Matthews, 1974: 109
Revision of Australian species of the dung beetle genus Lepanus (Coleoptera: Scarabaeidae: Scarabaeinae): description of 10 new species from the L. australis, L. monteithi, and L. villosus species groups
Gunter, Nicole L.
Weir, Thomas A.
Zootaxa
2019
2019-11-08
4695
3
201
252
6P7WD
Matthews, 1974
Matthews
1974
[151,572,1801,1828]
Insecta
Scarabaeidae
Lepanus
Animalia
Coleoptera
3
204
Arthropoda
species
australis
( Figs. 1C–D, F, K; 2A; 3A; 4A)
Lepanus australis Matthews, 1974: 109.
Type material examined. Holotype: ♂“ 5 mi. N. Batemans BayN.S.W. 16-VIII-67 E. Matthews” / “ Lepanus australis Det 1972M., E.G. Matthews HOLOTYPE” / “ ANICDatabase No. 25 057863” / “ HOLOTYPE”, in ANIC[ 35°40’0”S, 150°15”0”E]. Paratypes: Macquarie Pass, [ 34°34’0”S, 150°38’0”E], 29.xii.1967, E. Matthews(25-057864 1 specimen ♂, 4 specimenssex undetermined in ANIC); Clyde Mountain(general), [ 35°33’0”S, 149°57’0”E], 9.i.1968– 12.i.1968, E. Matthews(25-057865 1 specimensex undetermined in ANIC); 3–7 milesN of Eden, [ 37°0’0”S, 149°55’0”E], 13.i.1968, E. Matthews(25-057866 2 specimen ♂, 2 specimensex undetermined in ANIC); 5–17 milesS of Eden, [ 37°14’0”S, 149°55’0”E], 13.i.1968, E. Matthews(25-057867 1 specimensex undetermined in ANIC); Mallacoota National Park, [ 37°30’0”S, 149°46’0”E], 13.i.1968, E. Matthews(25-057868 1 specimensex undetermined in ANIC). Other material examined.Almost 600 other specimens were examined from around 150 localities. Queensland: St Lawrence, 18.5 kmW, 22°21’4”S, 149°20’33”E, 29.vi–22.x.2000, (T107480–T 107482 in QM); Fairlie’s Knob, 0.5 kmS, 25°30’32”S, 152°17’22”E, 20.x–20.xii.2000, (T 107490 in QM); Fairlie’s Knob, 0.5 kmS, 20.xii.2000– 23.iii.2001, (T107491, T107493–T 107495 in QM); Fairlie’s Knob, 0.5 kmS, 22–23.iii.2001, (T 107500 in QM); Lake Bowaddy[Lake Bowarrady], Fraser Island, [ 25°9’0”S, 153°13’0”E], 3.xii.1975, G. Thompson& J.A. Slater(25-058488 1 specimenin QM); Yidney Scrub, Fraser Island, [ 25°19’0”S, 153°11’0”E], 1.xii– 4.xii.1975, G. Thompson& J.A. Slater(25-058491 2 specimensin QM); Booloumba Creek State Forest RiverTrail, 26°38’50”S, 152°38’45”E, 350m, 15.i.2000, R.A. Anderson(CMNEN-00030001 in CMNC); Tinana Creek, 25°45’7”S, 152°41’38”E, 35 m, 22.xi–7.xii.2011, G.B. Monteith(25-0053884 1 specimenalso as COL 2345 in ANIC); Tallegalla Weir Junction, 25°45’25”S, 152°41’49”E, 35m, 6–7.xii.2011, G.B. Monteith(25-053887 1 specimenin ANIC); Nangur State Forest, 2nd site, 26°7’55”S, 151°58’43”E, 24.xi.1995– 3.ii.1996, (T 107570 in QM); Jack Smith[Scrub Conservation] Park on Boat Mountain, 26°9’17”S, 151°54’34”E, 26.i–20.iv.1995, (T107462 – T107464, T107533–T 107552 in QM); Jack Smith[Scrub Conservation] Park on Boat Mountain, 26°9’17”S, 151°54’34”E, 15.xii.1994– 26.i.1995, (T 107553 in QM); Kin Kin Creek, lower, 26°14’15”S, 152°58’52”E, 18– 19.ii.2006, (T 135125 in QM); Elanda Point, National Park Office, 26°15’28”S, 152°59’44”E, 17.ii.2006, (T135122– T 135124 in QM); Cooran Plateauvia Gympie, [ 26°21’0”S, 152°47’0”E], 1300 m, 10–11.iii.1973, R.I. Storey(25- 058486 x 12in QM, 25-058581, 2 specimensin QDPC); Cooran Plateau, 1300 feetvia Gympie, [ 26°21’0”S, 152°47’0”E], 396 m, 3.x–3.xi.1973, R.I. Storey(CMNEN-00030019 in CMNC); Kingaroy, 6 kmSE, 26°34’48”S, 151°52’35”E, 23–26.xii.2005,(T132946–T 132949in QM); Mapleton Falls National Park, 26°37’49”S, 152°50’21”E, 30.xi.1991- 8.i.1992, (T 107573 in QM); 4 kmW Buderim, [ 26°41’0”S, 153°0’0”E], 6.ix.1978, I.F.B. Common(25- 059200 x 1in ANIC); Chevallum, via Nambour, 26°42’3”S, 153°0’33”E, 12.v–27.ix.1978, (T 107594 in QM); Conondale Range, Site1, 26°42’46”S, 152°37’8”E, 20.iv-2.v.1995, (T107567–T 107568 in QM); Mary Cairncross Park, 26°46’52”S, 152°52’56”E, 29.xi.1991- 8.i.1992, (T 107574 in QM); Landsborough, 2.55 kmNE, 26°47’22”S, 152°58’41”E, 28.viii.2010, G.B. Monteith(25- 060783 1 specimenalso as COL 773 in ANIC); Saddletree Creek, 26°51’50”S, 151°38’2”E, 29–30.xii.2010, G.B. Monteith& K. Will(25- 060832 x2also as COL1464-COL 1465 in ANIC); Saddletree Creek, 26°51’50”S, 151°38’2”E, 22–29.xii.2010, Sota& Takami, (25- 060834 x10also as COL1467- COL 1468 in ANIC); Saddletree Creek, 26°51’50”S, 151°38’2”E, 22–30.xii.2010, G.B. Monteith& K. Will(25- 060835 7 specimensalso as COL 1469–1470in ANIC); 5 kmE Yarraman, [ 26°52’0”S, 152°3’0”E], 19.vi.1982, L. Hill, (25-059203 1 specimenin ANIC); Top of Blackbutt Range, 26°52’38”S, 152°11’32”E, 24.xi.1995- 3.ii.1996, (T107517–T 107532 in QM); Bunya Mountains, Dandabah, 26°52’47”S, 151°35’44”E, 20.ix.2007, (T 151105in QM); 3 kmN Upper Yarraman State Forest, 26°52’47”S, 151°53’4”E, 30.iii.1983, (T107571–T 107572 in QM); Bunya Mountain National Park, Westcott Campground, 26°51’51”S, 151°34’17”E, 1100 m, 18.i.2000, R.A. Anderson(CMNEN-00030002, CMNEN-00030004 in CMNC); Bunya Mountain National Park, Dandabah Picnic Area, 26°52’54”S, 151°35’23”E, 1100 m, 19.i.2000, R.A. Anderson(CMNEN-00030003 in CMNC); BlackbuttRange, 1 kmE Benarkin, [ 26°53’0”S, 152°10’0”E], 19.vi.1982, L. Hill, (25-059202 1 specimenin ANIC); Upper Yarraman State Forestvia Maidenwell, [ 26°54’0”S, 151°54’0”E], 11.i.1975, G.B. Monteith(25- 058493 3 specimensin QM); Wild Horse Mountain, 1.2 kmSW, 26°55’26”S, 152°59’24”E, 15–16.xi.2007, (T 151108in QM); Wild Horse Mountain, 1.2 kmSW, 26°55’26”S, 152°59’24”E, 23.i–13.ii.2008, (T154607– T 154614 in QM); Bribie Island, 1.3 kmNE of national park headquarters, 27°0’54”S, 153°7’30”E, 10 m, 8–9.x.2010, G.B. Monteith(25- 060876 x1also as COL 1141in ANIC); Bribie Island, 0.85 kmNE of national park headquarters, 27°1’5”S, 153°7’23”E, 8–9.x.2010, G.B. Monteith(25- 060877 1 specimenalso as COL 1142 in ANIC); Brisbane, Forest Park, Scrub CreekRoad, [ 27°1’0”S, 152°26’0”E], 18.ix–29.x.1998, P. Bouchard(25-058494 1 specimenin QM); Brisbane ForestPark, 27°25’41”S, 152°50’18”E, 1–15.x.2002, J. Skevington& J. Cumming(CMNEN- 00030008 in CMNC); Blackbutt Range( Bottom), 