Westwood, 1855 : 219
Thomson 1862 : 432
Parry 1863 : 450
Parry 1864b : 100
Harold 1868 : 965
Macleay 1871 : 173
Deyrolle 1874 : 414
Parry 1875 : 22
Felsche 1898 : 78
Van Roon 1910 : 52
Mjöberg 1916 : 13
Arrow 1943 : 139
Didier & Séguy 1953 : 180
Benesh 1950 : 53
Benesh 1955 : 64
Benesh 1960 : 26
Klausnitzer 1982 : 15
Bomans 1986 : 9
Maes 1992 : 36
Moore & Cassis 1992 : 10
Araya 2001 : 13
Krajcik 2001 : 11
Krajcik 2003 : 79
Klausnitzer & Sprecher-Uebersax 2008 : 29
Fujita 2010 : 395
Thomson, 1862 : 402
Parry 1863 : 450
Figulus ( Figulinus ) lilliputanus : Schoolmeesters 2015
Revision of the Australian species of Figulus MacLeay, 1819 (Coleoptera: Lucanidae)
Monte, Cinzia
Zilioli, Michele
Bartolozzi, Luca
Zootaxa
2016
4189
3
447
484
6HZRS
Westwood, 1855
Westwood
1855
[151,599,151,178]
Insecta
Lucanidae
Figulus
Animalia
Coleoptera
6
453
Arthropoda
species
lilliputanus
Figulus lilliputanus Westwood, 1855: 219; Thomson 1862: 432; Parry 1863: 450; Parry 1864b: 100; Harold 1868: 965; Macleay 1871: 173; Deyrolle 1874: 414; Parry 1875: 22; Felsche 1898: 78; Van Roon 1910: 52; Mjöberg 1916: 13; Arrow 1938: 53, 60, 61; Arrow 1943: 139; Didier & Séguy 1953: 180; Benesh 1950: 53; Benesh 1955: 64, 66; Benesh 1960: 26; Klausnitzer 1982: 15; Bomans 1986: 9( lectotypedesignation); Maes 1992: 36, 38; Moore & Cassis 1992: 10; Araya 2001: 13; Krajcik 2001: 11; Krajcik 2003: 79; Klausnitzer & Sprecher-Uebersax 2008: 29; Fujita 2010: 395; Maes 2014. Figulus clivinoides Thomson, 1862: 402, 432; synonymised by Parry 1863: 450. Figulus( Figulinus) lilliputanus: Schoolmeesters 2015.
Type material. Figulus lilliputanusWestwood, lectotype (designation by Bomans 1986): “S Austral [handwritten]”, “ Figulus/ Liliputanus(sic!) / Tr. Ent. Soc. n.s. III, pl. 12, fig. 5 [handwritten]”, “TYPE / WESTWOOD / Trans. Ent. / Soc. 1855 / P.219 T.12 f.5 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 333 1/ 2/ Figulus/ lilliputanus/ Westw. / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH]; Figulus lilliputanusWestwood, paralectotype: “S Austral [handwritten]”, “W [printed]”, “TYPE / WESTWOOD / Trans. Ent. Soc./ 1855 P.219. / T.12 fig.5 / Coll. Hope Oxon. [printed, handwritten]”, “TYPE COL: 333 2/ 2/ Figulus/ lilliputanus/ Westw. / HOPE DEPT. OXFORD [printed, handwritten]”, [OUMNH]; Figulus clivinoidesThomson: Lectotype here designated: Lectotype: “Th. / Type [printed]”, “ Clivinoides/ Type Thoms / Austr [handwritten]”, “Ex Musaeo / James Thomson [printed]”, “MUSÉUM PARIS / 1952 / COLL. R. OBERTHÜR [printed]”, “H.Bomans det., 1973 / = Figulus/ lilliputanusWest. [printed, handwritten]”, [MNHN]. Other material. Figulus lilliputanusWestwood(Ƌ): “ Lilliputanus/ West. Ent. J. / v. 3Feb [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]; Figulus lilliputanusWestwood(Ƌ): “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]; Figulus lilliputanusWestwood(♀): “ Lilliputanus/ West. / Clivinoides/ Thompson [handwritten]”, “Ex Musaeo / Parry [printed]”, “MUSÉUM PARIS / 1952 / Coll. R. OBERTHÜR [printed]”, [MNHN]. Note: this material was indicated as syntypes by S. Boucher (MNHN) in the loan form, but their status as type material is uncertain, as they have no red label or clear indication of being part of the original type series.
