1 Queensland 2448756210 1994-10-22 1995-02-02 1994-10-22 QM -27.33889 Samford 20 152.79779 Stony Creek 5 206 1 Queensland 2448757207 QM -27.33889 Samford 20 152.79779 Stony Creek 5 206 1 Queensland 2448759210 [363,1432,512,539] 2006-01-28 2006-01-29 2006-01-28 QM -27.3525 Mount Glorious 20 152.7686 5 206 1 Queensland 2448757209 [151,1300,548,575] 2003-10-01 30-12-21 2003-10-01 QM -27.402222 Site 20 152.78944 Boombana National Park 5 206 1 Queensland 2448759201 2004-02-15 32-12-21 2004-02-15 QM -27.402222 Site 20 152.78944 Boombana National Park 5 206 1 Queensland 2448756202 2004-01-02 95-12-27 2004-01-02 QM -27.402222 Site 20 152.78944 Boombana National Park 5 206 1 Queensland 2448758204 2012-02-18 2012-02-19 2012-02-18 ANIC G. B. Monteith 230 -27.431944 Boombana National Park 20 152.89694 5 206 1 Queensland 2448759203 1999-10-18 2000-02-06 1999-10-18 QM -27.432777 McAfee's Lookout 20 152.87056 5 206 1 Queensland 2448758203 1999-10-18 2000-02-06 1999-10-18 QM -27.4375 McAfee's Lookout 20 152.87056 5 206 1 Queensland 2448758205 1999-12-21 2000-01-27 1999-12-21 QM -27.443611 Site 20 152.90388 Enoggera Reservoir 5 206 1 Queensland 2448756206 2000-01-27 2000-03-15 2000-01-27 QM -27.443611 Site 20 152.90388 Enoggera Reservoir 5 206 1 Queensland 2448758211 2000-03-15 2000-05-18 2000-03-15 QM -27.443611 Site 20 152.90388 Enoggera Reservoir 5 206 1 Queensland 2448756212 1999-10-16 1999-12-21 1999-10-16 QM -27.445555 Site 20 152.92195 Enoggera Reservoir 5 206 1 Queensland 2448758213 [222,1316,872,899] 1999-12-21 2000-01-27 1999-12-21 QM -27.445555 Enoggera Reservoir 20 152.92195 5 206 1 Queensland 2448759202 2000-03-15 2000-05-18 2000-03-15 QM -27.445555 Enoggera Reservoir 20 152.92195 5 206 1 Queensland 2448756203 1999-07-06 1999-10-16 1999-07-06 QM -27.435555 McAfee's Lookout 20 152.87138 5 206 1 Queensland 2448757204 06-11-15 2007-10-14 06-11-15 QM -27.461945 Site 20 152.8725 Gold Creek Reservoir 5 206 1 Queensland 2448756205 07-11-15 2007-10-14 07-11-15 QM -27.462778 Gold Creek Reservoir 20 152.88167 5 206 1 Queensland 2448756213 [373,1228,1016,1043] 2001-02-06 2001-02-07 2001-02-06 QM -27.47889 Slaughter Falls 20 152.95944 5 206 1 Queensland 2448758202 2011-02-12 2011-02-13 2011-02-12 ANIC G. B. Monteith -27.4825 Brookfield 20 152.91695 Boscombe Road 5 206 1 Queensland 2448758214 [246,1223,1088,1115] 2003-11-04 31-12-21 2003-11-04 QM -27.468056 Site 20 152.87222 Gold Creek Reservoir 5 206 1 Queensland 2448757201 1975-08-24 1975-09-07 1975-08-24 QM R. I. Storey -27.483334 Brisbane 20 152.86667 Upper Brookfield Road 5 206 1 Queensland 2448758201 1975-09-22 1975-10-04 1975-09-22 QM R. I. Storey -27.483334 Brisbane 20 152.86667 Upper Brookfield Road 5 206 5 Queensland 2448756214 06-06-09 2010-11-17 06-06-09 ANIC -27.491112 Kenmore Kookaburra