Aphodius chapini Hinton, 1934: 189 Blackwelder 1944: 212 Aphodius ataenioides Hinton, 1938: 123 Blackwelder 1944: 212 Aphodius ( Blackburneus ) ataenioides Dellacasa 1988: 229 Aphodius ( Blackburneus ) chapini Dellacasa 1988: 351 Aphodius ( Blackburneus ) castanescens Petrovitz, 1973: 145 Dellacasa 1988: 233 Gonaphodiellus ( s. l. ) chapini Skelley et al. 2007: 4 Systematic revision of Gonaphodiellus taxa, with description of two new genera and fourteen new species (Coleoptera: Scarabaeidae: Aphodiinae) Dellacasa, Marco Dellacasa, Giovanni Gordon, Robert D. Insecta Mundi 2012 2012-04-06 2012 230 1 41 (Hinton, 1934) Dellacasa & Dellacasa & Gordon 2012 Hinton 1934 [192,744,200,219] Insecta Aphodiidae Gonaphodioides Animalia Coleoptera 18 17 Arthropoda species chapini comb. nov.  ( Fig. 73-77)      Aphodius chapini Hinton, 1934: 189;  Blackwelder 1944: 212.     Aphodius ataenioides Hinton, 1938: 123;  Blackwelder 1944: 212( new synonymy).    Aphodius( Blackburneus) ataenioides;  Dellacasa 1988: 229.    Aphodius( Blackburneus) chapini;  Dellacasa 1988: 351.     Aphodius( Blackburneus) castanescens Petrovitz, 1973: 145;  Dellacasa 1988: 233( new synonymy).    Gonaphodiellus( s. l.) chapini;  Skelley et al. 2007: 4.    Typelocality.Porto Bello [Puerto Bello, Panama].   Typerepository. United StatesNational Museum, Washington, DC, U.S.A.( typeexamined).   Redescription.Length 4.0-4.5 mm; oblong, moderately convex, shiny, glabrous. Chestnut brown; clypeal margin, pronotal sides and elytral apex somewhat paler; legs chestnut brown; antennal club yellowish. Head with epistome feebly convex on disc, superficially microreticulate, almost evenly subregularly coarsely punctured; punctation distally confusedly superficial, somewhat sparser on disc; clypeus angulately sinuate at middle, obtusely round at sides, very finely bordered, edge slightly upturned, glabrous; genae obtusely round, not ciliate; faintly protruding from the eyes; frontal suture finely impressed, front evenly, coarsely, regularly, not closely punctured throughout. Pronotum transverse, moderately convex, evenly, almost coarsely, almost regularly, not closely punctured; punctation somewhat sparser on disc; lateral margins feebly arcuate, finely bordered, edge glabrous; hind angles obtusely truncate; truncation slightly inwardly sinuate; base bisinuate, more or less strongly crenulate by a contiguous belt of coarse punctures. Scutellum elongate, almost smooth, distinctly striolate along lateral margins. Elytra oval elongate, moderately convex, not denticulate at shoulder; on disc, striae fine, superficially punctured, faintly crenulate; on preapical declivity, striae superficial, wider and not crenulate; on disc, interstriae convex, smooth, very shiny and with extremely fine sparse punctures; preapically, interstriae almost flat, superficially microreticulate, weakly shiny and more distinctly punctured. Hind tibiae upper spur shorter than first tarsal segment; first tarsal segment as long as following three segments combined. Male: aedeagus Fig. 76-77. Female: fore tibiae spur relatively more slender, elongate and somewhat outward curved.   Material examined.   BRAZIL: Acre, Nueva York( 1 ex., male, typeof  Aphodius (Blackburneus) castanescens Petrovitz, 1973, MHNG).   COSTA RICA: Alajuela, Penas Blancas, IX-X.1986, leg. E. Cruz, M.T. ( 1 ex., DCGI);  Cartago, Turrialba,  28.II.1980, leg. Howden H.& A. (37 exx., CNCI);  idem,  06.VI.1951, leg. Cartwright O. L.( 1 ex., USNM);  idem, m 650,  26.II.1980, leg. Howden H.& A. (5 exx., CNCI, DCGI);  idem,  26.II.1980, leg. Howden H.