Diagnosis.This is the smallest species of Australian Figulusand differs from all other Australian species by its antenna with only 9 antennomeres. The irregular transversal rows of dense and deep punctures with short erected setae directed downwards on the ventrites 2‒4 are secondary sexual characters, as in other Figulusspecies ( Bomans 1989).
Redescription.Measurements: size range ( n= 165): TL: 6.0–10.0 mm; PL: 1.6 1–2.49 mm; EL: 3.41–5.43 mm; PW: 1.69–2.97 mm; EW: 1.74–3.0 mm. Male ( Fig. 8). Body elongate, with subparallel sides, dark brown or reddish brown, glossy. Head: transverse, about twice as wide as long. Frontoclypeal region slightly concave at the anterior margin coarsely punctate with an evident median elevation and two large, glossy areas situated before the eyes. Labrum broad, short, with anterior margin slightly bilobed and with long, golden setae. Vertex slightly concave in the middle, feebly inclined anteriorly, with two small, circular elevations. Two longitudinal, slightly elevated carinae are present along the inner margin of eyes. Eyes small, divided into 2 parts by the canthi, the upper part smaller than the lower with distinct facets. Canthus slightly protruding, rounded laterally and angled behind; upper surface slightly depressed and smooth, with sparse, minute punctures on the lateral margin. Head surface densely punctate with superficial, large punctures smallest on canthi, anterior margin of the frons, longitudinal carinae and transversal elevations of vertex. Mandibles shorter than head, rounded laterally, apices acute; upper surface with external margin slightly raised. Internal margin of mandibles with a medial strong tooth, with apex blunted directed inwards. Antenna with 9 antennomeres, with scape as long as the remaining antennomeres combined; club of last three antennomeres, which are flatter and broader than the remaining antennomeres of the funicle. Mentumsubsquare ( Fig. 9), slightly concave medially; anterior margin slightly bilobed; base smooth inclined forwards, with sides slightly elevated at mid-length. Surface smooth, with mixed large punctures and wrinkles. Thorax: pronotum subsquare with subparallel sides; posterior width less than anterior width; anterior margin almost straight, with a small, blunt median tubercle; posterior margin slightly bisinuate; anterior angles produced with anterior margin rounded; posterior angles largely rounded, with slightly crenulate margin. Superficial longitudinal medial groove on pronotal disc, with some large, circular punctures. Pronotal surface glossy with dense, small, superficial punctures, smaller along the anterior margin of prothorax and with patch of large, close punctures at sides. Long, golden setae directed upwards along the lateral pronotal margin. Scutellum long, wedgeshaped; surface smooth and glossy. Elytra about twice as long as pronotum. Base concave, subparallel sides, apex regularly rounded. Humeral angles acute, with a small tubercle. Elytral surface glossy, with 9 visible subparallel striae, the dorsal striae deeper than the lateral striae; each stria formed by a row of large punctures well separated from each other, except for the first stria, deeper than others and where the punctures are confluent. Stria 1 the only stria than reaches apex; stria 2 ending slightly before; stria 3 curved outwards, converging with stria 6; striae 4 and 5 converging at declivity. Interstriae much larger than striae, slightly convex and smooth, with a longitudinal, irregular row of small punctures. Elytral margins and apex strongly punctate, with short, erect, red setae. Wings present. Legs: thin and straight. Protibiae with strong, elongate, apical fork, slightly dilated at the point