The dung beetle fauna of the Big Bend region of Texas (Coleoptera: Scarabaeidae: Scarabaeinae) Edmonds, W. D. Insecta Mundi 2018 2018-07-27 642 1 30 7ZMF9 Schaeffer Schaeffer 1906 [192,585,747,771] Insecta Scarabaeidae Copris Animalia Coleoptera 14 13 Arthropoda species arizonensis McCleve and Kohlmann 2005 [1134,1242,1311,1334] Mammalia Muridae Neotoma Animalia Rodentia 14 13 Chordata genus   Fig. 64–69   Diagnosis.Black, surface lustrous, sides of body more-or-less parallel. Length 13–22 mm. Male ( Fig. 64–66) – Head of large individuals bearing posteriorly curved horn, tip of which approaches bifurcated median process of pronotum; flanks of pronotum each with large, acute, forward-directed, blade-like process separated from apically bifurcated median process by broad, declivitous concavity (in smaller individuals horn and processes progressively attenuated, Fig. 65). Female Fig. 67–69) – Head with short, erect horn, apex widened, scoop-like; pronotum convex except for pair of blunt tubercles, one on each side near anterior angle and separated by thick transverse ridge. Matthews (1961)provides a full re-description of this species.   Big Bend collection sites(altitudinal range: 1375–1800 m).   Brewster Co.: [1] Alpine, Sul Ross State Universitycampus, 30°22′04″N 103°38′52″W, 1375 m;  [2]* Big Bend National Park, Basin area, 29°16′04″N 103°17′39″W,  1660 m(May);  [3]* Alpine, Sunny Glen, 30°22′41″N 103°45′06″W,  1445 m(Jun).   Jeff Davis Co.: [1]* Davis Mountains State Park, 30°35′43″N 103°56′5″W, 1540 m(Jun, Aug);  [2] *  Davis MountainsResort, 30°37′30″N 104°05′30″W,  1800 m(Jun–Aug).   Surface activity.Nocturnal.  Habitat.Montane, juniper-pinyon woodlands and arroyos in association with the wood rat,  Neotoma albigula(and possibly also  N. mexicanaBaird).   Comments.This is by far the rarest dung beetle in the Big Bend. All examined specimens from the region were collected at lights.  Copris arizonensisis also known from various locations in the mountains of southeastern Arizona and adjacent areas of southwestern New Mexicoand northern Chihuahua, Mexico[ Matthews 1961; McCleve and Kohlmann 2005; Warner 1990].) Figure 64depicts a specimen from southeastern Arizona (Dragoon Mts., Cochise Co.) exhibiting maximum expression of male armament, which I have not observed in Big Bend specimens ( Fig. 65). The occurrence of  C. arizonensisin scattered higher mountainous areas of northern Chihuahuaand the desert southwest is doubtless the result of shrinking and vertical isolation of suitable, high-elevation habitat during post-Pleistocene desertification of the region. 2575871655 [232,1391,1132,1158] Brewster Co. 30.367779 Sul Ross State University 20 -103.64777 Alpine 14 13 1 2575871674 1660 29.267778 Big Bend National Park 20 -103.29417 Basin area 14 13 1 2575871697 1445 30.378056 Alpine 20 -103.75166 Sunny Glen 14 13 1 2575871686 [232,1387,1228,1254] Jeff Davis Co. 30.595278 Davis Mountains State Park 20 -103.93472 Davis Mountains State Park 14 13 1 2575871682 Mountains Resort 1800 30.625 Davis Mountains Resort 20 -104.09167 Davis Mountains 14 13 1