Typhlops fornasini Bianconi, 1849a: 107
C. Fornasini
Bianconi, 1849b: 203
Bocage, 1895: 64
1896a: 99
Branch 1986: 285
Wallach, 1993b: 33
Frank & Ramus, 1995: 253
McDiarmid et al., 1999: 102
Typhlops fornasinii Bianconi, 1850a: 183
Jan, 1863: 12
Jan & Sordelli, 1864c: 4
Peters, 1882: 94
Boulenger, 1893: 38
1910: 497
Werner, 1921: 313
Schmidt, 1923: 33
Barbour & Loveridge, 1929: 357
Rose, 1950: 252
1955: 74
1962: 320
Brynard & Pienaar, 1960: 31
Isemonger, 1968: 81
1983: 84
Broadley, 1973: 20
Roux-Estève, 1974: 146
Broadley & Cock, 1975: 101
Bruton & Haacke, 1975: 26
1980: 262
Bourquin, 1977: 46
1989: 20
Hahn, 1977: 37
1980: 58
Poynton & Broadley, 1978: 941
Broadley & Blake, 1979: 10
Welch, 1982: 128
1994: 67
Hughes, 1983: 354
Haagner & Els, 1986: 20
Branch, 1986: 290
1994: 47
Marais, 1992: 178
Meirte, 1992: 22
Wallach, 1993a: 217
1993b: 33
1998a: 183
1998b: 370
Haagner, 1994: 34
Broadley & Kalk, 1995: 322
Mattison, 1999: 147
McDiarmid et al., 1999: 102
Webb et al., 2001: 559
Murphy, 2003: 552
Brogard, 2005: 115
Schneider et al., 2005: 116
Marais, 2007: 52
Maritz & Alexander, 2009: 6
Onychocephalus mossambicus Peters, 1854
fide Bauer et al., 2002: 161
Peters, 1855: 50
1882: 93
Lichtenstein, 1856: 21
Günther, 1866: 25
Barbour & Loveridge, 1929: 310
Roux-Estève, 1974: 146
Hahn, 1980: 58
Bauer et al., 1995: 78
McDiarmid et al., 1999: 102
Onychocephalus trilobus Peters, 1854
Peters, 1855: 50
1860: 80
Lichtenstein, 1856: 21
Barbour & Loveridge, 1929: 310
Hahn, 1980: 58
Bauer et al., 1995: 78
McDiarmid et al., 1999: 102
Ophtalmidion (sic) mossambicus Jan, 1857: 43
Onychocephalus tettensis Peters, 1860
Peters, 1882: 92
Roux-Estève, 1974: 146
Hahn, 1980: 58
Bauer et al., 1995: 78
McDiarmid et al., 1999: 102
Typhlops (Ophthalmidion) bianconii Jan & Sordelli, 1860: 1
Jan, 1863: 13
fide Jan, 1864: 23
fide Jan & Sordelli, 1860: 1
Jan, 1863: 13
Hahn, 1980: 58
McDiarmid et al., 1999: 102
Typhlops (Typhlops) fornasinii
Jan, 1863: 12
Hahn, 1980: 58
Typhlops (Ophthalmidion) mossambicus
Jan, 1863: 13
McDiarmid et al., 1999: 102
Typhlops bianconii
Jan, 1863: 13
Roux-Estève, 1974: 146
Typhlops mossambicus
Jan & Sordelli, 1864c: 4
Peters, 1882: 93
Boulenger, 1887:174
1893: 41
1908a: 227
1910: 498
1915b: 615
Bocage, 1896a: 99
Sclater, 1898: 97
Gough, 1908: 18
Sternfeld, 1908b: 242
Werner, 1921: 314
Schmidt, 1923: 33
Loveridge, 1924: 3
Essex , 1927: 925
Barbour & Loveridge, 1929: 357
Rose, 1950: 252
1955: 74
1962: 320
Brynard & Pienaar, 1960: 31
Isemonger, 1962: 93
1968: 83
1983: 87
Pooley, 1965: 47
Hahn, 1977: 35
1980: 58
Welch, 1982: 128
McDiarmid et al., 1999: 102
Typhlops tettensis
Peters 1882: 92
Boulenger, 1915b: 615
Werner, 1921: 314
Schmidt, 1923: 33
Broadley, 1961: 3
Hahn, 1980: 58
McDiarmid et al., 1999: 102
Brogard, 2005: 116
Typhlops trilobus
Typhlops tetensis
Bocage, 1896a: 99
Typhlops mossombicus
Werner, 1921: 336
Hahn, 1980: 58
McDiarmid et al., 1999: 102
Typhlops bianconi
Rhinotyphlops fornasinii
Broadley & Wallach, 2000: 167
2255
Broadley, Donald G.