Street 20 152.92307 5 206 1 Queensland 2448756209 2011-01-20 2011-01-21 2011-01-20 ANIC G. B. Monteith -27.491112 Kenmore Hills 20 152.92334 5 206 2 Queensland 2448756208 2011-01-20 2011-01-21 2011-01-20 ANIC G. B. Monteith -27.506666 Kenmore 20 152.93112 5 206 2 Queensland 2448759209 2010-11-16 89-06-08 2010-11-16 ANIC G. B. Monteith -27.506945 Kenmore 20 152.93112 Mabb Street 5 206 1 Queensland 2448759207 1999-01-13 1999-05-16 1999-01-13 QM -27.5075 The Knobby 20 152.58833 5 206 1 Queensland 2448758208 1999-01-13 1999-05-16 1999-01-13 QM -27.508055 Cameron's Scrub 20 152.73166 5 206 1 Queensland 2448759206 2012-04-30 2012-05-01 2012-04-30 ANIC G. B. Monteith 75 -27.508888 Cameron's Scrub 20 152.73 5 206 7 Queensland 2448757205 1999-11-01 1999-11-04 1999-11-01 QM -27.509445 Long Pocket 20 152.99806 5 206 1 Queensland 2448758207 1999-01-13 1999-05-16 1999-01-13 QM -27.509722 Cameron's Scrub 20 152.73166 5 206 1 Queensland 2448756211 1998-11-11 1999-01-13 1998-11-11 QM -27.509722 Cameron's Scrub 20 152.73166 5 206 1 Queensland 2448757208 1999-01-13 1999-05-16 1999-01-13 QM -27.5125 Cameron's Scrub 20 152.7275 5 206 1 Queensland 2550750804 [220,1303,1592,1619] 1997-12-12 1998-05-07 1997-12-12 QM -27.513334 Belmont Hill Belmont Hill 5 206 1 2550750808 1999-01-13 1999-05-16 1999-01-13 QM -27.51361 Snig 152.7286 Scrub 5 206 1 2550750802 1998-11-11 1999-01-13 1998-11-11 QM -27.51361 Snig 152.7286 Scrub 5 206 1 2550750811 2001-04-20 2001-04-22 2001-04-20 QM -27.516111 Carnelian Street 153.07805 Whites Hill Reserve 5 206 1 2550750812 2001-04-09 2001-04-11 2001-04-09 QM -27.516388 Pullenvale State Forest 152.86223 5 206 1 2550750801 2003-07-28 91-12-05 2003-07-28 QM -27.5025 Carindale Bulimba Creek 5 206 1 2448758209 2003-05-27 96-12-27 2003-05-27 QM -27.5025 Carindale Bulimba Creek 5 206 1 Queensland 2448756201 2001-04-20 2001-04-22 2001-04-20 QM -27.516945 Camp Hill 20 153.08499 Sankey's Scrub 5 206 1 Queensland 2448756204 2005-11-24 2005-12-21 2005-11-24 QM -27.518333 Cubberla Creek 20 152.9611 Rainbow Forest 5 206 1 Queensland 2448757203 2005-11-24 2005-11-25 2005-11-24 QM -27.518333 Cubberla Creek 20 152.9611 Rainbow Forest 5 206 1 Queensland 2448758206 1999-01-13 1999-05-16 1999-01-13 QM -27.603056 Perry's Knob 20 152.60472 5 206 1 Queensland 2448759204 1998-11-11 1999-01-13 1998-11-11 QM -27.603056 Perry's Knob 20 152.60472 5 206 1 Queensland 2448756215 1997-12-12 1998-05-07 1997-12-12 QM 6 207 -27.616112 Scott's Dam 20 153.20583 Mount Cotton 5 206 1 Queensland 2448759236 1997-12-12 1998-05-07 1997-12-12 QM -27.616112 Mount Cotton 20 153.2089 6 207 1 Queensland 2448758237 1997-09-03 1997-12-12 1997-09-03 QM -27.616112 Mount Cotton 20 153.20583 6 207 1 Queensland 2448757231 2015-11-02 2015-11-20 2015-11-02 ANIC G. B. Monteith & K. Ebert 