where it articulates a long, strong spur; lateral margin with 3‒4 small teeth, increasing in size distally. Mesotibiae with 4 long, acute apical teeth, and 2 apical spurs, 1 larger than the other; lateral margin with strong submedial tooth, preceded by 1‒2 smaller teeth; ventral lateral margin of mesotibiae with 2‒3 small teeth. Metatibiae with 4 acute apical teeth, and 2 long apical spurs, 1 larger than the other; lateral margin with strong submedial tooth. Prosternum subpentagonal in shape behind the coxae; surface with small punctures at anterior edge and with smaller punctures medially and at posterior edge. Metasternal disc smooth, with a long medial groove, deep at posterior edge; sides of metasternum with large and dense, horse-shoe shaped, superficial hollows. Abdomen: first abdominal ventrite with some large and dense, horse-shoe shaped, superficial hollows with a long, golden seta in the middle. Ventrites 2‒4 with some deep longitudinal depressions along the anterior margin, with short, golden setae emerging from the anterior margin; they are not present in the anteromedial area where there are 1‒3 irregular transversal rows of dense and deep punctures, each with a short seta, erected and directed downwards. Last ventrite with some punctures, which are smaller near the posterior margin. FIGURES 8‒11. Figulus lilliputanusWestwood.8, dorsal view of adult (scale bar: 5 mm); 9, mentum; 10, male genitalia, dorsal view (scale bar: 0.5 mm); 11, male genitalia, lateral view. Male genitalia: penis tubular ( Figs. 10–11), short and narrow, moderately curved in lateral view, strongly bent proximally close to articulation with basal piece, almost completely covered by a dorsal translucid extension of the basal piece. Permanently everted internal sac drawn out into a narrow colorless or yellow flagellum, slightly longer than the tegmen (basal piece + parameres), with two subsequent forkings in the basal half, tapering towards the apex ( Fig. 10). Parameres slightly shorter than basal piece, with marginal fringe of setae entirely enclosed in the cuticle (not represented in figure); ventrally the base of the parameres ends in a long spine-shaped apophysis articulated with basal piece ( Figs. 10–11). Basal piece ventrally largely membranous, dorsally sclerotized with a moderately large (about one-half of basal piece length), translucent cuticular area along the midline. Two long, slender struts run along basal piece, not reaching its base. Female.All external morphological characters as in male, except for sternites 2‒4 smooth and glabrous, without irregular, transverse punctation or short setae. Female genitalia: hemisternites elongate, broader anteriorly, narrower posteriorly; anteriorly broad up to about half of their length; outer edge notched; medial edge convex throughout length. Bursa copulatrix represented by a slight expansion of the wall of spermathecal duct near junction with the median oviduct. Spermathecal duct short, ending with a curve and long expansion, proximally colorless, distally pigmented. Intraspecific variation.The specimens examined do not show significant morphological variability, except for the body size (see “Measurements” section).