8 milesE Blackbutt, [ 27°3’0”S, 151°55’0”E], 22–23.xii.1973, R.I. Storey(25-058580 2 specimensin QDPC); Mount Deongwar, 3 kmS, Site3, 27°13’41”S, 152°15’36”E, 30.xii.1998– 26.iii.1999, (T107285–T 107292 in QM); Mount Deongwar, 3 kmS, Site3, 27°13’41”S, 152°15’36”E, 14.x–30.xii.1998, (T 107391 in QM); Mount Deongwar, 3 kmS, Site2, 27°13’49”S, 152°15’25”E, 30.xii.1998– 26.iii.1999, (T107282–T 107284 in QM); Mount Deongwar, 3 kmS, Site2, 27°13’49”S, 152°15’25”E, 14.x– 31.xii.1998, (T 107390 in QM); Mount Glorious, barracks, 27°19’54”S, 152°45’12”E, 16–28.iv.2007, (T 148366 in QM); Mount Glorious, barracks, 27°19’54”S, 152°45’12”E, 12–15.iii.2008, (T 156700 in QM); Crows Nest National Park, Perseverance Section, Site1, 27°19’8”S, 152°6’’27”E, 4–5.xii.2003, (T 120685–T 120687in QM); Mount Glorious, [ 27°20’0”S, 152°46’0”E], 630 m, 14.xi.1986– 30.i.1987, T. Hiller, (25-059197 3 specimens, 25- 059198, 2 specimensin ANIC); Mount Glorious, [ 27°20’0”S, 152°46’0”E], 635 m, ii.1989, H.F. & A.T. Howden(CMNEN-00030007 in CMNC); Stony Creek, via Samford, 27°20’20”S, 152°47’52”E, 22.x.1994- 2.ii.1995, (T107554–T 107558 in QM); Stony Creek, via Samford, 27°20’20”S, 152°47’52”E, 2.ii–iv.1995, (T107559– T 107566 in QM); Mount Glorious, 2 kmS, 27°21’9”S, 152°46’7”E, 28–29.i.2006, (T133758–T 133761 in QM); Boombana National Park, Site1, 27°24’8”S, 152°47’22”E, 1.x.2003– 30.x.2003, (T 122130in QM); Boombana National Park, Site1, 27°24’8”S, 152°47’22”E, 15–16.ii.2004, (T 122132in QM); Boombana National Park, Site1, 27°24’8”S, 152°47’22”E, 2.i.2004– 30.i.2004, (T 122794–T 122795in QM); Boombana National Park, 27°25’55”S, 152°53’49”E, 230 m, 18.ii.2012– 19.ii.2012, G.B. Monteith(25-053911 x 1in ANIC); McAfee’s Lookout, 0.5 kmSW, 27°25’58”S, 152°52’14”E, 18.x.1999- 6.ii.2000, (T 107475 in QM); McAfee’s Lookout, 1 kmSW, 27°26’15”S, 152°52’14”E, 18.x.1999– 6.ii.2000, (T 107476 in QM); Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E, 21.xii.1999– 27.i.2000, (T107471–T 107473 in QM); Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E, 27.i– 15.iii.2000, (T 107474 in QM); Enoggera Reservoir, Site8, 27°26’37”S, 152°54’14”E, 15.iii–18.v.2000, (T107478– T 107479 in QM); Enoggera Reservoir, Site5, 27°26’44”S, 152°55’19”E, 16.x–21.xii.1999, (T107467–T 107469 in QM); Enoggera Reservoir, site 5, 27°26’44”S, 152°55’19”E, 21.xii.1999– 27.i.2000, (T 107470 in QM); Enoggera Reservoir, site 3, 27°26’44”S, 152°55’19”E, 15.iii–18.v.2000, (T 107477 in QM); McAfee’s Lookout, 0.7 kmSW, 27°26’8”S, 152°52’17”E, 6.vii–16.x.1999, (T107393–T 107394 in QM); Gold Creek Reservoir, Site4, 27°27’43”S, 152°52’21”E, 12–14.x.2007, (T 151106in QM); Gold Creek Reservoir, 27°27’46”S, 152°52’54”E, 12–14.x.2007, (T 151107in QM); Slaughter Falls, 27°28’44”S, 152°57’34”E, 6–7.ii.2001, (T 107483 in QM); Boscombe Road, Brookfield, 27°28’57”S, 152°55’1”E, 12–13.ii.2011, G.B. Monteith(25-053877 1 specimenalso as COL 1678 in ANIC); Gold Creek Reservoir, Site2, 27°28’5’’S, 152°52’20”E, 4.xi.2003, (T 122131in QM); Upper Brookfield Road, Brisbane, [ 27°29’0”S, 152°52’0”E], 24.viii–7.ix.1975, R.I. Storey(25-058489 1 specimenin QM); Upper Brookfield Road, Brisbane, [ 27°29’0”S, 152°52’0”E], 22.ix–4.x.1975, R.I. Storey(25-058490 5 specimensin QM); Kenmore Kookaburra Street, 27°29’28”S, 152°55’23”E, 16–17.xi.2010, <collector unknown>, (25- 060906 1 specimenalso as COL 1197 in ANIC); Kenmore Hills, 27°29’28”S, 152°55’24”E, 20–21.i.2011, G.B. Monteith(25- 065490 2 specimensin ANIC); Kenmore, 27°30’24”S, 152°55’52”E, 20–21.i.2011, G.B. Monteith(25-065489 2 specimensin ANIC); Mabb Street, Kenmore, 27°30’25”S, 152°55’52”E, 16–17.xi.2010, G.B. Monteith(25- 060889 1 specimenalso as COL 1196 in ANIC); The Knobby, 4 kmW Glamorgan Vale, 27°30’27”S, 152°35’18”E, 13.i– 16.v.1999, (T 107279 inQM–T 107281 in QM); Cameron’s Scrub, top ridge, 27°30’29”S, 152°43’54”E, 13.i– 16.v.1999, (T107303–T 107310 in QM); Cameron’s Scrub, 27°30’32”S, 152°43’48”E, 75 m, 30.iv–1.v.2012, G.B. Monteith(25-057574 2 specimens, 25-075575 5 specimensin ANIC); Long Pocket, river bank, 27°30’34”S, 152°59’53”E, 1.xi–4.xi.1999, (T 107395 in QM); Cameron’s Scrub, gully, 27°30’35”S, 152°43’54”E, 13.i–16.v.1999, (T107311–T 107347 in QM); Cameron’s Scrub, gully, 27°30’35”S, 152°43’54”E, 11.xi.1998– 13.i.1999, (T107349– T 107369 in QM); Cameron’s Scrub, below road, 27°30’45”S, 152°43’39”E, 13.i–16.v.1999, (T107293–T 107295 in QM); Belmont Hill, top, 27°30’48”S, 153°6’’59”E, 12.xii.1997– 7.v.1998, (T107417–T 107433 in QM); Cameron’s Scrub, Snigtrack, 27°30’49”S, 152°43’43”E, 13.i–16.v.1999, (T107296–T 107302 in QM); Cameron’s Scrub, Snigtrack, 27°30’49”S, 152°43’43”E, 11.xi.1998– 13.i.1999, (T 107389 in QM); Whites Hill Reserve, Carnelian Street, 27°30’58”S, 153°4’41”E, 20–22.iv.2001, (T 107499 in QM); Pullenvale State Forest, 27°30’59”S, 152°51’44”E, 9–11.iv.2001, (T107465–T 107466 in QM); Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E, 28.vii–1.ix.2003, (T 120590–T 120591in QM); Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E, 27.v–30.vi.2003, (T 120592in QM); Bulimba Creek, Carindale, 27°30’9”S, 153°6’’34”E, 1.xii.2003- 2.i.2004, (T 122796in QM); Sankey’s Scrub, Camp Hill, 27°31’1”S, 153°5’6”E, 20–22.iv.2001, (T 107497 in QM); Rainbow Forest, Cubberla Creek, 27°31’6”S, 152°57’40”E, 24.xi–21.xii.2005, (T132653–T 132654 in QM); Rainbow Forest, Cubberla Creek, 27°31’6”S, 152°57’40”E, 24–25.xi.2005, (T132655–T 132657 in QM); Perry’s Knob, 4 kmNNE Rosewood, 27°36’11”S, 152°36’17”E, 13.i.1999– 16.v.1999, (T107252–T 107278 in QM); Perry’s Knob, 4 kmNNE Rosewood, 27°36’11”S, 152°36’17”E, 11.xi.1998– 13.i.1999, (T107348, T107370–T 107388 in QM); Mount Cotton, Scott’s Dam, 27°36’58”S, 153°12’21”E, 12.xii.1997– 7.v.1998, (T107396–T 107402 in QM); Mount Cotton, upper gully, 27°36’58”S, 153°12’32”E, 12.xii.1997– 7.v.1998, (T107403–T 107416 in QM); Mount Cotton, 27°36’58”S, 153°12’21”E, 