410 -27.635 Burtons 20 152.02805 6 207 6 Queensland 2448759235 1984-03-03 CMNC L. Masner 700 -27.886389 Mount Tamborine National Park 20 153.18083 6 207 1 Queensland 2448759234 1973-03-01 1973-03-02 1973-03-01 QM, QDPC R. I. Storey -28.05 Cunninghams Gap 20 152.4 6 207 2 Queensland 2448756238 [401,1432,368,395] 1994-02-20 1994-04-03 1994-02-20 QM -28.074446 Spicer's Gap 20 152.42307 Moss' Well 6 207 1 Queensland 2448757240 [151,1199,404,431] 2000-10-06 2000-12-03 2000-10-06 QM -28.125555 Black Rock Scrub 20 152.66333 6 207 1 Queensland 2448756234 2000-12-02 2001-05-13 2000-12-02 QM -28.125555 Black Rock Scrub 20 152.66333 6 207 1 Queensland 2448759233 2007-03-30 QM -28.134 Site Lamington National Park 6 207 1 Queensland 2448758244 2007-01-18 2007-01-23 2007-01-18 QM -28.142 Site 1 153.133 Lamington National Park 6 207 1 Queensland 2448759241 2006-10-16 2006-10-21 2006-10-16 QM -28.154999 Site 15 153.13899 Lamington National Park 6 207 1 Queensland 2448757239 2012-02-29 2012-03-01 2012-02-29 ANIC G. B. Monteith 610 -28.188889 Binna Burra 20 153.18611 6 207 1 Queensland 2448759237 1975-10-05 1976-02-22 1975-10-05 QM -28.223612 Boonah 20 152.47916 Teviot Falls 6 207 1 Queensland 2448759238 2006-04-01 2006-04-02 2006-04-01 QM -28.24 Mount Wunburra 20 153.2675 Springbrook 6 207 1 Queensland 2448756241 1975-10-05 1976-02-22 1975-10-05 QM -28.315832 Boonah 20 152.84805 Mount French 6 207 1 Queensland 2448757238 1987-01-02 1987-01-11 1987-01-02 ANIC A. Newton & M. Thayer Unumgar State Forest 430 -28.4 Pole Bridge 20 152.66667 Woodenbong 6 207 4 New South Wales 2448757241 1973-02-24 1973-02-25 1973-02-24 QM R. I. Storey -28.4 Mount Lindsay Hwy 20 152.86667 6 207 1 New South Wales 2448757237 1993-04-13 ANIC D. S. Chandler 350 -28.45 Border Ranges National Park 20 152.88333 6 207 1 New South Wales 2448757233 1982-08-25 ANIC S. & J. Peck 50 -28.816668 Wilson Park 20 153.26666 6 207 2 New South Wales 2448756236 1998-11-20 1998-11-24 1998-11-20 GWC G. Williams -29.4 Iluka Nature Reserve 20 153.35 6 207 1 New South Wales 2448756235 1999-03-21 79-06-07 1999-03-21 AM S. Lassau & C. Lemann Kangaroo River State Forest 400 -30.075 Lowana 20 152.86806 Burns Road 6 207 1 New South Wales 2448757236 1997-12-02 GWC G. Williams -30.366667 Toormina 20 153.1 Toormina Park 6 207 1 New South Wales 2448758240 2011-09-23 2011-09-26 2011-09-23 ANIC N. Gunter Scotchman State Forest -30.423056 Horseshoe Road 20 152.82916 Grahams 6 207 2 New South Wales 2448759239 2011-09-23 2011-09-26 2011-09-23 ANIC N. Gunter -30.528055 Hungry Head Road 20 153.0061 Newry State Forest 6 207 6 New South Wales 2448757230 1993-01 81-06-07 1993-01 AM I. Oliver -30.629444 Walcha Survey Big Hill 20 152.235 6 207 2 New South Wales 2448758236 1999-01-20 GWC G. & T. Williams -31.25 Racecourse