Additional material examined. Australia[state unknown]: 8 specimens, 29.IX.1929, E.Sutton Collection, donated Dec.1964( QMB); 1 specimen, ex Musaeo Ed. Brown, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, ex Musaeo W.Möllenkamp, Muséum Paris1952 Collection R.Oberthür ( MNHN); 2 specimens, det. A.M.Lea (MEL); 5 specimens, Nat. Mus. Victoria, COL-85173, COL-85174, COL-85175, COL-85176, and COL- 8517 ( NMVM); 1 specimen (GTC); 1 specimen, ex Musaeo Mniszech, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, Nouvelle Hollande, R.I.Sc.N.B. 21.418, Collection P.de Moffarts, 4764, Comp.au type, H.E.Bomans BMNH(E) 1999-248, 404764 (handwritten) ( NHM). AUSTRALIAN CAPITALTERRITORY: 1 specimen, Black Mountain, 1.II.1968, light trap, M.S.Upton( ANIC). NEW SOUTH WALES: 1 specimen, New South Wales, Macleay Museum Universityof Sydney2006 Australia, 11384, H.E.Bomans BMNH(E) 1999-248, 413884 (handwritten) ( NHM); 1 specimen, New South Wales, Nat. Mus. Victoria, COL-85784 ( NMVM); 1 specimen, Albury, 16.II.1970, R.Parrot, Collection R.E.Parrot ( MEL); 1Ƌ, Avoca Stationoff Pomonaroad 20 kmN of Wentworth, 13.VII.1993, COL-86865 (NMVM); 1 specimen, Ballimore, F.H.Taylor (ANIC); 1♀, 1 specimen, N Belmore Road, 3.2 kmSSW of Busbys Flat, 26.I. 1999, 170 m, at mv light, S.Watkins, S.Watkins Collectiondonated 2001 ( ANIC); 1Ƌ, Bungabbee State Forest, SEKyogle, 29.I.1997, mv light, S.G.Watkins, 97:199 Ƌ, S.G.Watkins Collection, donated 2001 ( ANIC); 1 specimen, Como, Dr. K.K.Spence, K.K.Spence Collection, K293062 (AMS); 1 specimen, Coonabarabran, XII.1929, H.J.Carter( ANIC); 1♀, Fortis CreekNational Park, Southboundary, N of Grafton, 4.I. 2000, 50 m, at mv light, S.Watkins Collectiondonated 2001 ( ANIC); 2ƋƋ, Fortis CreekNational Park, Southboundary, N of Grafton, 17.XII. 1998, 50 m, mv light, 98:837Ƌ and 98:838 Ƌ, S.Watkins Collectiondonated 2001 ( ANIC); 2 specimens, Maitland, 26.XII.1962, A NeboissCOL-85777 ( NMVM); 1 specimen, Mogriguy, Goonoo State Forest, 13.II.1981, R.H.M., R.H.Mulder Collection, K 293063 (AMS); 1 specimen, Mullaly, XII.1929, H.J.Carter( ANIC); 2 specimens, Sydney, Hyde Park, XII.1985, K 293074 (AMS); 2 specimens, Wagga, 1937, C.Oke, COL-85913 (NMVM); 2 specimens, Wagga, 1938, C.Oke, Presentedby C.G.Oke 1.IX.1948, COL-85883 ( NMVM). QUEENSLAND: 1 specimen, Queensland, Nat. Mus. Victoria, COL-85916 ( NMVM); 1 specimen, Queensland, Nouvelle Hollande, Fry Collection1905-100, BMNH(E) # 607642 ( NHM); 1 specimen, Blackdown Tableland, via Dingo, XII.1978, Maywald& Czechura( QMB); 1Ƌ, Boggomoss 15 infield, Taroom District, 25°28'S, 150°01'E, 12.XI.1996, P.Lawless, 191, QM Reg. No T 34260 (QMB); 1Ƌ, Boggomoss number 8 via Taroom, 25°27'S, 150°02'E, 12.XI.1996, Queensland Mus.survey, day hand collection, 0 39, QM Reg. No T 34259 ( QMB); 1Ƌ, 2 specimens, Boyne Island, via Gladstone, 23°55'S, 151°20'E, 1995, C.Q. University, 5022 ( QMB); 1Ƌ, Bribie Island, XII.1925, H.Hacker( QMB); 1 specimen, Brisbane, 29.X.1917and 1 specimen, 4.III.1920, H.Hacker (QMB); 2 specimens, Brisbane (QMB); 2 specimens, Brisbane, Mount Coot-tha, I.1966, C.OkeCOL-85890 ( NMVM); 1 specimen, Cape York, 3851 ( QMB); 6 specimens, Clermont, XII.1929, Dr. K.K.Spence, K.K.Spence Collection, K293070 K 293071, and K 293072 (AMS); 2 specimens, Cooktown, I.1971, J.G.Brooks, J.G.Brooks Bequest1976 ( ANIC); 1 specimen, Dawson District, Barnard Collection, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, Dawson River, Macleay Museum Universityof Sydney2006, 11383 (handwritten), H.E.Bomans1999-248, 411383 ( NHM); 1Ƌ, 2♀♀, Dunmore State Forest, 27°38'S, 150°58'E, 26.XII.1998, 98:883 Ƌ, mv light, S.Watkins Collectiondonated 2001 ( ANIC); 1 specimen, Garradunga Stone Creek, 15.I.1997, J.Hasenpusch, R.deKeyzer Insect Collection, K293325 (AMS); 1 specimen, 15 kmS of Greenvale, 9.I.1911, B.B.Howery( ANIC); 1Ƌ, Gurgeena Plateau, 25°27'S, 151°23'E, 8.VI. 1999, 360 m, open forest, 7741, G.B.Monteith& G.I.Thompson( QMB); 1 specimen, Hann River, 15°11'S, 143°52'E(GPS), 18.XII.1993‒ 14.I.1994, flight intercept trap, P.Zborowski, E.D. Edwards( ANIC); 1 specimen, 14 kmENE Heathlands, 11°41'S, 142°42'E, 8.XII.1992, at light rainforest, P. Zoborowski & W.Dressler (ANIC); 1 specimen, Iron Range, W Claudie Rg, 12°45'S, 143°14'E, 3.XII. 