& A., dung mega trap, Florenciasur woods ( 1 ex. CNCI);  idem, m 600,  19-21.V.1979, leg. Howden H.& A., CATI (33 exx., CNCI);  Heredia Pr., La Selva, nr. P.to Viejo, m 50,  19.II.1980, leg. Howden H.& A. (12 exx., CNCI);  LimonP., Valle de la Estrella Pandora,  14.II.1984, leg. Howden H.& A. ( 1 ex., CNCI);  Punt., Monteverde, m 140,  24.V.1979, leg. Howden H.& A. (8 exx., CNCI);  Puntarenas Prov., Rincon de Osa,  10-15.VIII.1966, leg. Peck B.( 1 ex., CNCI);  San JoséP., 2 Km S Colon, m 1100,  01-16.II.1984, leg. Howden H.& A. ( 1 ex., CNCI).   ECUADOR: Pich[incha]. Prov., Rio Palenque,  24.II.1976, leg. Nealis V.(6 exx. CNCI, DCGI);  Pich[incha]. Prov., 16 Km E Santo Domingo, Tinalandia, m 680,  04.V-25.VII.1985, leg. Peck S.& J., malaise-FIT, rain forest ( 1 ex., CNCI);   MEXICO: Oaxaca: Ixtlán de Juárez, 17°20’07"N– 96°29’02"W,  19.IV.1993( 1 ex., DCGI).   PANAMA: Canal Zone,  12-13.IX.1975, leg. Young P., human feces trap( 1 ex., DCGI);  Canal Zone, Ciricito,  03.V.1930( 1 ex., USNM);  Cerro Campana, m 3000,  02.VIII.1970, leg. Howden H.& A. (5 exx., CNCI);  Chiriqui Prov., Cerro Pelota, m 1500,  01-14.VII.1982, leg. Gill B.(2 exx., DCGI, CNCI);  idem, 4 Km N Sta. Clara,  VIII.1952, leg. Gill B.(2 exx., CNCI);  Chiriqui Prov., 2 Km N Sta. Clara, Hartmann’s Finca, 8°51’N– 82°46’W, m 1300,  24- 25.V.1977, leg. Howden H.& A. (2 exx., DCGI, CNCI);  Chiriqui Prov., 2 KmN Sta. Clara, m 1300, W. Hartman’s Finca,  30-31.V.1977, leg. Howden H.& A. (8 exx., CNCI);  idem, 8°51’N– 82°36’W, m 1300,  30- 31.V.1977, leg. Howden H.& A. ( 1 ex., CNCI);  Chiriqui Prov., Hartmann’s Finca,  04-07.VII.1977, leg. Morris& Wappes, feces baited pitfall trap(2 exx., MCGA);  Colon Prov., Santa Rita Ridge, m 270,  10- 11.VI.1977, leg. Howden H.& A. (9 exx., CNCI);  Panama Prov., Cerro Azul,  13-18.II.1996, leg. Turnbow& Wappes, swine feces baited trap(2 exx., RHTC);  idem,  12-13.V.1996, leg. Wappes, Huether& Morris( 1 ex., ISEA);  idem,  21-27.V.1996, leg. Wappes, Huether& Morris( 1 ex., MCGA);  Panama Prov., Cerro Jefe,  30.VI.1997, leg. Turnbow R.(2 exx., DCGI, RHTC);  Panama[Prov.], Chepo-Carti Rd, m 400,  VI.1982, leg Gill B., flight interception trap(2 exx., CNCI);  Panama Prov., K 8-13 El Llano-Carti Rd,  10-13.V.1996, leg. Wappes, Huether& Morris( 1 ex., MCGA);  idem,  21-24.V.1996, leg. Wappes, Huether& Morris(2 exx., MCGA);  Panama Prov., Pipeline Rd K1-12,  15-22.II.1999, leg. Wappes J. E., feces baited pitfall trap( 1 ex., CNCI); [ Panama Prov.], Porto Bello[= Puerto Bello],  11.II.1911, leg. Schwarz E. A., ex horse manure ( 1 ex., male, typeof  Aphodius chapini Hinton, 1934, USNM);  Pan[ama]. [Prov.], Potrellos[=Potrerillos?],  II.1934( 1 ex., male, typeof  Aphodius ataenioides Hinton, 1938, USNM).   PERU: Pucalpa,  11.VII.1992, leg. Peña L. E.(2 exx., DCGI).   VENEZUELA: Carab[obo]., Borbureta, P. Cabella, m 200,  13.X.1985, leg. Bordon C.( 1 ex., DCGI);  Miranda, Los Altos, m 1300,  24.XII.1983, leg. Bordon C.( 1 ex., DCGI);  Miranda, El Amarillo, S. Antoniode los Altos, sector Los Moralitos, 10°21’N– 66°56’W, m 50,  20.II.2002, leg. Zunino M.(2 exx., DCGI);  Parque Nacional de Rancho Grande, m 1300,  19.VII.1964, leg. Bordon C.( 1 ex., DCGI).   Distribution. Brazil, Costa Rica, Ecuador, Mexico( Oaxaca), Panama, Peru, Venezuela.   Bionomics.Poorly known; the majority of specimens examined were collected in late winter and early spring.   