Headland 20 152.96666 6 207 1 New South Wales 2448756243 2010-10-23 88-06-08 2010-10-23 ANIC G. & T. Williams -31.316668 Limeburners Creek National Park 20 152.96666 Point Plomer 6 207 1 New South Wales 2448758235 GWC D. Bray 530 -31.566668 Doyles River 20 152.1 Winmurra 6 207 1 New South Wales 2448759240 1981-10-08 ANIC, GWC G. & T. Williams -31.733334 Yarratt State Forest 20 152.43333 Eastern 6 207 3 New South Wales 2448757232 2010-10-17 87-06-08 2010-10-17 ANIC G. & T. Williams -31.75 Lorien Wildlife Refuge 20 152.53334 6 207 1 New South Wales 2448756242 1986-05-12 1986-06-12 1986-05-12 CMNC H. F. & A. T. Howden -31.75 Lorein 20 152.53334 6 207 1 New South Wales 2448756239 1987-04-01 CMNC H. F. & A. T. Howden -31.75 Lorien 20 152.53334 Lansdowne 6 207 1 New South Wales 2448758238 1981-04-12 1981-04-14 1981-04-12 ANIC G. & T. Williams -31.783333 Lansdowne 20 152.53334 Newbys Lane 6 207 1 New South Wales 2448758239 1981-04-10 1981-04-12 1981-04-10 GWC G. & T. Williams -31.866667 Lansdowne 20 152.48334 Lorien 6 207 2 New South Wales 2448756237 1984-10-24 1984-10-25 1984-10-24 GWC G. Williams Lansdowne Recreational Reserve -31.783333 Taree 20 152.53334 Lansdowne 6 207 1 New South Wales 2448757229 1991-12-19 1991-12-20 1991-12-19 GWC Williams -31.816668 24 km NNW Gloucester 20 151.86667 6 207 1 New South Wales 2448757228 1990-08-18 78-06-07 1990-08-18 AM Williams -31.833334 3.5 km NE of Harrington 20 152.71666 6 207 1 New South Wales 2448758242 1982-09-28 GWC G. & T. Williams -31.85 Crowdy 20 152.75 6 207 2 New South Wales 2448757235 1976-11-14 SAM G. Williams -31.866667 Harrington 20 152.7 6 207 2 New South Wales 2448757234 1976-12-25 GWC G. Williams -31.866667 Harrington 20 152.7 6 207 1 New South Wales 2448758241 1976-12-20 ANIC G. Williams -31.866667 Harrington 20 152.7 6 207 1 New South Wales 2448756240 [331,1432,1909,1936] 1976-10-10 ANIC G. Williams -31.866667 Harrington 20 152.7 6 207 1 New South Wales 2448758243 [151,1319,1945,1972] 1992-05-25 ANIC T. Gush -31.95 Kiwarrak State Forest 20 152.45 6 207 1 New South Wales 2448757227 1979-09-19 86-06-07 1979-09-19 AM W. King -31.966667 Taree 20 152.43333 Kiwarrak State Forest 6 207 1 New South Wales 2448756233 1993-05-29 1993-06-01 1993-05-29 ANIC V. Lorimer 7 208 -31.966667 Kiwarrak State Forest 20 152.46666 6 207 1 New South Wales 2448756232 1982-02-17 ANIC, GWC G. & T. Williams -31.983334 Taree 20 152.41667 Kiwarrak State Forest 7 208 3 New South Wales 2448758233 1981-10-22 GWC G. & T. Williams -32.0 Old Bar 20 152.56667 Saltwater 7 208 1 New South Wales 2448758234 1981-10-13 GWC G. & T. Williams -32.066666 Hallidays Point 20 152.55 Black Head 7 208 2 New South Wales 2448756231 1981-11-12 1981-11-14 1981-11-12 ANIC T. A. Weir -32.066666 Barrington Tops National Park 20 151.68333 