3.ix–12.xii.1997, (T107450–T107461, T 107496 in QM); Burtonsfirebreak RF (B6), 27°38’6”S, 152°1’41”E, 410m, 2–20.xi.2015, G.B. Monteith& K. Ebert, (25-065488 6 specimensin ANIC); Mount Tamborine National Park, [ 27°53’11”S, 153°10’51”E], 700 m, 3.iii.1984, L. Masner(CMNEN-00030000 in CMNC); Cunninghams Gap, 28°3’0”S, 152°24’0”E], 1–2.iii.1973, R.I. Storey(25-058487 1 specimenin QM, 25-058582 1 specimenin QDPC); Moss’ Wellat Spicer’s Gap, 28°4’28”S, 152°25’23”E, 20.ii–3.iv.1994, (T 107569 in QM); Black Rock Scrub, 28°7’32”S, 152°39’48”E, 6.x–3.xii.2000, (T107484–T 107489 in QM); Black Rock Scrub, 28°7’32”S, 152°39’48”E, 2.xii.2000– 13.v.2001, (T107503–T107510, T107513, T107515–T 107516 in QM); Lamington National Park, IBISCA SiteIQ–700-OF, 28°8’2.4”S, 153°6’’46.8”E,– 30.iii.2007, (T108395–T 108396 in QM); Lamington National Park, IBISCA SiteIQ–300-D, 28°8’31.2”S, 153°7’58.8”E, 18–23.i.2007, (T 145450 in QM); Lamington National Park, IBISCA SiteIQ–300-B, 28°9’18”S, 153°8’20.4”E, 16–21.x.2006, (T145447– T 145449 in QM); Binna Burra, below, 28°11’20”S, 153°11’10”E, 610 m, 29.ii–1.iii.2012, G.B. Monteith(25- 053908 1 specimenalso as COL 2391 in ANIC); Teviot Falls, via Boonah, 28°13’25”S, 152°28’45”E, 5.x.1975– 22.ii.1976, (T107592–T 107593 in QM); Springbrook, Mount Wunburra, 28°14’24”S, 153°16’3”E, 1–2.iv.2006, (T 135121 in QM); Mount French, via Boonah, 28°18’57”S, 152°50’53”E, 5.x.1975– 22.ii.1976, (T107575–T 107591 in QM); New South Wales: Unumgar State Forestnear Woodenbong, Pole Bridgeroad 788, [ 28°24’0”S, 152°40’0”E], 430 m, 2–11.i.1987, A. Newton& M. Thayer(25-059195 3 specimens, 25-059196 1 specimenin ANIC); Mount Lindsay Hwy, 13 mi.S of Rathdowney, [ 28°24’0”S, 152°52’0”E], 24–25.ii.1973, R.I. Storey(25- 058492 1 specimenin QM); Border Ranges National Park 1 kmSE Richmond Gap, [ 28°27’0”S, 152°53’0”E], 350 m, 13.iv.1993, D.S. Chandler(25-059190 1 specimenin ANIC); Wilson Park, 3 kmSE of Lismore, [ 28°49’0”S, 153°16’0”E], 50 m, 25.viii.1982, S. & J. Peck(25-059188 2 specimensin ANIC); Iluka Nature Reserve, [ 29°24’0”S, 153°21’0”E], 20–24.xi.1998, G. Williams(25-058088 1 specimenin GWC); Kangaroo River State Forestalong Burns Roadfrom Lowana, 30°4’30”S, 152°52’5”E, 400m, 21.iii–8.v.1999, S. Lassau& C. Lemann(25- 060779 1 specimenalso as COL 760 in AM); vicinity of Toormina Park, Toormina, [ 30°22’0”S 153°6’0”E], 2.xii.1997, G. Williams(25-058095 1 specimenin GWC); Scotchman State Forest, Grahamstrail off Horseshoe Road, 30°25’23”S, 152°49’45”E, 23–26.ix.2011, N. Gunter(25-057582 1 specimenalso as COL2006, 25-057583 1 specimenalso as COL 2007 in ANIC); Newry State Forest, Hungry Head Road, 30°31’41”S, 153°0’22”E, 23–26.ix.2011, N. Gunter(25-057580 5 specimensalso as COL2004, 25-057581 1 specimenalso as COL 2005 in ANIC); Walcha Survey Big Hill, 30°37’46”S, 152°14’6”E, i.1993, I. Oliver, (25- 060781 2 specimensin AM); Racecourse Headland S of Crescent Head[Racecourse Head], [ 31°15’0”S, 152°58’0”E], 20.i.1999, G. & T. Williams(25-058089 1 specimenin GWC); Point Plomer, Limeburners Creek National Park N of Port Macquarie, [ 31°19’0”S, 152°58’0”E], 23– 24.x.2010, G. & T. Williams(25- 060888 1 specimenalso as COL 1195 in ANIC); Winmurra, approximately 50 kmNW Taree, Doyles River, [ 31°34’0”S, 152°6’0”E], 530 m, 2002, 2008, D. Bray(25-053894 1 specimenin GWC); N end of Easternfire trail, Yarratt State Forest, NW of Wingham, [ 31°44’0”S, 152°26’0”E], 8.x.1981, G. & T. Williams(25-057875 1 specimenin ANIC, 25-058090 2 specimensin GWC); Lorien Wildlife Refuge 3 kmN Lansdowne[ 31°45’0”S, 152°32’0”E], 17.x.2010, G. & T. Williams(25- 060887 1 specimenalso as COL 1194 in ANIC); 4 kmW Lansdowne, Lorein, 152°32’0”E, 31°45’0”S, 12.v–12.vi.1986, H.F. & A.T. Howden, (CMNEN-00030009- 00030010 in CMNC); 4kmW. Lansdowne, Lorien, 152°32’0”E, 31°45’0”S, 1.iv.1987, H.F. & A.T. Howden, (CMNEN-00030011-00030018 in CMNC); Newbys Lane, Lansdowne, [ 31°47’0”S, 152°32’0”E], 12–14.iv.1981, G. & T. Williams(25-057874 1 specimenin ANIC); Lorien, Lansdowne, [ 31°52’0”S, 152°29’0”E], 10–12.iv.1981, G. & T. Williams(25-058097 2 specimensin GWC); Lansdowne Recreational ReserveSE Lansdownevia Taree, [ 31°47’0”S, 152°32’0”E], 24–25.x.1984, G. Williams(25-058092 1 specimenin GWC); approximately 24 kmNNW Gloucester, [ 31°49’0”S 151°52’0”E], 19–20.xii.1991, G. & T. Williams(25-058094 1 specimenin GWC); 3.5 kmNE of Harrington, [ 31°50’0”S, 152°43’0”E], 18.viii.1990, G. & T. Williams(25- 060778 1 specimenalso as COL 759 in AM); Crowdyhead approximately 7 kmN of Harrington, [ 31°51’0”S, 152°45’0”E], 28.ix.1982, G. & T. Williams(25-058100 2 specimensin GWC); Harrington, [ 31°52’0”S, 152°42’0”E], 14.xi.1976, G. WilliamsSAMA25-027647 2 specimensin SAM); Harrington, [ 31°52’0”S, 152°42’0”E], 25.xii.1976, G. Williams(25- 058104 1 specimenin GWC); Harrington, [ 31°52’0”S, 152°42’0”E], 20.xii.1976, G. Williams(25-059193 1 specimenin ANIC); Harrington, [ 31°52’0”S, 152°42’0”E], 10.x.1976, G. Williams(25-059194 1 specimenin ANIC); Kiwarrak State Forest, [ 31°57’0”S, 152°27’0”E], 25.v.1992, T. Gush(25-059191 1 specimenin ANIC); Kiwarrak State Forestnear Taree, [ 31°58’0”S, 152°26’0”E], 19.ix.1979, W. King(25- 060786 1 specimenalso as COL777 and K 266951 in AM); Kiwarrak State Forest, [ 31°58’0”S, 152°28’0”E], 29.v–1.vi.1993, V. Lorimer(25-058712 2 spec- imens in ANIC); Kiwarrak State Forest, via Taree, approximately 1 kmN of Breakneck Lookout, [ 31°59’0”S, 152°25’0”E], 17.ii.1982, G. & T. Williams(25-057873 1 specimenin ANIC, 25-058102 2 specimensin GWC); Saltwatervicinity of Old Bar, [ 32°0’0”S, 152°34’0”E], 22.x.1981, G. & T. Williams(25-058096 1 specimenin GWC); Black Head( Hallidays Point), [ 32°4’0”S, 152°33’0”E], 13.x.1981, G. & T. Williams(25-058101 2 specimensin GWC); Gloucester River, Barrington Tops National Park, [ 32°4’0”S, 151°41’0”E], 12–14.xi.1981, T.A. Weir(25-059189 x 1in ANIC); on Mountain Road 0.2 kmoff junction with Kunungra