The millipede genus Antichiropus (Diplopoda: Polydesmida: Paradoxosomatidae), part 3: species of the Pilbara bioregion of Western Australia Car, Catherine A. Harvey, Mark S. Hillyer, Mia J. Huey, Joel A. Zootaxa 2019 2019-06-17 4617 1 1 71 Car Car & Harvey & Hillyer & Huey 2019 [151,462,837,864] Diplopoda Paradoxosomatidae Antichiropus Animalia Polydesmida 41 42 Arthropoda species nimbus sp. nov.   ZooBank LSID: urn:lsid:zoobank.org:act: F4C915A3-8686-47D0-A2CA-E977E47DF52D   Type material examined.   Australia:  Western Australia: holotypemale, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  89 kmWSW of Nullagine,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T144574).  Paratype: 1 male, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T109073, GenBank accession number 12S,  MK735758; 28S,  MK735818; COIII,  MK735940)  Other material examined.   Australia:  Western Australia: 1 female, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October 2010–  3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T109072);  1 male, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T109074, GenBankaccession number 28S,  MK735819);  2 males, 4 females, 5 juveniles, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T109075);  1 male, Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T 112610);  1 male, Newman, ca.  130 kmNW of Newman, BFS2, Site08, 22°17’14.20”S, 119°01’48.20”E,  6 June–16 July 2010, wet trap09, M.K. Curran( WAMT  121031, GenBankaccession number 12S,  MK735761; 28S,  MK735821; COI,  MK735887; COIII,  MK735942; CytB,  MK735993);  2 females Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T146719);  1 male, 4 females, 15 juveniles(some damaged), Cloudbreak, 22°18’52.62”S, 119°18’07.04”E,  26 October–3 December 2010, wet pitfall trap, S. Whiteand N. Dight( WAM T146720);  2 juveniles(damaged), Newman, ca.  130 kmNW of Newman, BFS2, Site08, 22°17’14.20”S, 119°01’48.20”E,  6 June–16 July 2010, wet trap09, M.K. Curran( WAM T146721).   Diagnosis.Gonopod:  Antichiropus nimbusCar,  n. sp.may easily be separated from other Pilbara species by its distinctive, elaborate second femoral process (fp1) that appears to hug the femorite.   Description. Male holotype: Bodyca. 12 mmlong; midbody ring ca. 1 mmwide, with distinct, smooth waist, prozonite and metazonite of similar widths.  Colour(in alcohol) brown Fig 24A) leg colour brown. No paranota ( Fig 24B).  Sterniteswithout obvious processes/tubercles, sternal lamella squat, broad and rounded. Leg coxal processes absent. Anterior spiracles at midbody ovoid, flat.  Headsmooth, with no sculpturing; frons smooth, with few setae; face narrow, maximum width ca. 3xthe distance between antennal sockets; sockets separated by ca. 2xwidth of socket. Antennaeshort, reaching to posterior edge of collum, relatively robust, antennomeres all similar shape, apical antennomere slightly stouter.  Collumca. 1xlength of head ( Fig 24A).  Gonopodof medium length, reaching ring 5; coxa (C) robust, squat, shorter than femorite, with very slight ridge on anterior surface; prefemur (PF) robust and broad, with almost triangular prefemoral lip; femorite (F) short, slightly curved, broadening near apex; main femoral process (MFP) long, robust, curved, pointed ca. 1/3 solenomere length; second femoral process (fp1) long, asymmetrically pointed, appearing toothed in lateral view, curling round femorite at right angles to femorite; prolongation of femorite apex (prof) absent; solenomere (S) long, forming>1 loop/circle, similar width along length; solenomere tip short, asymmetrically pointed; solenomere process (sp1) ca. halfway along solenomere, pointed, two-pronged ( Figs 24C–F).  Female:Similar to male, but stouter (lateral view), with shorter, more slender legs (WAM T109075).   Distribution.This species is known only from the Cloudbreak area, WSW. of Nullagine in the Pilbara region ( Fig 33).   Etymology.The species is named for the location in which it was collected (Latin, noun,  nimbus, cloud). 2382464073 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T144574 1 Western Australia holotype 2382464072 2010-10-26 2010-12-03 2010-10-26 WAM, MK, COIII S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T109073 1 1 Western Australia paratype 2382464076 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T109072 1 1 Western Australia 2382464074 2010-10-26 2010-12-03 2010-10-26 WAM, MK S. White & N. Dight & GenBank Australia -22.314615 Cloudbreak 1 119.301956 41 42 T109074 1 1 Western Australia 2382464075 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T109075 11 4 5 2 Western Australia 2382464071 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T112610 1 1 Western Australia 2382464070 2010-06-06 31-12-10 2010-06-06 WAM, MK, COI, COIII M. K. Curran & GenBank Australia -22.287277 Site 1 119.03006 Newman 41 42 1 1 Western Australia 2382464079 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T146719 2 2 Western Australia 2382464077 2010-10-26 2010-12-03 2010-10-26 WAM S. White & N. Dight Australia -22.314615 Cloudbreak 1 119.301956 41 42 T146720 20 4 15 1 Western Australia 2382464078 2010-06-06 2010-07-16 2010-06-06 WAM M. K. Curran Australia -22.287277 Site 1 119.03006 Newman 41 42 T146721 2 2 Western Australia