Gloucester River 7 208 1 New South Wales 2448756230 1993-02-04 75-06-07 1993-02-04 AM G. Cassis & M. Gray -32.139446 Kunungra Road 20 151.73445 Mountain Road 7 208 1 New South Wales 2448756224 1981-09-30 GWC G. & T. Williams South -32.216667 Forster 20 152.56667 Cape Hawke 7 208 1 New South Wales 2448759231 1982-06-11 1982-08-27 1982-06-11 ANIC S. & J. Peck 50 -32.333332 O'Sullivan Gap Reserve 20 152.28334 7 208 2 New South Wales 2448758231 1993-05-01 ANIC V. Lorimer -32.5 Bombah Point Swamp 20 152.28334 7 208 1 New South Wales 2448757225 1981-10-26 ANIC, GWC G. & T. Williams -32.533333 Myall Lakes National Park 20 152.31667 Mungo Brush 7 208 4 New South Wales 2448758230 1993-05-01 ANIC V. Lorimer -32.6 Mungo Brush 20 152.26666 7 208 2 New South Wales 2448759228 2002-03-05 2002-03-09 2002-03-05 ANIC G. Hangay -32.833332 Olinda Station 20 150.35 Mount Coricudgy 7 208 3 New South Wales 2448757226 [151,1263,656,683] 1976-07-25 ANIC B. B. Lowery -33.2 Gowan 20 149.36667 7 208 4 New South Wales 2448756228 1980-01-05 GWC G. & T. Williams -33.483334 Mount Irvine 20 150.46666 7 208 1 New South Wales 2448756229 1985-02-04 1985-02-05 1985-02-04 GWC G. Williams South -33.6 Hat Head 19 150.33333 Hat Hill 7 208 2 New South Wales 2448759224 1998-12 ANIC G. Hangay -33.683334 Katandra Bushland Sanct. 19 151.26666 Ingleside 7 208 1 New South Wales 2448759225 1979-12-26 GWC G. & T. Williams -33.7 Springwood 19 150.53334 Sassafras Gully 7 208 2 New South Wales 2448757224 1995-12 80-06-07 1995-12 AM G. O'Reilly -33.75 West Pennant Hills 19 151.03334 7 208 1 New South Wales 2448759232 [252,1333,872,899] 1991-07-07 ANIC T. Gush -33.766666 West Killara 19 151.11667 7 208 2 New South Wales 2448757223 1967-12-15 SAMA Matthews -35.666668 5 mi. N Batemans Bay 19 150.25 7 208 2 New South Wales 2448759230 1978-03-06 CMNC Peck -35.666668 5 mi. N Batemans Bay 19 150.25 7 208 1 New South Wales 2448758232 1974-12-22 CMNC H. F. & A. T. Howden -34.775833 Gerroa 19 150.79695 7 208 1 New South Wales 2448759229 1999-03-12 87-06-07 1999-03-12 ANIC L. Wilkie & R. Harris & Smith -36.28 Princes Highway Corunna State Forest 19 150.12584 7 208 50 New South Wales 2448759226 1968-01-13 SAMA E. Matthews -37.5 Mallacoota National Park 19 149.76666 7 208 1 Victoria 2448756227 1978-05-21 1978-05-26 1978-05-21 ANIC S. & J. Peck -37.5 Mallacoota National Park 19 149.76666 7 208 1 Victoria 2448756226 1978-05-26 ANIC S. & J. Peck -37.5 Mallacoota National Park 19 149.76666 7 208 1 Victoria 2448756225 1993-02-10 1993-02-23 1993-02-10 ANIC A. Newton & M. Thayer 20 -37.516666 South West Arm Road 19 149.68333 Croajingolong National Park 7 208 3 Victoria 2448759227 1978-05-22 1978-05-26 1978-05-22 CMNC S. & J. Peck 3 -37.52972 Double Creek 19 149.68695 Mallacoola National Park 7 208 1 Victoria