lata Dall, 1889b: 324–325 Aclis lata : Dall (1889a: 126 Hemiaclis lata : Bartsch (1947: 18 Umbilibalcis lata : Bouchet & Warén (1986: 444–445 Revision of the deep-sea Eulimidae (Gastropoda, Caenogastropoda) from off Northeast Brazil Souza, Leonardo Santos De Pimenta, Alexandre Dias Barros, José Carlos Nascimento De Zootaxa 2021 2021-02-16 4927 4 451 504 7DGTT (Dall, 1889 b) Dall 1889 [151,526,1680,1707] Gastropoda Eulimidae Umbilibalcis Animalia Littorinimorpha 42 493 Mollusca species lata  ( Figure 24A–I)     Aclis( supranitida var.?)  lata Dall, 1889b: 324–325, plate 18, figure 8.     Aclis lata: Dall (1889a: 126, plate 18, figure 8) [reproduced from original illustration].     Hemiaclis lata: Bartsch (1947: 18, plate 4, figure 4)     Umbilibalcis lata: Bouchet & Warén (1986: 444–445, figure 1056–1057).   Type material.  Holotype USNM 126790( Figure 24A). Type locality. Barbados,   183 m.   FIGURE 24.A–I.  Umbilibalcis lata( Dall, 1889b). A. Holotype of  Aclis lata Dall, 1889b, USNM 126790. B–I. IBUFRJ 14399. E. Detail of teleoconch sculpture. H. Detail of protoconch, frontal view. I. Detail of protoconch, apical view. Scale bars: A–D= 1.0 mm; E, H= 100 µm; F, G= 500 µm; I= 50 µm.  Material examined.  Typematerial; USA: USFC stn. 2404 ( 28°44′N 85°16′W,  110 m), coll.  15/iii/1885: USNM 323963[1 dd]; USBF stn. 20030 (~24°25′N 82°58′,  164 m):  USNM 407033[1 dd]. Antigua and Barbuda: English Harbor, S.U.I.stn. 496 (  219 m): USNM 459689[1 dd]. Barbados:  Off Pelican Islands, S.U.I. Expedition stn. 454 (  182 m): USNM 459677[1 dd]; Off Lazareto, S.U.I. Expedition stn. 497 (  171 m):  USNM 459708[1 dd]. Brazil: Rio Grande do Norte: ( 04°51′S 35°06′W,  375 m),  24/xi/2001: MNRJ 35200† [1 dd];  Bahia: ( 11°35′31″S 37°12′18″W,  510 m),  30/x/2000:  MNRJ 35268† [3 dd]; REVIZEE-Central V stn. 504 ( 14°28′55″S 38°54′03″W,  278 m), coll. Supply Boat Astro Garoupa,  02/vii/2001:  IBUFRJ 14399† [5 dd]; Espírito Santo: REVIZEE-Central V stn. 25a ( 19°33′28″S 38°45′28″W,  230 m), coll. Supply Boat Astro Garoupa,  28/vi/2001:  IBUFRJ 19717† [3 dd]; Rio de Janeiro: Campos Basin( 22°18′53″S 40°11′59″W,  200 m), coll. Supply Boat Astro Garoupa,  18/xii/2004:  MNRJ 31867† [2 dd];  Campos Basin( 22°42′33″S 40°41′24″W,  110 m), coll.  19/ix/2003:  MNRJ 31881† [2 dd], MNRJ 31895† [1 dd]; PADCT stn. 6541:  MNRJ 28285† [3 dd]; PADCT stn. 6627: MNRJ 28287† [1 dd]; Cabo Frio VII stn. 6194 ( 24°03′36″S 44°07′36″W,  134 m), coll. RV Almirante Saldanha,  01/iv/1983:  IBUFRJ 19727† [1 dd]; S„o Paulo: REVIZEE Sulstn. 6672: MNRJ 28289† [1 dd]; PADCT stn. 6573: MNRJ 23226† [1 dd];  Paraná: REVIZEE Sulstn. 6653: MNRJ 28288† [1 dd];  Santa Catarina: REVIZEE Sulstn. 6695: MNRJ 28290† [1 dd]; PADCT stn. 6595: MNRJ 28283† [7 dd]; PADCT stn. 6606: MNRJ 28286† [1 dd];  Rio Grande do Sul: off Bojuru(  220 m), coll. RV Almirante Saldanha,  xi/1979: MORG 21416† [1 dd].   Redescription ( Holotype).Shell conical, straight, reaching about 5.4 mmlong, 2.4 mmwide. Larval shell brownish, partially broken. Teleoconch with about 8.0 whorls, slightly convex outline; suture distinct, sloping; subsutural region not visible; surface polished, presenting slightly opisthocline axial lines; incremental scars slightly demarcated, appearing at irregular intervals. Last whorl about 45% of the shell length; base rounded.Aperture wide, pear shaped, acute posteriorly and rounded anteriorly; outer lip thin, slightly prosocline, sinuous, slightly retracted near the suture, reaching a maximum projection in the middle of its height, retracted at distal region; inner lip sinuous, sloping, reflected and well demarcated. Umbilicus present. Shell whitish.  