Road, 32°8’22”S, 151°44’4”E, 4.ii–9.iv.1993, G. Cassis& M. Gray(25- 060775 1 specimenalso as COL756 and K 266946 in AM); Southbase of Cape Hawkevia Forster, [ 32°13’0”S, 152°34’0”E], 30.ix.1981, G. & T. Williams(25-058099 1 specimenin GWC); O’Sullivan Gap Reserve, 11 kmNE Bulahdelah, [ 32°20’0”S, 152°17’0”E], 50 m, 11.vi–27.viii.1982, S. & J. Peck(25-059187 2 specimensin ANIC); Bombah Point Swamp[ 32°30’0”S, 152°17’0”E], 1.v.1993, V. Lorimer(25- 058713 1 specimenin ANIC); Mungo Brush, Myall Lakes National Park, [ 32°32’0”S, 152°19’0”E], 26.x.1981, G. & T. Williams(25-057872 2 specimensin ANIC, 25-058103 2 specimensin GWC); Mungo Brush, [ 32°36’0”S, 152°16’0”E], 1.v.1993, V. Lorimer(25-058711 2 specimensin ANIC); Mount Coricudgynear Olinda Station, [ 32°50’0”S, 150°21’0”E], 5–9.iii.2002, G. Hangay(25-057878 2 specimens, 25-057879 1 specimenin ANIC); Gowan, [ 33°12’0”S, 149°22’0”E], 25.vii.1976, B.B. Lowery(25-057871 4 specimensin ANIC); Mount Irvine, [ 33°29’0”S, 150°28’0”E], 5.i.1980, G. & T. Williams(25-058098 1 specimenin GWC); Southslope, Hat Hill, Hat Head, [ 33°36’0”S, 150°20’0”E], 4–5.ii.1985, G. Williams(25-058091 2 specimensin GWC); Ingleside, Katandra Bushland Sanct., [ 33°41’0”S, 151°16’0”E], xii.1998, G. Hangay(25-059192 1 specimenin ANIC); Sassafras Gullyvia Springwood, [ 33°42’0”S, 150°32’0”E], 26.xii.1979, G. & T. Williams(25-058093 2 specimensin GWC); West Pennant Hills, [ 33°45’0”S, 151°2’0”E], xii.1995, G. O’Reilly(25- 060780 1 specimenalso as COL761 and K 266947 in AM); West Killara, [ 33°46’0”S, 151°7’0”E], 7.vii.1991, T. Gush(25-057880 2 specimensin ANIC); 5 mi.N Batemans Bay, [ 35°40’0”S, 150°15’0”E], 15.xii.1967, E. Matthews(25-027645 1 specimen, 25-027646 1 specimenin SAMA); 5 mi.N Batemans Bay, [ 35°40’0”S, 150°15’0”E], 6.iii.1978, S. & J. Peck(CMNEN-00030005 in CMNC); Gerroa, [ 34°46’33”S, 150°47’49”E], 22.xii.1974, H.F. & A.T. Howden(CMNEN-00030020-00030024 in CMNC); Princes Highway Corunna State Forest, 36°16’48”S, 150°7’33”E, 12.iii.1999, L. Wilkie, R. Harris, & H. Smith(25- 060787 50 specimensand K266949 also as COL778-COL 780 in ANIC); Victoria: Mallacoota National Park, [ 37°30’0”S, 149°46’0”E], 13.i.1968, E. Matthews(25-027649 1 specimenin SAMA); Mallacoota National Park, [ 37°30’0”S, 149°46’0”E], 21–26.v.1978, S. & J. Peck(25-057869 1 specimenin ANIC); Mallacoota National Park, [ 37°30’0”S, 149°46’0”E], 26.v.1978, S. & J. Peck(25-057870 1 specimenin ANIC); Croajingolong National Park, South West Arm Road, 11 kmNW Mallacoota, [ 37°31’0”S, 149°41’0”E], 20 m, 10–23.ii.1993, A. Newton& M. Thayer(25-057876 2 specimens, 25-057877 1 specimenin ANIC); Mallacoola National Park, Double Creek, [ 37°31’47”S, 149°41’13”E], 3 m, 22.v.1978– 26.v.1978, S. & J. Peck(CMNEN-00030006 in CMNC).
Description. Holotypeuniformly dark brown almost black in colour. Rest of specimens range from light brown to black. Antennal clubs grey. Total length: 2.7–3.5 mm. Holotypemeasurements: Total length 3.1 mm, elytral width 2.0 mm. Male. Head: Surface with fine punctation. Basal carina absent. Dorsal part of the eyes narrow, separated by an interocular space approximately 16 times the eye width (40: 2.5); eye canthus dividing eye. Head 1.4 x wider than long (49:35). Prothorax: Pronotal surface with very fine punctation anteriorly to fine punctation posteriorly. Anterior angles sharp (less than 90°). Basal margin slightly angulate. Hypomeral surface reticulate. Hypomeral striae over half the length of hypomeron and very close to lateral margin, almost appearing double. Pronotum 1.7 x wider than long (40:24). Elytra: Surface smooth with impressed, impunctate striae, striae 1–3 or 1–4 appearing double, 5–7 appearing single; intervals very finely punctate. Stria 6 extending to elytral base and 7 not extending to elytral base. Striae 8 slightly curving inwards posteriorly. Elytra almost equal in width and length (48:45). Legs: Front edge of protibiae deeply angulate. Abdomen: Pygidium simple, smooth, finely punctured and slightly convex. Abdominal ventrites reticulate right across. Ventrite 6 very finely punctate. Pterothorax: Medial lobe of metaventrite punctate and broadly margined between the mesocoxae. Lateral lobe of metaventrite punctate, inner border next to mesocoxae anteriorly straight and lateral border virtually absent. Mesoventrite notched anteriorly. Mesoventrite smooth and nitid with very fine punctation. Mesepimeron very finely reticulate. Metanepisternum reticulate. FIGURE 1. Some diagnostic characters of the Lepanus australisspecies group. A, Male protibia of Lepanus eungelladotted line depicts triangular apical digit (grey arrow) and truncate front edge; B, female protibia of Lepanus eungelladotted line depicts truncate front edge, black arrow points to apical spur; C, male protibia of Lepanus australisdotted line depicts elongate apical digit (grey arrow) and deeply angulate front edge; D, female protibia of Lepanus australisdotted line depicts shortened apical digit (grey arrow) and deeply angulate front edge, black arrow points to apical spur; E, lateral view of Lepanus dukungarriwhite arrow points to row of punctures on metanepisternum; F, lateral view of Lepanus australis, dotted line indicates anteriorly straight inner border of lateral lobe of metaventrite, note the absence of puntures on metanepisternum and straight lower edge of elytral epipleura; G, lateral view of Lepanus oxleyiwhite arrow points to row of fine punctures on metanepisternum, grey arrow inducates curved lower edge of elytral epipleura; H, male metafemora of Lepanus dukungarrishowing medial expansion; I, Head of Lepanus oxleyiwhite arrow points to interrupted basal carina; J, lateral view of Lepanus latheticusdotted line indicates anteriorly curved inner border of lateral lobe of metaventrite; K, Lepanus australisat bird droppings (image: Geoff Williams). Female. As per L. australisspecies group.
