Merriam, 1897 :229 Merriam, 1897 :229 Jackson, 1928 :158 Jackson, 1928 : 159 Woodman et al. 2012 : 214 The taxonomic status of Long-tailed shrews (Mammalia: genus Sorex) from Nuclear Central America Matson, John O. Ordóñez-Garza, Nicté Zootaxa 2017 4236 3 461 483 Merriam, 1897 Merriam 1897 [151,503,603,629] Mammalia Soricidae Sorex Animalia Soricomorpha 15 476 Chordata species salvini     Sorex salvini  Merriam, 1897:229. Type locality: “Calel, Totonicapan [sic], Guatemala (alt., 10200 ft. = 3100 meters).”    Sorex godmani  Merriam, 1897:229. Type locality: “Volcano Santa Maria, Quezaltenango [sic], Guatemala (alt., 9000 ft. = 2740 meters).    Sorex saussurei godmani:  Jackson, 1928:158    Sorex saussurei salvini:  Jackson, 1928: 159    Sorex salvini:  Woodman et al.2012: 214     Holotype. Number77035, U. S.Nat. Mus., Biological Survey Collection; adult, female, skin and skull; collected  January 12, 1896, by E. W. Nelsonand E. A. Goldman, original number 9 0 57.   Distribution.Highlands of northwestern, central, and eastern  Guatemala.   Diagnosis.  Sorex salviniis a member of the  Sorex salvinispecies group. It is the largest of this species group in body measurements (except for  S. sclateri), but is intermediate in skull size ( Table 1). The color of the dorsum is more gray/brown than in  S. cristobalensis, about the same color below; tail is gray/brown above, slightly paler below. Specimens from Yaiquich, Huehuetenango, are more chocolate brown, as is a topotype (USNM 77071). U3 is usually taller than U4, but the typespecimen has U3 = U4.   Description.The body size of  S. salviniaverages larger than in  S. cristobalensis, S. mccarthyi, or  S. stizodon, but smaller than  S. sclateri. The skull averages smaller in most measurements than in  S. cristobalensisor  S. sclateri, but larger than in  S. mccarthyior  S. stizodon. Sorex salvinihas a relatively short mandible, longer toothrow, and shorter rostrum compared to  S. cristobalensisand  S. mccarthyi.  Ecology.  Sorex salviniis found in broad-leaved and coniferous cloud forests of Alta Verapaz, Huehuetenango, Quetzaltenango, and Totonicapán. Depending upon the mountain range, known small mammal associates include  Cryptotis mam, C. lasertosus, C. oreoryctes, Habromys lophurus, Handleyomyssaturatior, Heteromys desmarestianus, Marmosamexicana, Microtus guatemalensis, Nyctomys sumichrasti, Oligoryzomysfulvescens, Peromyscus beatae, P. grandis, P. guatemalensis, P. mayensis, Reithrodontomys microdon, R. sumichrasti, R. tenuirostris, Scotinomysteguina, and  Sorex veraepacis( Carleton & Huckaby 1975; Woodman 2010, Woodman 2011b; Matson et al.2014). Nothing is known about its reproductive habits.    Specimensexamined (14). Guatemala, Alta Verapaz: Finca Chinaux, Campamentode los Helechos,  2040 m( CM 120109– 120110);  Huehuetenango: San Mateo Ixtatán, Cerro Bobí(  USAC26);  San Mateo Ixtatán, Aldea Chiloala Zum(  USAC27);  Todos Santos(  USNM77023);  Yiaquich,  4 kmNW Sta Eulalia, San Mateo Ixtatán,  2950 m(  UMMZ117849–117850);  Quiche: Chimel Grande, 12.6 kmN,  9 kmE Uspantan,  2490 m, 15.46208 N, 90.78811 W(  MVZ226939);  Quetzaltenango: Volcán Santa María  9000 ft. (  USNM77044–77046);  4 kmSE Zunil( Finca la Chingada)  2720 m(  USNM569591);  Totonicapán, Calel,  10,200 ft. (  USNM77035, 77071). 1437075708 1896-01-12 E. W. Nelson & E. A. Goldman United States of America Number Biological Survey Collection 15 476 1 holotype 1437075711 Guatemala Specimens 2040 Campamento Finca Chinaux 15 476 1 Alta Verapaz 1437075718 [423,1123,1635,1660] USAC Guatemala Cerro Bobi San Mateo Ixtatan 15 476 USAC 26 1 Huehuetenango 1437075724 USAC Guatemala Aldea Chiloala Zum San Mateo Ixtatan 15 476 USAC 27 1 Huehuetenango 1437075734 [453,788,1671,1696] USNM Guatemala Todos Santos 15 476 USNM 77023 1 Huehuetenango 1437075738 UMMZ Guatemala 2950 San Mateo Ixtatan Yiaquich 15 476 UMMZ 117849-117850 1 Huehuetenango 1437075748 MVZ Guatemala 2490 15.46208 Chimel Grande 1 -90.78811 15 476 MVZ 226939 1 Quiche 1437075757 [483,1261,1742,1767] USNM Guatemala 2743 Volcan Santa Maria 15 476 USNM 77044-77046 1 Quetzaltenango 1437075764 USNM Finca la Chingada Guatemala 2720 Zunil 15 476 USNM 569591 1 Quetzaltenango 1437075775 [677,1282,1777,1802] USNM Guatemala 3109 Calel 15 476 USNM 77035, 77071 1 Totonicapan