1985, 50 m, rainforest, stick brushing, G.B.Monteith, QM Berlesate No 690 (QMB); 1 specimen, same locality, 3.XII. 1985, 50 m, rainforest, Monteith& Cook( QMB); 1 specimen, Iron Range National Park, Gordon Creek( QMB); 1 specimen, Iron Range, 28.X.1981, J.Hasenpusch, R.deKeyzer Insect Collection, K293323 (AMS); 1Ƌ, Iron Range, Cape YorkPeninsula, VI‒VII.1977, G.B.Monteith (QMB); 1 specimen, Kuranda, 13.I.1987, R.deKeyzer Insect Collection, K293322 (AMS); 1 specimen, Maryborough, E.W.Fischer, ex Collection Australian Museum (ANIC); 2 specimens, Moreton Bay, K 15076(handwritten), K 293064 (AMS); 1 specimen, Moreton Bay, ex Musaeo Ed. Brown, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, Nipps, COL-85915 (NMVM); 1 specimen, Normanton, II.1982, R.deKeyzer Insect Collection, K293324 (AMS); 1 specimen, Rockhampton, ex Musaeo Ed. Brown, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, Rocky River, Silver Plains, E Coast, Cape YorkPeninsula, 6.I.1960, J.L.Wassel( ANIC); 2 specimens, Stanthorpe, E.Sutton, F.E.Wilson Collection, COL-85885 (NMVM); 5 specimens, Stanthorpe, 29.VII.‒4.IX.1924, E.Sutton Collection, donated Dec.1964( QMB); 2 specimens, Station Creek, N Queensland, 30.XII.1964, J.G.Brooks( ANIC); 1Ƌ, 2 specimens, Station Creek, 30.XII.1969, J.G.Brooks, J.G.Brooks Bequest1976 ( ANIC); 1 specimen, Toomba, 19°58'S, 145°34'E, 14‒16.XII.2007, Site 1, 390 m, 14652, mv light, vinescrub edge/paddock, Monteith& Wright, QMReg. No T147162 ( QMB); 1 specimen, Townsville, K.K.Spence, K.K.Spence Collection, K 293069 (AMS); 3 specimens, 9 kmENE of Mount Tozer, 12°43'S, 143°17'E, 5‒10.VII.1986, rotten log, T.Weir& A.Calder( ANIC); 1Ƌ, Woodstock, 3.VII.1992, B.P.Moore( LBC). SOUTH AUSTRALIA: 2 specimens, South Australia, ex Macleay Museum Universityof Sydney2006 Australia( GTC, LBC); 1Ƌ, South Australia, R. Mus. Hist. Nat. Belg., I.G. 12.595 ( IRSNB); 1 specimen, Adelaide, BMNH(E) # 607646 (NHM); 2 specimens, Adelaide Hills, V.1930, C.Oke, Presentedby C.G.Oke 1.IX.1948, COL- 85892 ( NMVM); 1 specimen, Adelaide, ex Musaeo Ed. Brown, Muséum Paris1952 Collection Oberthür( MNHN); 1 specimen, Adelaide, Griffith 0 4 (ANIC); 1 specimen, Adelaide, K 28038(handwritten), K 293073 (AMS); 1 specimen, Barossa, A.H.Elston, A.H.Elston Collection, K 293075 (AMS); 1 specimen, Buckaringa Gorge, Flinders Ra. C. 30 kmNNW of Quorn, 19.XII.1985, Reid, Gullan& Lewis, at light trap( ANIC). VICTORIA: 4ƋƋ, Ballarat, VIII.1937, C.Oke, Presentedby C.G.Oke 1.IX.1948, COL-85875 and COL-85884 ( NMVM); 1Ƌ, 1♀, Hattah, C.Oke COL-85914 (NMVM); 2 specimens, Lake Hattah, XI.1924, C.Oke, COL-85785 and COL-85786 ( NMVM); 1 specimen, Mount Hope, 27.IX.1947, C.Oke, Presentedby C.G.Oke 1.IX.1948, COL-85881 ( NMVM); 2 specimens, Piangil, 20.VII.1919, C.Oke, COL-85912 ( NMVM); 1 specimen, Sea Lake, III.1912, J.C.G., COL- 85894 ( NMVM); 1 specimen, Sea Lake, Goudie, 583, COL-85901 ( NMVM). WESTERNAUSTRALIA: 1 specimen, Bakers Hill, 27.III.1973, light trap, D.Weeldenburg (ANIC); 4 specimens, Blackwood River( ANIC); 2 specimens, Bridgetown, L.J.Newman, F.E.Wilson Collection, COL-85850 (NMVM); 1 specimen, Champion Bay, ex CollectionV-de-P-, Sig. K. vV. Steinwehr Dec./ Nov.1935, Museum Koenig Bonn, 5368, H.E.Bomans BMNH(E) 1999-248, 405368 (handwritten) ( NHM); 3ƋƋ, Gingin, H.J.Carter Collection P. 20.4.22., COL-85891 (NMVM); 1 specimen, Gray River, 6.III.1983, at light, K. & E.Carnaby( ANIC); 1 specimen, 2 mi. NW Margaret River, 23.XII.1970, mv lamp, G.A.Holloway& H.Hughes, K 293085 (AMS); 1 specimen, Nannup, 26.XII.2004, D.Lord, R.deKeyzer Insect Collection, K293326 (AMS); 1Ƌ, Pipehead Dam, 15 milesSSE of Armadale, 26.I.1967, M.S.Upton (ANIC); 3 specimens, Wilga, I.1973and 1 specimen, 21.IV.1973, K. & Carnaby (ANIC).