Distribution. Lepanus australishas the one of the broadest distributions of the genus ( Fig. 5A) and has been recorded in eastern Australiafrom as far north as St Lawrence, Queenslandto Mallacoota, Victoria( 2000 kmsouth). It is known from a variety of habitats (rainforest, palm forest, wet and dry sclerophyll forests) in the following Interim Biogeographic Regionalisation for Australia(IBRA) bioregions: South Eastern Queensland(SEQ), NSW North Coast (NNC), New EnglandTablelands (NET), Sydney Basin (SYB), South East Corner (SEC), South Eastern Highlands (SEH), the most easterly extents of the Brigalow Belt North (BBN), and Brigalow Belt South (BBS). BIOCLIM modelling suggests appropriate climatic conditions extend further inland than currently recorded, however despite extensive sampling in some of the adjacent IBRA regions, L. australishas not been collected.
Comments.Most specimens have been collected in baited pitfall traps (mushroom, dung, or carrion), but it has also been collected in berlesed litter, at faeces, and in flight intercept traps. Molecular sequences have been generated from numerous locations across the range. The phylogeny of Gunter & Weir (2017)includes representatives from three Queenslandpopulations: Kenmore, Brisbane (25- 060906also as COL1197), Saddletree Creek, Bunya Mountains (25- 060835also as COL1470), and Tinana Creek near Maryborough (25-0053884 also as COL2345) but other specimens from New South have also been sequenced and confirm the wide distribution of the species. The specimen from Kenmore was included in the phylogeny of Gunter et al.(2019) and it formed a relationship with an undescribed species in the L. pisoniaespecies group informally referred to as Lepanusspecies NQ19. Lepanus australisis one of several species of Lepanusthat have been observed at bird droppings ( Fig. 1K), previously thought to be a food source underutilised by dung beetles.
2448758245
1967-08-16
ANIC
N. Batemans Bay & E. Matthews
Batemans Bay
3
204
1
1
holotype
2448757220
1967-12-29
ANIC
E. Matthews
-34.566666
Macquarie Pass
19
150.63333
4
205
5
1
paratype
2448759213
1968-01-09
1968-01-12
1968-01-09
ANIC
E. Matthews
-35.55
Clyde Mountain
19
149.95
4
205
1
paratype
2448756216
1968-01-13
ANIC
E. Matthews
-37.0
Eden
19
149.91667
4
205
4
2
paratype
2448756223
1968-01-13
ANIC
E. Matthews
-37.233334
Eden
19
149.91667
4
205
1
paratype
2448756221
1968-01-13
ANIC
E. Matthews
-37.5
Mallacoota National Park
19
149.76666
4
205
1
paratype
2448758228
2000-06-29
2000-10-22
2000-06-29
QM
-22.351112
St Lawrence
21
149.3425
4
205
1
Queensland
2448757221
[220,1217,476,503]
2000-10-20
2000-12-20
2000-10-20
QM
-25.508888
Fairlie's Knob
21
152.28944
4
205
1
Queensland
2448756217
2000-12-20
2001-03-23
2000-12-20
QM
Fairlie's Knob
4
205
1
Queensland
2448759212
2001-03-22
2001-03-23
2001-03-22
QM
Fairlie's Knob
4
205
1
Queensland
2448757217
1975-12-03
QM
G. Thompson & J. A. Slater
-25.15
Fraser Island
21
153.21666
Lake Bowaddy
4
205
1
Queensland
2448759216
1975-12-01
1975-12-04
1975-12-01
QM
G. Thompson & J. A. Slater
-25.316668
Fraser Island
21
153.18333
Yidney Scrub
4
205
2
Queensland
2448756219
2000-01-15
CMNC
R. A. Anderson
350
-26.647223
Booloumba Creek State Forest River
20
152.64583
4
205
1
Queensland
2448756218
2011-11-22
2011-12-07
2011-11-22
ANIC
G. B. Monteith
35
-25.751944
Tinana Creek
21
152.6939
4
205
1
Queensland
2448759223
2011-12-06
2011-12-07
2011-12-06
ANIC
G. B. Monteith
35
-25.756945
Tallegalla Weir Junction
21
152.69695
4
205
1
Queensland
2448758226
[337,1432,764,791]
1995-11-24
1996-02-03
1995-11-24
QM
-26.131945
Nangur State Forest
20
151.9786
4
205
1
Queensland
2448759222
1995-01-26
1995-04-20
1995-01-26
QM
-26.154722
Park on Boat Mountain
20
151.90944
Jack Smith
4
205
1
Queensland
2448758223
1994-12-15
1995-01-26
1994-12-15
QM
-26.154722
Park on Boat Mountain
20
151.90944
Jack Smith
4
205
1
Queensland
2448758219
2006-02-18
2006-02-19
2006-02-18
QM
-26.237501
Kin Kin Creek
20
152.98111
4
205
1
Queensland
2448758224
2006-02-17
QM
-26.257778
National Park Office
20
152.99556
Elanda Point
4
205
1
Queensland
2448757218
1973-03-10
2012-10
1973-03-10
QM, QDPC
R. I. Storey
1300
-26.35
Gympie
20
152.78334
Cooran Plateau
4
205
2
Queensland
2448758221
1973-10-03
1973-11-03
1973-10-03
CMNC
R. I. Storey
396
-26.35
Gympie
20
152.78334
Cooran Plateau
4
205
1
Queensland
2448758218
2005-12-23
2005-12-26
2005-12-23
QM
-26.580002
Kingaroy
20
151.87639
4
205
1
Queensland
2448758220
1991-11-30
1992-01-08
1991-11-30
QM
-26.630278
Mapleton Falls National Park
20
152.83916
4
205
1
Queensland
2448759219
1978-09-06
ANIC
Common
-26.683332
4 km W Buderim
20
153.0
4
205
1
Queensland
2448759221
[416,1432,1124,1151]
1978-05-12
1978-09-27
1978-05-12
QM
-26.700834
Nambour
20
153.00917
Chevallum
4
205
1
Queensland
2448758227
[151,1236,1160,1187]
1995-04-20
1995-05-02
1995-04-20
QM
-26.712778
Site
20
152.6189
Conondale Range
4
205
1
Queensland
2448758217
1991-11-29
1992-01-08
1991-11-29
QM
-26.78111
Mary Cairncross Park
20
152.88222
4
205
1
Queensland
2448759211
2010-08-28
83-06-07
2010-08-28
ANIC
G. B. Monteith
-26.789444
Landsborough
20
152.97804
4
205
1
Queensland
2448758216
2010-12-29
2010-12-30
2010-12-29
ANIC
G. B. Monteith & K. Will
-26.86389
Saddletree Creek
20
151.63388
4
205
1
Queensland
2448757222
2010-12-22
2010-12-29
2010-12-22
ANIC
Sota & Takami
-26.86389
Saddletree Creek
20
151.63388
4
205
1
Queensland
2448759217
2010-12-22
35-06-08
2010-12-22
ANIC
G. B. Monteith & K. Will
-26.86389
Saddletree Creek
20
151.63388
4
205
7
Queensland
2448757214
1982-06-19