Measurements. HolotypeUSNM 126790, 10 whorls (protoconch partially broken), SL= 5.4 mm; BWL= 2.5 mm; AL= 1.6 mm; SW= 2.4 mm; AW= 1.2 mm. IBUFRJ 14399, 10 whorls, SL= 5.0 mm; BWL= 2.3 mm; AL= 1.4 mm; SW= 2.2 mm; AW= 1.1 mm.   Remarks.The protoconch is partially broken in the holotypeof  Umbilibalcis lata( Figure 24A), but the general shape, dimensions and sculpture fits perfectly with specimens from Brazil( Figure 24B–I). The protoconch of the shells from Brazilhave about 3.0 whorls, the first (embryonic) whorl with a smooth surface, the remaining whorls with strongly opisthocline axial lines ( Figure 24H–I). The protoconch is about 270 μm wide and 390 μm in height. A weak brownish band close to suture is present on the protoconch of fresh specimens. The outer lip of the holotypeis also partially broken, but it seems uniform in all specimens that the proximal area is slightly advanced, giving a prosocline outline.   Umbilibalcis latadiffers from  U. crassula Bouchet & Warén, 1986and  U. subumbilicata(Jeffreys, 1883)by the higher number of protoconch whorls (3.0 vs. 1.5 in  U. crassulaand  U. subumbilicata) and by the straighter outer lip ( Bouchet & Warén 1986).  Geographical distribution. USA: Florida (present study); Antigua and Barbuda(present study); Barbados( Dall 1889b); Brazil: Rio Grande do Norte, Bahia, Espírito Santo, Rio de Janeiro, S„o Paulo, Paraná, Santa Catarina, Rio Grande do Sul(present study).  Bathymetric distribution.From 60 mto  375 m. 3041480308 USNM Barbados 183 Barbados 42 493 USNM 126790 183 holotype 3041480325 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 43 494 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480343 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 44 495 -24.06 Campos Basin 21 -44.126663 Campos Basin 43 494 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480361 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480310 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480306 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480346 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480351 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480338 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480366 [331,1090,367,395] 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480301 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480356 [327,536,402,429] 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480319 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480368 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480386 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480385 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480315 1885-03-15 2004-12-18 1885-03-15 USNM, MNRJ, IBUFRJ, MORG Antigua and Barbuda Off Pelican Islands 110 -24.06 Campos Basin 21 -44.126663 Campos Basin 44 495 USNM 323963, USNM 407033, USNM 459689, USNM 459677, USNM 459708, MNRJ 35200, MNRJ 35268, IBUFRJ 14399, IBUFRJ 19717, MNRJ 31867, MNRJ 31881, MNRJ 31895, MNRJ 28285, MNRJ 28287, IBUFRJ 19727, MNRJ 28289, MNRJ 23226, MNRJ 28288, MNRJ 28290, MNRJ 28283, MNRJ 28286, MORG 21416 1 Rio Grande do Norte holotype 3041480326 Antigua and Barbuda Rio Grande do Norte 44 495 1 Rio Grande do Norte