Distribution. Australia: Australian CapitalTerritory, New South Wales, Queensland, Western Australia, Victoria, South Australia. The species is widely distributed in Australia, with the exception of the NorthernTerritory. Natural history data.According to label data, adults were collected thoughout the year, mostly from December to April. Most of them were collected at light trap with mercury vapor light; a single specimen was captured in a flight intercept trap (Hann River, XII.1993– I.1994). Few other specimens were found in rotten logs, in vine scrub. The elevation ranges from 50 to 390 m. The larvae and host plant(s) are unknown.
Remarks.Concerning Figulus lilliputanusWestwoodand Figulus clivinoidesThomson, Bomans (1986: 9)wrote that: “- Il n’est pas absolument certain que ces deux insects soient synonymes. L’examen des types permet de constater que le premier est nettament plus petit que le second, 6,2 et 7,5 mmpour 8 mm. De plus lilliputanusa été découvert en Australie méridionale et occidentale, et clivinoidesou du moins l’espèce la plus grande, en Nouvelle Galle du Sud et au Queensland” [It is not sure that the two species are synonyms. Examination of types shows that the first is clearly smaller than the second, 6.2 and 7.5 mminstead of 8 mm. Moreover F. lilliputanushas been discovered in southern and western Australia, and F. clivinoides, or at least the largest species, in New South Wales and Queensland]. We dissected the male labeled "Type" of F. clivinoides, and we compared the genitalia with those of F. lilliputanus. We do not see any significant difference between F. clivinoidesand F. lilliputanus, both in external morphology and in genitalia characters ( e.g., in the flagellum shape). Therefore we consider F. clivinoidesas a junior synonym of F. lilliputanus( new synonymy). We found in the collection of NHM a specimen labeled “Tasmania” (A.Simson leg., 8116, 4765, H.E.Bomans), but this locality is most probably erroneous, since we have never seen any other specimens of this species from Tasmania.
1324963449
1929-09-29
1964-12
1929-09-29
QMB
E. Sutton Collection
Australia
Australia
8
455
8
1324963455
[460,1432,799,824]
Musaeo Ed. Brown & Museum Paris & Collection Oberthur
Australia
MNHN
8
455
1
MNHN
1324963454
[151,1228,836,861]
Musaeo W. & Museum Paris & Collection R.
Australia
MNHN
8
455
1
MNHN
1324963462
NMVM
Nat. Mus.
Australia
Victoria
8
455
5
Victoria
1324963444
Musaeo Mniszech & Museum Paris & Collection Oberthur
Australia
MNHN
8
455
1
MNHN
1324963447
BMNH, NHM
R. I. Sc. & Collection P. & Comp. & H. E. Bomans
Netherlands
Hollande
8
455
1
holotype
1324963452
1968-02-01
ANIC
M. S. Upton
Netherlands
Black Mountain
8
455
1
AUSTRALIAN CAPITAL TERRITORY
holotype
1324963465
[943,1371,1015,1040]
Netherlands
NEW SOUTH WALES
8
455
1
NEW SOUTH WALES
holotype
1324963461
BMNH, NHM
Macleay Museum University & Sydney & H. E. Bomans
Australia
New South Wales
8
455
1
New South Wales
holotype
1324963464
[518,1017,1087,1112]
Nat. Mus
Australia
New South Wales
8
455
1
New South Wales
holotype
1324963443
[1033,1404,1087,1112]
NMVM
Australia
Victoria
8
455
1
Victoria
holotype
1324963442
1970-02-16
MEL
R. Parrot & Collection R. E. & Pomona
Australia
Avoca Station
Albury
8
455
1
Victoria
holotype
1324963450
1999-01-26
ANIC
N Belmore Road & S. Watkins & S. Watkins Collection
Australia