ANIC
Hill
-26.866667
5 km E Yarraman
20
152.05
4
205
1
Queensland
2448759215
1995-11-24
1996-02-03
1995-11-24
QM
-26.877222
Top of Blackbutt Range
20
152.19223
4
205
1
Queensland
2448756220
05-11-15
2007-09-20
05-11-15
QM
-26.879723
Dandabah
20
151.59555
Bunya Mountains
4
205
1
Queensland
2448758222
1983-03-30
QM
-26.879723
Upper Yarraman State Forest
20
151.88445
4
205
1
Queensland
2448757215
2000-01-18
CMNC
R. A. Anderson
1100
-26.864166
Westcott Campground
20
151.57138
Bunya Mountain National Park
4
205
1
Queensland
2448757213
2000-01-19
CMNC
R. A. Anderson
1100
-26.881666
Dandabah Picnic Area
20
151.58972
Bunya Mountain National Park
4
205
1
Queensland
2448756222
1982-06-19
ANIC
L. Hill
-26.883333
Blackbutt
20
152.16667
4
205
1
Queensland
2448757216
1975-01-11
QM
G. B. Monteith
-26.9
Maidenwell
20
151.9
Upper Yarraman State Forest
4
205
3
Queensland
2448759218
08-11-15
2007-11-16
08-11-15
QM
-26.92389
Wild Horse Mountain
20
152.99
4
205
1
Queensland
2448757219
2008-01-23
2008-02-13
2008-01-23
QM
-26.92389
Wild Horse Mountain
20
152.99
4
205
1
Queensland
2448758225
2010-10-08
2010-10-09
2010-10-08
ANIC
G. B. Monteith
10
-27.015
Bribie Island
20
153.125
4
205
1
Queensland
2448759214
2010-10-08
77-06-08
2010-10-08
ANIC
G. B. Monteith
-27.018055
Bribie Island
20
153.12306
4
205
1
Queensland
2448757212
1998-09-18
1998-10-29
1998-09-18
QM
P. Bouchard
Brisbane
-27.016666
Scrub Creek
20
152.43333
Forest Park
4
205
1
Queensland
2448758229
2002-10-01
2002-10-15
2002-10-01
CMNC
J. Skevington & J. Cumming
-27.428055
Brisbane Forest
20
152.83833
4
205
1
Queensland
2448759220
1973-12-22
1973-12-23
1973-12-22
QDPC
R. I. Storey
-27.05
Bottom
20
151.91667
Blackbutt Range
4
205
2
Queensland
2448758215
1998-12-30
1999-03-26
1998-12-30
QM
5
206
-27.228056
Site
20
152.26
Mount Deongwar
4
205
1
Queensland
2448756207
1998-10-14
1998-12-30
1998-10-14
QM
-27.228056
Site
20
152.26
Mount Deongwar
5
206
1
Queensland
2448759205
1998-12-30
1999-03-26
1998-12-30
QM
-27.230278
Site
20
152.25694
Mount Deongwar
5
206
1
Queensland
2448759208
1998-10-14
1998-12-31
1998-10-14
QM
-27.230278
Site
20
152.25694
Mount Deongwar
5
206
1
Queensland
2448758210
2007-04-16
2007-04-28
2007-04-16
QM
-27.331667
Mount Glorious
20
152.75333
5
206
1
Queensland
2448757210
[223,1236,296,323]
2008-03-12
2008-03-15
2008-03-12
QM
-27.331667
Mount Glorious
20
152.75333
5
206
1
Queensland
2448757202
2003-12-04
87-12-06
2003-12-04
QM
Crows Nest National Park
-27.31889
Site
Perseverance Section
5
206
1
Queensland
2448758212
1986-11-14
1987-01-30
1986-11-14
ANIC
T. Hiller
630
-27.333334
Mount Glorious
20
152.76666
5
206
5
Queensland
2448757206
1989-02
CMNC
H. F. & A. T. Howden
635
-27.333334
Mount Glorious
20
152.76666
5
206
1
Queensland
2448756210
1994-10-22
1995-02-02
1994-10-22
QM
-27.33889
Samford
20
152.79779
Stony Creek
5
206
1
Queensland
2448757207
QM
-27.33889
Samford
20
152.79779
Stony Creek
5
206
1
Queensland
2448759210
[363,1432,512,539]
2006-01-28
2006-01-29
2006-01-28
QM
-27.3525
Mount Glorious
20
152.7686
5
206
1
Queensland
2448757209
[151,1300,548,575]
2003-10-01
30-12-21
2003-10-01
QM
-27.402222
Site
20
152.78944
Boombana National Park
5
206
1
Queensland
2448759201
2004-02-15
32-12-21
2004-02-15
QM
-27.402222
Site
20
152.78944
Boombana National Park
5
206
1
Queensland
2448756202
2004-01-02
95-12-27
2004-01-02
QM
-27.402222
Site
20
152.78944
Boombana National Park
5
206
1
Queensland
2448758204
2012-02-18
2012-02-19
2012-02-18
ANIC
G. B. Monteith
230
-27.431944
Boombana National Park
20
152.89694
5
206
1
Queensland
2448759203
1999-10-18
2000-02-06
1999-10-18
QM
-27.432777
McAfee's Lookout
20
152.87056
5
206
1
Queensland
2448758203
1999-10-18
2000-02-06
1999-10-18
QM
-27.4375
McAfee's Lookout
20
152.87056
5
206
1
Queensland
2448758205
1999-12-21
2000-01-27
1999-12-21
QM
-27.443611
Site
20
152.90388
Enoggera Reservoir
5
206
1
Queensland
2448756206
2000-01-27
2000-03-15
2000-01-27
QM
-27.443611
Site
20
152.90388
Enoggera Reservoir
5
206
1
Queensland
2448758211
2000-03-15
2000-05-18
2000-03-15
QM
-27.443611
Site
20
152.90388
Enoggera Reservoir
5
206
1
Queensland
2448756212
1999-10-16
1999-12-21
1999-10-16
QM
-27.445555
Site
20
152.92195
Enoggera Reservoir
5
206
1
Queensland
2448758213
[222,1316,872,899]
1999-12-21
2000-01-27
1999-12-21
QM
-27.445555
Enoggera Reservoir
20
152.92195
5
206
1
Queensland
2448759202
2000-03-15
2000-05-18
2000-03-15
QM
-27.445555
Enoggera Reservoir
20
152.92195
5
206
1
Queensland
2448756203
1999-07-06
1999-10-16
1999-07-06
QM
-27.435555
McAfee's Lookout
20
152.87138
5
206
1
Queensland
2448757204
06-11-15
2007-10-14
06-11-15
QM
-27.461945
Site
20
152.8725
Gold Creek Reservoir
5
206
1
Queensland
2448756205
07-11-15
2007-10-14
07-11-15
QM
-27.462778
Gold Creek Reservoir
20
152.88167
5
206
1
Queensland
2448756213
[373,1228,1016,1043]
2001-02-06
2001-02-07
2001-02-06
QM
-27.47889
Slaughter Falls
20
152.95944
5
206
1
Queensland
2448758202
2011-02-12
2011-02-13
2011-02-12
ANIC
G. B. Monteith
-27.4825
Brookfield
20
152.91695
Boscombe Road
5
206
1
Queensland
2448758214
[246,1223,1088,1115]
2003-11-04
31-12-21
2003-11-04
QM
-27.468056
Site
20
152.87222
Gold Creek Reservoir
5
206
1
Queensland
2448757201
1975-08-24
1975-09-07
1975-08-24
QM
R. I. Storey
-27.483334
Brisbane
20
152.86667
Upper Brookfield Road
5
206
1
Queensland
2448758201
1975-09-22
1975-10-04
1975-09-22
QM
R. I. Storey
-27.483334
Brisbane
20
152.86667
Upper Brookfield Road
5
206
5
Queensland
2448756214
06-06-09
2010-11-17
06-06-09
ANIC
-27.491112
Kenmore Kookaburra Street
20
152.92307
5
206
1
Queensland