Wallach, Van
Zootaxa
2009
2009-10-08
2255
1
100
(Bianconi)
Bianconi
1847
[151,571,151,177]
Reptilia
Typhlopidae
Afrotyphlops
Animalia
Squamata
26
27
Chordata
species
fornasinii
( Fig. 4A) Fornasini’s dwarf blind-snake
Typhlops fornasini Bianconi, 1849a: 107(incorrect original spelling). Type locality: “ Mozambico” [= Mozambique], collected by C. Fornasini, holotypeMBI; Bianconi, 1849b: 203; Bocage, 1895: 64, 1896a: 99; Branch 1986: 285; Wallach, 1993b: 33; Frank & Ramus, 1995: 253; McDiarmid et al.,1999: 102. Typhlops fornasinii Bianconi, 1850a: 183, pl. 6, fig. 1, 1850b in1850–1870: 13, pl. 3, fig. 1; Jan, 1863: 12, 1864: 17; Jan & Sordelli, 1864c: 4, pl. 5, fig. 5, pl. 6, fig. 5; Peters, 1882: 94, pl. 15, fig. 3; Boulenger, 1893: 38, 1910: 497; FitzSimons, F., 1912 & 1919: 52; Werner, 1921: 313; Schmidt, 1923: 33; Barbour & Loveridge, 1929: 357; Rose, 1950: 252, 1955: 74, 1962: 320; Brynard & Pienaar, 1960: 31; FitzSimons, V., 1962: 65, fig. 4, 1966a: 37, 1970 & 1974: 65, 1978: 55; Isemonger, 1968: 81, 1983: 84; Laurent, 1964: 421, 1968: 1; Broadley, 1973: 20, 1978: 29, 1983: 40, fig. 4, 1998c: 385, 1990a: 40 & 377, fig. 4, 1990b: 485; 1992: 543; Roux-Estève, 1974: 146, figs. 13.23 & 97, 1975a: 492, fig. 30; Broadley & Cock, 1975: 101; Bruton & Haacke, 1975: 26, 1980: 262; Bourquin, 1977: 46, 1989: 20; Hahn, 1977: 37, 1980: 58; Poynton & Broadley, 1978: 941; Broadley & Blake, 1979: 10; Welch, 1982: 128, 1994: 67; Hughes, 1983: 354; Haagner & Els, 1986: 20; Branch, 1986: 290, 1988a& 1994: 47, pl. 40, 1988b: 11, 1998: 54, pl. 40; Marais, 1992: 178, 3 figs., 286, 2 figs.; Meirte, 1992: 22; Wallach, 1993a: 217, 1993b: 33, 1998a: 183, 1998b: 370; Haagner, 1994: 34; Broadley & Kalk, 1995: 322; Mattison, 1999: 147; McDiarmid et al.,1999: 102; Webb et al.,2001: 559; Murphy, 2003: 552; Brogard, 2005: 115; Schneider et al.,2005: 116; Alexan- der & Marais, 2007: 52, 2 figs.; Maritz & Alexander, 2009: 6. Onychocephalus mossambicus Peters, 1854, Ber. Bekanntmach. Geeignet. Verhandl. Königl.-Preuss. Akad. Wiss. Berlin, 1854(11): 621. Type locality: “Insulae Mossambique et Anjoana” [= Mozambique Islandand Nzwani, Comoro Islands], syntypesMCZ 21007 (formerly ZMB 3948), ZMB 3949–50, 63441 (formerly ZMB 3949), collected by W.C.H. Peters, June 1843– March 1847(Anjoana syntypepresumably lost fideBauer et al.,2002: 161); Peters, 1855: 50, 1882: 93, pl. 15, fig. 2; Lichtenstein, 1856: 21; Günther, 1866: 25; Barbour & Loveridge, 1929: 310; Fitz- Simons, V., 1962, 66, 1966a: 38; Roux-Estève, 1974: 146; Hahn, 1980: 58; Bauer et al.,1995: 78, 2002: 161; McDiarmid et al.,1999: 102. Onychocephalus trilobus Peters, 1854, Ber. Bekanntmach. Geeignet. Verhandl. Königl.