Busbys Flat
8
455
2
1
Victoria
holotype
1324963445
1997-01-29
ANIC
Bungabbee State Forest, SE & S. G. Watkins & D, S. G. Watkins Collection & Como & Dr. K. K. & Spence, K. K. & Spence Collection, K
Australia
Bungabbee State Forest
8
455
1
Victoria
holotype
1324963460
[337,993,1303,1328]
1929-12
ANIC
H. J. Carter
Australia
Coonabarabran
8
455
1
Victoria
holotype
1324963458
2000-01-04
ANIC
S. Watkins Collection
Australia
Fortis Creek National Park
South
Fortis Creek
8
455
1
1
Victoria
holotype
1324963466
1998-12-17
ANIC
D, S. Watkins Collection
Australia
Fortis Creek National Park
South
Fortis Creek
8
455
1
Victoria
holotype
1324963446
1962-12-26
NMVM
A Neboiss
Australia
Maitland
8
455
2
Victoria
holotype
1324963451
1981-02-13
R. H. M., R. H. Mulder Collection
Australia
Goonoo State Forest
Mogriguy
8
455
1
Victoria
holotype
1324963448
[245,816,1484,1509]
1929-12
ANIC
H. J. Carter
Australia
Mullaly
8
455
1
Victoria
holotype
1324963456
1985-12
Australia
Hyde Park
Sydney
8
455
2
Victoria
holotype
1324963457
1948-09-01
NMVM
C. Oke & C. G. Oke
Australia
Presented
Wagga
8
455
2
Victoria
holotype
1324963453
[701,1048,1556,1581]
Australia
QUEENSLAND
8
455
1
QUEENSLAND
holotype
1324963459
[1063,1324,1556,1581]
Australia
Queensland
8
455
1
Queensland
holotype
1324963463
NMVM
Australia
Victoria
8
455
1
Victoria
holotype
1324963488
BMNH, NHM
Queensland & Fry Collection
Netherlands
Hollande
8
455
1
holotype
1324963479
1978-12
QMB
Maywald & Czechura & Boggomoss
Netherlands
Blackdown Tableland
-25.466667
Taroom District
1246
150.01666
Dingo
8
455
1
holotype
1324963472
1996-11-12
QMB
Mus.
Netherlands
Queensland
8
455
1
Queensland
holotype
1324963471
QMB
Gladstone & C. Q. University
Iceland
-23.916666
Island
1254
151.33333
8
455
2
holotype
1324963468
[602,1431,1772,1797]
1925-12
QMB
H. Hacker & Brisbane
Iceland
Island
8
455
1
holotype
1324963475
1966-01
NMVM
C. Oke
Iceland
Mount Coot-tha
Brisbane
8
455
2
holotype
1324963474
[814,1267,1844,1869]
QMB
Iceland
Cape York
8
455
1
Cape York
holotype
1324963489
1929-12
Dr. K. K. & Spence, K. K. & Spence Collection, K
Iceland
Clermont
8
455
6
Cape York
holotype
1324963467
1971-01
ANIC
J. G. Brooks & J. G. Brooks Bequest
Iceland
Cooktown
8
455
2
Cape York
holotype
1324963491
Barnard Collection & Museum Paris & Collection Oberthur
Iceland
Dawson District
8
455
1
MNHN
holotype
1324963469
NHM
Macleay Museum University & Sydney & H. E. Bomans
Iceland
Dawson River
8
455
1
MNHN
holotype
1324963482
1998-12-26
ANIC
S. Watkins Collection
Iceland
9
456
-27.633333
Dunmore State Forest
1235
150.96666
8
455
2
2
MNHN
holotype
1324963478
1997-01-15
J. Hasenpusch & Insect Collection, K
Iceland
Garradunga Stone Creek
9
456
1
MNHN
holotype
1324963480
1911-01-09
ANIC
B. B. Howery & Gurgeena Plateau
Iceland
-25.45
Greenvale
1246
151.38333
9
456
1
MNHN
holotype
1324963481
1999-06-08
QMB
G. B. Monteith & G. I. Thompson
Iceland
-15.183333
Hann River
1286
143.86667
9
456
1
MNHN
holotype
1324963477
1994-01-14
ANIC
P. Zborowski & E. D. Edwards
Iceland
-11.683333
Heathlands
1295
142.7
9
456
1
MNHN
holotype
1324963484
1985-12-03
QMB
Monteith & Cook
Iceland
-11.683333
Heathlands
1295
142.7
9
456
1
MNHN
holotype
1324963486
[317,1016,439,464]
QMB
Iron Range National Park
Iceland
Gordon Creek
9
456
1
MNHN
holotype
1324963476
1981-10-28
J. Hasenpusch & Insect Collection, K & Iron Range
Iceland
Iron Range