2448756209
2011-01-20
2011-01-21
2011-01-20
ANIC
G. B. Monteith
-27.491112
Kenmore Hills
20
152.92334
5
206
2
Queensland
2448756208
2011-01-20
2011-01-21
2011-01-20
ANIC
G. B. Monteith
-27.506666
Kenmore
20
152.93112
5
206
2
Queensland
2448759209
2010-11-16
89-06-08
2010-11-16
ANIC
G. B. Monteith
-27.506945
Kenmore
20
152.93112
Mabb Street
5
206
1
Queensland
2448759207
1999-01-13
1999-05-16
1999-01-13
QM
-27.5075
The Knobby
20
152.58833
5
206
1
Queensland
2448758208
1999-01-13
1999-05-16
1999-01-13
QM
-27.508055
Cameron's Scrub
20
152.73166
5
206
1
Queensland
2448759206
2012-04-30
2012-05-01
2012-04-30
ANIC
G. B. Monteith
75
-27.508888
Cameron's Scrub
20
152.73
5
206
7
Queensland
2448757205
1999-11-01
1999-11-04
1999-11-01
QM
-27.509445
Long Pocket
20
152.99806
5
206
1
Queensland
2448758207
1999-01-13
1999-05-16
1999-01-13
QM
-27.509722
Cameron's Scrub
20
152.73166
5
206
1
Queensland
2448756211
1998-11-11
1999-01-13
1998-11-11
QM
-27.509722
Cameron's Scrub
20
152.73166
5
206
1
Queensland
2448757208
1999-01-13
1999-05-16
1999-01-13
QM
-27.5125
Cameron's Scrub
20
152.7275
5
206
1
Queensland
2550750804
[220,1303,1592,1619]
1997-12-12
1998-05-07
1997-12-12
QM
-27.513334
Belmont Hill
Belmont Hill
5
206
1
2550750808
1999-01-13
1999-05-16
1999-01-13
QM
-27.51361
Snig
152.7286
Scrub
5
206
1
2550750802
1998-11-11
1999-01-13
1998-11-11
QM
-27.51361
Snig
152.7286
Scrub
5
206
1
2550750811
2001-04-20
2001-04-22
2001-04-20
QM
-27.516111
Carnelian Street
153.07805
Whites Hill Reserve
5
206
1
2550750812
2001-04-09
2001-04-11
2001-04-09
QM
-27.516388
Pullenvale State Forest
152.86223
5
206
1
2550750801
2003-07-28
91-12-05
2003-07-28
QM
-27.5025
Carindale
Bulimba Creek
5
206
1
2448758209
2003-05-27
96-12-27
2003-05-27
QM
-27.5025
Carindale
Bulimba Creek
5
206
1
Queensland
2448756201
2001-04-20
2001-04-22
2001-04-20
QM
-27.516945
Camp Hill
20
153.08499
Sankey's Scrub
5
206
1
Queensland
2448756204
2005-11-24
2005-12-21
2005-11-24
QM
-27.518333
Cubberla Creek
20
152.9611
Rainbow Forest
5
206
1
Queensland
2448757203
2005-11-24
2005-11-25
2005-11-24
QM
-27.518333
Cubberla Creek
20
152.9611
Rainbow Forest
5
206
1
Queensland
2448758206
1999-01-13
1999-05-16
1999-01-13
QM
-27.603056
Perry's Knob
20
152.60472
5
206
1
Queensland
2448759204
1998-11-11
1999-01-13
1998-11-11
QM
-27.603056
Perry's Knob
20
152.60472
5
206
1
Queensland
2448756215
1997-12-12
1998-05-07
1997-12-12
QM
6
207
-27.616112
Scott's Dam
20
153.20583
Mount Cotton
5
206
1
Queensland
2448759236
1997-12-12
1998-05-07
1997-12-12
QM
-27.616112
Mount Cotton
20
153.2089
6
207
1
Queensland
2448758237
1997-09-03
1997-12-12
1997-09-03
QM
-27.616112
Mount Cotton
20
153.20583
6
207
1
Queensland
2448757231
2015-11-02
2015-11-20
2015-11-02
ANIC
G. B. Monteith & K. Ebert
410
-27.635
Burtons
20
152.02805
6
207
6
Queensland
2448759235
1984-03-03
CMNC
L. Masner
700
-27.886389
Mount Tamborine National Park
20
153.18083
6
207
1
Queensland
2448759234
1973-03-01
1973-03-02
1973-03-01
QM, QDPC
R. I. Storey
-28.05
Cunninghams Gap
20
152.4
6
207
2
Queensland
2448756238
[401,1432,368,395]
1994-02-20
1994-04-03
1994-02-20
QM
-28.074446
Spicer's Gap
20
152.42307
Moss' Well
6
207
1
Queensland
2448757240
[151,1199,404,431]
2000-10-06
2000-12-03
2000-10-06
QM
-28.125555
Black Rock Scrub
20
152.66333
6
207
1
Queensland
2448756234
2000-12-02
2001-05-13
2000-12-02
QM
-28.125555
Black Rock Scrub
20
152.66333
6
207
1
Queensland
2448759233
2007-03-30
QM
-28.134
Site
Lamington National Park
6
207
1
Queensland
2448758244
2007-01-18
2007-01-23
2007-01-18
QM
-28.142
Site
1
153.133
Lamington National Park
6
207
1
Queensland
2448759241
2006-10-16
2006-10-21
2006-10-16
QM
-28.154999
Site
15
153.13899
Lamington National Park
6
207
1
Queensland
2448757239
2012-02-29
2012-03-01
2012-02-29
ANIC
G. B. Monteith
610
-28.188889
Binna Burra
20
153.18611
6
207
1
Queensland
2448759237
1975-10-05
1976-02-22
1975-10-05
QM
-28.223612
Boonah
20
152.47916
Teviot Falls
6
207
1
Queensland
2448759238
2006-04-01
2006-04-02
2006-04-01
QM
-28.24
Mount Wunburra
20
153.2675
Springbrook
6
207
1
Queensland
2448756241
1975-10-05
1976-02-22
1975-10-05
QM
-28.315832
Boonah
20
152.84805
Mount French
6
207
1
Queensland
2448757238
1987-01-02
1987-01-11
1987-01-02
ANIC
A. Newton & M. Thayer
Unumgar State Forest
430
-28.4
Pole Bridge
20
152.66667
Woodenbong
6
207
4
New South Wales
2448757241
1973-02-24
1973-02-25
1973-02-24
QM
R. I. Storey
-28.4
Mount Lindsay Hwy
20
152.86667
6
207
1
New South Wales
2448757237
1993-04-13
ANIC
D. S. Chandler
350
-28.45
Border Ranges National Park
20
152.88333
6
207
1
New South Wales
2448757233
1982-08-25
ANIC
S. & J. Peck
50
-28.816668
Wilson Park
20
153.26666
6
207
2
New South Wales
2448756236
1998-11-20
1998-11-24
1998-11-20
GWC
G. Williams
-29.4
Iluka Nature Reserve
20
153.35
6
207
1
New South Wales
2448756235
1999-03-21
79-06-07
1999-03-21
AM
S. Lassau & C. Lemann
Kangaroo River State Forest
400
-30.075
Lowana
20
152.86806
Burns Road
6
207
1
New South Wales
2448757236
1997-12-02
GWC
G. Williams
-30.366667
Toormina
20
153.1
Toormina Park
6
207
1
New South Wales
2448758240
2011-09-23
2011-09-26
2011-09-23
ANIC
N. Gunter
Scotchman State Forest
-30.423056
Horseshoe Road
20
152.82916
Grahams
6
207
2
New South Wales
2448759239
2011-09-23
2011-09-26
2011-09-23
ANIC
N. Gunter
-30.528055
Hungry Head Road
20
153.0061
Newry State Forest
6
207
6
New South Wales
2448757230
1993-01
81-06-07
1993-01
AM
I. Oliver
-30.629444
Walcha Survey Big Hill
20
152.235
6
207
2
New South Wales
2448758236
1999-01-20
GWC
G. & T. Williams
-31.25
Racecourse Headland
20
152.96666
6