-Preuss. Akad. Wiss. Berlin, 1854(11): 621. Type locality: “Lourenzo-Marques, Inhambane” [= Maputoand Inhambane, Mozambique], syntypesMCZ 21006 (formerly ZMB 3917), ZMB 3915–16, collected by W.C.H. Peters, July–November 1846; Peters, 1855: 50, 1860: 80; Lichtenstein, 1856: 21; Barbour & Loveridge, 1929: 310; FitzSimons, V., 1962, 65, 1966a: 37; Roux- Estève, 1974: 146; Hahn, 1980: 58; Bauer et al.,1995: 78, 2002: 162; McDiarmid et al.,1999: 102. Ophtalmidion (sic) mossambicus Jan, 1857: 43. Onychocephalus tettensis Peters, 1860, Mber. Königl. Akad. Wiss. Berlin, 1860(2): 80. Type locality: “Mossambique, Tette” [= Tete, Mozambique], holotypeZMB 3944, collected by W.C.H. Peters, December 1844– September 1845; Peters, 1882: 92, pl. 15, fig. 1; Roux-Estève, 1974: 146; Hahn, 1980: 58; Bauer et al.,1995: 78, 2002: 162; McDiarmid et al.,1999: 102. Typhlops (Ophthalmidion) bianconii Jan & Sordelli, 1860: 1, pl. 5, fig. 3, pl. 6, fig. 3. Type locality: “Mosambique,” by restriction of type locality by Jan, 1863: 13, holotypeMZUT fideJan, 1864: 23( contraMSNM fide Jan & Sordelli, 1860: 1); Jan, 1863: 13, 1864: 23; Hahn, 1980: 58; McDiarmid et al.,1999: 102. Typhlops (Typhlops) fornasinii— Jan, 1863: 12; Hahn, 1980: 58. Typhlops (Ophthalmidion) mossambicus— Jan, 1863: 13; McDiarmid et al.,1999: 102. Typhlops bianconii— Jan, 1863: 13; Roux-Estève, 1974: 146. Typhlops mossambicus— Jan & Sordelli, 1864c: 4, pl. 5, fig. 3, pl. 6, fig. 3; Jan, 1864: 22; Peters, 1882: 93, pl. 15, fig. 2; Boulenger, 1887:174, 1893: 41, 1908a: 227, 1910: 498, 1915b: 615; Bocage, 1896a: 99; Sclater, 1898: 97; Gough, 1908: 18; FitzSimons, F., 1910: 26, 1912 & 1919: 52; Sternfeld, 1908b: 242; Werner, 1921: 314; Schmidt, 1923: 33; Loveridge, 1924: 3; Essex, 1927: 925, figs. 77–78; Barbour & Loveridge, 1929: 357; Pitman, 1938& 1974: pl. 1, fig. 1; Rose, 1950: 252, 1955: 74, 1962: 320; Brynard & Pienaar, 1960: 31; FitzSimons, V., 1962: 66, fig. 5, 1966a: 38, 1970 & 1974: 66, 1978: 56; Isemonger, 1962: 93, 1968: 83, 1983: 87; Laurent, 1964: 421; Pooley, 1965: 47; Hahn, 1977: 35, 1980: 58; Welch, 1982: 128; McDiarmid et al.,1999: 102. Typhlops tettensis— Peters 1882: 92, pl. 15, fig. 1; Boulenger, 1915b: 615; Werner, 1921: 314; Schmidt, 1923: 33; Broadley, 1961: 3; Hahn, 1980: 58; McDiarmid et al.,1999: 102; Brogard, 2005: 116. Typhlops trilobus— Peters 1882, pl. 15, fig. 3. Typhlops tetensis(sic)— Bocage, 1896a: 99. Typhlops mossombicus(sic)— Werner, 1921: 336; Hahn, 1980: 58; McDiarmid et al.,1999: 102. Typhlops bianconi—FitzSimons, V., 1966a: 38. Rhinotyphlops fornasinii— Broadley & Wallach, 2000: 167.
Description.Snout rounded, prominent. Rostral broad and slightly pointed behind; frontal small, subhexagonal; supraocular transverse, its lateral apex between preocular and ocular; eye beneath or slightly behind the anterior border of the ocular; nasal often completely divided, the nasal sulcus arising from the first labial; SIP T-II-O (N2, P/O, O, O); MSR 22–26, reduction A/B 0 to 2, B/C 0–4; MD 232–286 (mean 260.7, n= 35); vertebrae 144–160 (mean 154.3, n= 24); MD/V ratio 1.52–1.77 (mean 1.69, n= 23); L/D ratio 28–45 (mean 33.6, n= 28). Dark grey to black, with irregular yellowish patches below, especially on throat and cloacal region. Largest specimen (PEM 12121—Shangwane, Zululand, South Africa) has a total length of 185 mm.