9
456
1
Cape York
holotype
1324963470
1987-01-13
Insect Collection, K
Iceland
Kuranda
9
456
1
Cape York
holotype
1324963473
Iceland
Moreton Bay
9
456
2
Cape York
holotype
1324963487
Musaeo Ed. Brown & Museum Paris & Collection Oberthur
Iceland
Moreton Bay
9
456
1
MNHN
holotype
1324963490
1982-02
Insect Collection, K
Iceland
Normanton
9
456
1
MNHN
holotype
1324963483
Rockhampton & Musaeo Ed. Brown & Museum Paris & Collection Oberthur
Iceland
MNHN
9
456
1
MNHN
holotype
1324963485
[586,1167,692,717]
Silver Plains, E
Iceland
Silver Plains
Rocky River
9
456
1
Coast
holotype
1324963507
1960-01-06
ANIC
J. L. Wassel
Iceland
Cape York
9
456
1
Cape York
holotype
1324963503
1924-07-29
1964-12
1924-07-29
QMB
E. Sutton Collection
Iceland
Stanthorpe
9
456
5
Cape York
holotype
1324963496
1964-12-30
ANIC
Station Creek, N & J. G. Brooks
Iceland
Station Creek
9
456
2
Queensland
holotype
1324963494
1969-12-30
ANIC
J. G. Brooks & J. G. Brooks Bequest
Iceland
Station Creek
9
456
2
Queensland
holotype
1324963493
2007-12-14
2007-12-16
2007-12-14
QMB
Site & Monteith & Wright, QM
Iceland
1
-19.966667
Toomba
1269
145.56667
9
456
1
Queensland
holotype
1324963499
1986-07-05
1986-07-10
1986-07-05
ANIC
T. Weir & A. Calder
Iceland
-12.716666
Mount Tozer
1292
143.28334
9
456
3
Queensland
holotype
1324963504
[900,1432,943,968]
1992-07-03
LBC
B. P. Moore
Iceland
Woodstock
9
456
1
Queensland
holotype
1324963502
[151,578,980,1005]
Iceland
SOUTH AUSTRALIA
9
456
2
SOUTH AUSTRALIA
holotype
1324963492
GTC, LBC
Macleay Museum University & Sydney
Australia
South Australia
9
456
1
South Australia
holotype
1324963509
[323,1154,1015,1040]
IRSNB
R. Mus. Hist. Nat. Belg.
Australia
South Australia
9
456
1
South Australia
holotype
1324963495
1930-05
1948-09-01
1930-05
NMVM
C. Oke & Presented & C. G. Oke
Australia
Adelaide Hills
9
456
2
South Australia
holotype
1324963506
[351,1432,1087,1112]
Adelaide & Musaeo Ed. Brown & Museum Paris & Collection Oberthur
Australia
MNHN
9
456
1
MNHN
holotype
1324963516
1985-12-19
ANIC
Flinders Ra. C. & Reid & Gullan & Lewis
Australia
Buckaringa Gorge
9
456
1
MNHN
holotype
1324963512
1937-08
1948-09-01
1937-08
NMVM
C. Oke & Presented & C. G. Oke
Australia
Ballarat
9
456
1
VICTORIA
holotype
1324963511
1924-11
NMVM
C. Oke
Australia
Lake Hattah
9
456
2
VICTORIA
holotype
1324963501
[274,1432,1303,1328]
1947-09-27
1948-09-01
1947-09-27
NMVM
C. Oke & Presented & C. G. Oke
Australia
Mount Hope
9
456
1
VICTORIA
holotype
1324963514
[151,900,1340,1365]
1919-07-20
NMVM
C. Oke
Australia
Piangil
9
456
2
VICTORIA
holotype
1324963515
1912-03
NMVM
H.J.Carter
Australia
Sea Lake
9
456
1
VICTORIA
holotype
1324963500
[363,1068,1375,1400]
NMVM
Goudie
Australia
Sea Lake
9
456
1
VICTORIA
holotype
1324963498
Bakers Hill
Australia
WESTERN AUSTRALIA
9
456
1
WESTERN AUSTRALIA
holotype
1324963497
[960,1410,1412,1437]
ANIC
Australia
Blackwood River
9
456
4
WESTERN AUSTRALIA
holotype
1324963508
1935-11
BMNH, NHM
Sig. K. & Museum Koenig Bonn & H. E. Bomans
Australia
Collection
Champion Bay
9
456
1
vV
holotype
1324963510
[275,1059,1556,1581]
1983-03-06
ANIC
K. & E. Carnaby
Australia
Gray River
9
456
1
vV
holotype
1324963513
1970-12-23
NW Margaret River & G. A. Holloway & H. Hughes
Australia
Margaret River
9
456
1
vV
holotype
1324963505
2004-12-26
D. Lord & Insect Collection, K & Pipehead Dam
Australia
Nannup
9
456
1
vV
holotype