Remarks.The new synonymies are established with the thorough study of the typesof involved taxa. 3766785455 MHNG Brazil Nueva York 18 17 1 Acre 3766785308 DCGI E. Cruz Costa Rica Penas Blancas 18 17 1 Alajuela 3766785554 [419,1220,1301,1320] 1980-02-28 CNCI Howden H. Costa Rica Turrialba 18 17 1 Cartago 3766785357 1951-06-06 USNM Cartwright O. L. Costa Rica Turrialba 18 17 1 Cartago 3766785513 [632,1436,1333,1352] 1980-02-26 CNCI, DCGI Howden H. Costa Rica Turrialba 18 17 1 Cartago 3766785424 [192,1274,1365,1384] 1980-02-26 CNCI Howden H. & Florencia Costa Rica Turrialba 18 17 1 Cartago 3766785380 1979-05-19 1979-05-21 1979-05-19 CNCI Howden H. Costa Rica Turrialba 18 17 1 Cartago 3766785553 1980-02-19 CNCI La Selva & Howden H. Costa Rica Viejo Pr. 18 17 1 Heredia 3766785530 1984-02-14 CNCI Valle de la Estrella Pandora & Howden H. Costa Rica Limon 18 17 1 Limon 3766785543 [559,1436,1461,1480] 1979-05-24 CNCI Howden H. Costa Rica Monteverde Punt. 18 17 1 3766785562 [192,1113,1493,1512] 1966-08-10 1966-08-15 1966-08-10 CNCI Peck B. Costa Rica Rincon de Osa 18 17 1 Puntarenas 3766785339 1984-02-01 1984-02-16 1984-02-01 CNCI Km S Colon & Howden H. Costa Rica San Jose 18 17 1 San Jose 3766785378 1976-02-24 CNCI, DCGI Nealis V. Ecuador Pich Rio Palenque Prov. 18 17 1 3766785391 1985-05-04 1985-07-25 1985-05-04 CNCI Km E Santo Domingo & Peck S. Ecuador Pich Tinalandia Prov. 18 17 1 3766785406 1993-04-19 DCGI Mexico 17.335278 Ixtlan de Juarez 21 -96.48389 18 17 1 Oaxaca 3766785490 1975-09-12 1975-09-13 1975-09-12 DCGI Young P. Panama Canal Zone 18 17 1 3766785317 [777,1356,1653,1672] 1930-05-03 USNM Panama Ciricito Canal Zone 18 17 1 3766785417 1970-08-02 CNCI Howden H. Panama Cerro Campana 18 17 1 3766785523 1982-07-01 1982-07-14 1982-07-01 DCGI, CNCI Gill B. Panama Cerro Pelota 18 17 1 Chiriqui 3766785384 1952-08 1982-07-14 1982-07-01 CNCI Km N Sta. Clara & Gill B. Panama Cerro Pelota 18 17 1 Chiriqui 3766785467 1977-05-24 1977-05-25 1977-05-24 DCGI, CNCI Howden H. & A. Panama 8.85 Chiriqui Prov. 1300 -82.76667 18 17 1 Chiriqui 3766785355 1977-05-30 1977-05-31 1977-05-30 CNCI W. Hartman's Finca & Howden H. Panama Sta. Clara Km 18 17 1 Chiriqui 3766785492 1977-05-30 1977-05-31 1977-05-30 CNCI Howden H. Panama 8.85 Sta. Clara 1300 -82.6 Km 18 17 1 Chiriqui 3766785363 1977-07-04 1977-07-07 1977-07-04 MCGA Morris & Wappes Panama Hartmann's Finca 18 17 1 Chiriqui 3766785485 1977-06-10 1977-06-11 1977-06-10 CNCI Howden H. Panama 19 18 Santa Rita Ridge 18 17 1 Colon 3766785434 1996-02-13 1996-02-18 1996-02-13 RHTC Turnbow & Wappes Panama Cerro Azul 19 18 1 Panama 3766785343 1996-05-12 1996-05-13 1996-05-12 ISEA Wappes & Huether & Morris Panama Cerro Azul 19 18 1 Panama 3766785364 [326,1119,264,283] 1996-05-21 1996-05-27 1996-05-21 MCGA Wappes & Huether & Morris Panama Cerro Azul 19 18 1 Panama 3766785450 1997-06-30 DCGI, RHTC Turnbow R. Panama Cerro Jefe 19 18 1 Panama 3766785528 1982-06 CNCI Gill B. Panama Chepo-Carti Rd 19 18 1 Panama 3766785305 1996-05-10 1996-05-13 1996-05-10 MCGA Wappes & Huether & Morris Panama Panama Prov. 19 18 1 Panama 3766785514 1996-05-21 1996-05-24 1996-05-21 MCGA Wappes & Huether & Morris Panama Panama Prov. 19 18 1 Panama 3766785393 1999-02-15 1999-02-22 1999-02-15 CNCI Pipeline Rd K & Wappes J. E. Panama Panama Prov. 19 18 1 Panama 3766785369 1911-02-11 USNM Schwarz E. A. Panama Porto Bello 19 18 1 Panama 3766785401 1934-02 USNM Panama Potrellos Pan 19 18 1 3766785452 1992-07-11 DCGI Pena L. E. Peru Pucalpa 19 18 1 3766785501 1985-10-13 DCGI Borbureta, P & Bordon C. Venezuela Cabella Carab 19 18 1 3766785536 1983-12-24 DCGI Bordon C. Venezuela Los Altos 19 18 1 Miranda 3766785519 2002-02-20 DCGI Zunino M. Venezuela 10.35 Los Moralitos 1298 -66.933334 Altos 19 18 1 Miranda 3766785399 1964-07-19 DCGI Bordon C. Venezuela Parque Nacional de Rancho Grande 19 18 1