207
1
New South Wales
2448756243
2010-10-23
88-06-08
2010-10-23
ANIC
G. & T. Williams
-31.316668
Limeburners Creek National Park
20
152.96666
Point Plomer
6
207
1
New South Wales
2448758235
GWC
D. Bray
530
-31.566668
Doyles River
20
152.1
Winmurra
6
207
1
New South Wales
2448759240
1981-10-08
ANIC, GWC
G. & T. Williams
-31.733334
Yarratt State Forest
20
152.43333
Eastern
6
207
3
New South Wales
2448757232
2010-10-17
87-06-08
2010-10-17
ANIC
G. & T. Williams
-31.75
Lorien Wildlife Refuge
20
152.53334
6
207
1
New South Wales
2448756242
1986-05-12
1986-06-12
1986-05-12
CMNC
H. F. & A. T. Howden
-31.75
Lorein
20
152.53334
6
207
1
New South Wales
2448756239
1987-04-01
CMNC
H. F. & A. T. Howden
-31.75
Lorien
20
152.53334
Lansdowne
6
207
1
New South Wales
2448758238
1981-04-12
1981-04-14
1981-04-12
ANIC
G. & T. Williams
-31.783333
Lansdowne
20
152.53334
Newbys Lane
6
207
1
New South Wales
2448758239
1981-04-10
1981-04-12
1981-04-10
GWC
G. & T. Williams
-31.866667
Lansdowne
20
152.48334
Lorien
6
207
2
New South Wales
2448756237
1984-10-24
1984-10-25
1984-10-24
GWC
G. Williams
Lansdowne Recreational Reserve
-31.783333
Taree
20
152.53334
Lansdowne
6
207
1
New South Wales
2448757229
1991-12-19
1991-12-20
1991-12-19
GWC
Williams
-31.816668
24 km NNW Gloucester
20
151.86667
6
207
1
New South Wales
2448757228
1990-08-18
78-06-07
1990-08-18
AM
Williams
-31.833334
3.5 km NE of Harrington
20
152.71666
6
207
1
New South Wales
2448758242
1982-09-28
GWC
G. & T. Williams
-31.85
Crowdy
20
152.75
6
207
2
New South Wales
2448757235
1976-11-14
SAM
G. Williams
-31.866667
Harrington
20
152.7
6
207
2
New South Wales
2448757234
1976-12-25
GWC
G. Williams
-31.866667
Harrington
20
152.7
6
207
1
New South Wales
2448758241
1976-12-20
ANIC
G. Williams
-31.866667
Harrington
20
152.7
6
207
1
New South Wales
2448756240
[331,1432,1909,1936]
1976-10-10
ANIC
G. Williams
-31.866667
Harrington
20
152.7
6
207
1
New South Wales
2448758243
[151,1319,1945,1972]
1992-05-25
ANIC
T. Gush
-31.95
Kiwarrak State Forest
20
152.45
6
207
1
New South Wales
2448757227
1979-09-19
86-06-07
1979-09-19
AM
W. King
-31.966667
Taree
20
152.43333
Kiwarrak State Forest
6
207
1
New South Wales
2448756233
1993-05-29
1993-06-01
1993-05-29
ANIC
V. Lorimer
7
208
-31.966667
Kiwarrak State Forest
20
152.46666
6
207
1
New South Wales
2448756232
1982-02-17
ANIC, GWC
G. & T. Williams
-31.983334
Taree
20
152.41667
Kiwarrak State Forest
7
208
3
New South Wales
2448758233
1981-10-22
GWC
G. & T. Williams
-32.0
Old Bar
20
152.56667
Saltwater
7
208
1
New South Wales
2448758234
1981-10-13
GWC
G. & T. Williams
-32.066666
Hallidays Point
20
152.55
Black Head
7
208
2
New South Wales
2448756231
1981-11-12
1981-11-14
1981-11-12
ANIC
T. A. Weir
-32.066666
Barrington Tops National Park
20
151.68333
Gloucester River
7
208
1
New South Wales
2448756230
1993-02-04
75-06-07
1993-02-04
AM
G. Cassis & M. Gray
-32.139446
Kunungra Road
20
151.73445
Mountain Road
7
208
1
New South Wales
2448756224
1981-09-30
GWC
G. & T. Williams
South
-32.216667
Forster
20
152.56667
Cape Hawke
7
208
1
New South Wales
2448759231
1982-06-11
1982-08-27
1982-06-11
ANIC
S. & J. Peck
50
-32.333332
O'Sullivan Gap Reserve
20
152.28334
7
208
2
New South Wales
2448758231
1993-05-01
ANIC
V. Lorimer
-32.5
Bombah Point Swamp
20
152.28334
7
208
1
New South Wales
2448757225
1981-10-26
ANIC, GWC
G. & T. Williams
-32.533333
Myall Lakes National Park
20
152.31667
Mungo Brush
7
208
4
New South Wales
2448758230
1993-05-01
ANIC
V. Lorimer
-32.6
Mungo Brush
20
152.26666
7
208
2
New South Wales
2448759228
2002-03-05
2002-03-09
2002-03-05
ANIC
G. Hangay
-32.833332
Olinda Station
20
150.35
Mount Coricudgy
7
208
3
New South Wales
2448757226
[151,1263,656,683]
1976-07-25
ANIC
B. B. Lowery
-33.2
Gowan
20
149.36667
7
208
4
New South Wales
2448756228
1980-01-05
GWC
G. & T. Williams
-33.483334
Mount Irvine
20
150.46666
7
208
1
New South Wales
2448756229
1985-02-04
1985-02-05
1985-02-04
GWC
G. Williams
South
-33.6
Hat Head
19
150.33333
Hat Hill
7
208
2
New South Wales
2448759224
1998-12
ANIC
G. Hangay
-33.683334
Katandra Bushland Sanct.
19
151.26666
Ingleside
7
208
1
New South Wales
2448759225
1979-12-26
GWC
G. & T. Williams
-33.7
Springwood
19
150.53334
Sassafras Gully
7
208
2
New South Wales
2448757224
1995-12
80-06-07
1995-12
AM
G. O'Reilly
-33.75
West Pennant Hills
19
151.03334
7
208
1
New South Wales
2448759232
[252,1333,872,899]
1991-07-07
ANIC
T. Gush
-33.766666
West Killara
19
151.11667
7
208
2
New South Wales
2448757223
1967-12-15
SAMA
Matthews
-35.666668
5 mi. N Batemans Bay
19
150.25
7
208
2
New South Wales
2448759230
1978-03-06
CMNC
Peck
-35.666668
5 mi. N Batemans Bay
19
150.25
7
208
1
New South Wales
2448758232
1974-12-22
CMNC
H. F. & A. T. Howden
-34.775833
Gerroa
19
150.79695
7
208
1
New South Wales
2448759229
1999-03-12
87-06-07
1999-03-12
ANIC
L. Wilkie & R. Harris & Smith
-36.28
Princes Highway Corunna State Forest
19
150.12584
7
208
50
New South Wales
2448759226
1968-01-13
SAMA
E. Matthews
-37.5
Mallacoota National Park
19
149.76666
7
208
1
Victoria
2448756227
1978-05-21
1978-05-26
1978-05-21
ANIC
S. & J. Peck
-37.5
Mallacoota National Park
19
149.76666
7
208
1
Victoria
2448756226
1978-05-26
ANIC
S. & J. Peck
-37.5
Mallacoota National Park
19
149.76666
7
208
1
Victoria
2448756225
1993-02-10
1993-02-23
1993-02-10
ANIC
A. Newton & M. Thayer
20
-37.516666
South West Arm Road
19
149.68333
Croajingolong National Park
7
208
3
Victoria
2448759227
1978-05-22
1978-05-26
1978-05-22
CMNC
S. & J. Peck
3
-37.52972
Double Creek
19
149.68695
Mallacoola National Park
7
208
1
Victoria