Distribution.Coastal Mozambiqueand its offshore islands, extending into southeastern Zimbabweand northern KwaZulu-Natal, NSL- 65 m( Fig. 14). Localities. MOZAMBIQUE. Noprecise locality MNHN 178, 3213; Bazaruto Island NMZB 10002, 11444; Benguerua Island NMZB 11395–97; Inhaca Island NMZB-UM 2301; Inhambane MCZ21006 ( syntypeof Onychocepphalus trilobusPeters); Inharrime NMZB-UM 25586; Lourenco-Marques MCZ41947; Magaruque Island NMZB 10012; Maputo NMZB 11931–32; TMP 4829, 4831–35, 4841, 4843–47, 4849–50; ZMB 3915–16; Matola CZL 1; Mauele CZL 1157–59; Maxixe SAM44553; Mozambique Island MCZ21007 ( syntypeof Onychocephalus mossambicusPeters), ZMB 3949(2), 3950 ( syntypesof Onychocephalus mossambicusPeters); Tete(?) ZMB 3944; Vundiça TMP29461. ZIMBABWE. Malugwe Pan, Gonarezhou National Park NMZB-UM 12329. SOUTH AFRICA( Kwazulu-Natal). Mseleni TMP 4830, 4836–38, 4848; Shangwane, Tembe Elephant Park PEM 12119–21, 12123, 12173–77; Sibaya Research Station TMP46022; Sodwara BayPark TMP46001.
[339,1036,152,178]
MNHN
Mozambique
No
28
29
MNHN 178, 3213
1
NMZB
Mozambique
Bazaruto Island
28
29
NMZB 10002, 11444
1
[240,671,192,218]
NMZB
Mozambique
Benguerua Island
28
29
NMZB 11395-97
1
[685,1067,192,218]
Mozambique
Inhaca Island
28
29
NMZB-UM 2301
1
MCZ 21006
MCZ
Mozambique
Inhambane
28
29
1
Inhambane
syntype
[672,1029,232,258]
Mozambique
Inharrime
28
29
NMZB-UM 25586
1
syntype
MCZ 41947
[1044,1432,232,258]
MCZ
Mozambique
Lourenco-Marques
28
29
1
syntype
[151,540,272,298]
NMZB
Mozambique
Magaruque Island
28
29
NMZB 10012
1
syntype
[552,863,272,298]
NMZB
Mozambique
Maputo
28
29
NMZB 11931-32
1
Maputo
syntype
[875,1432,272,298]
TMP
Mozambique
true
Maputo
28
29
TMP 4829, 4831-35, 4841, 4843-47, 4849-50
1
Maputo
syntype
[151,328,312,338]
ZMB
Mozambique
28
29
ZMB 3915-16
1
syntype
[339,507,312,338]
CZL
Mozambique
Matola
28
29
CZL 1
1
syntype
[522,784,312,338]
CZL
Mozambique
Mauele
28
29
CZL 1157-59
1
syntype
SAM 44553
[796,1040,312,338]
SAM
Mozambique
Maxixe
28
29
1
syntype
MCZ 21007
MCZ
Mozambique
Mozambique Island
28
29
1
syntype
[751,923,352,378]
ZMB
Mozambique
28
29
ZMB 3949
1
syntype
ZMB
Mozambique
Tete
28
29
ZMB 3944
1
Tete
syntype
TMP 29461
[576,825,392,418]
TMP
Mozambique
Vundica
28
29
1
syntype
Zimbabwe
Gonarezhou National Park
Malugwe Pan
28
29
NMZB-UM 12329
1
syntype
[459,1338,432,458]
TMP
South Africa
Mseleni
28
29
TMP 4830, 4836-38, 4848
1
KwaZulu-Natal
syntype
PEM 12119
PEM
South Africa
Tembe Elephant Park
Shangwane
28
29
PEM 12119-21, 12123, 12173-77
1
syntype
TMP 46022
[922,1367,472,498]
TMP
South Africa
Sibaya Research Station
28
29
1
syntype
TMP 46001
TMP
South Africa
Sodwara Bay
28
29
1
syntype