Classification, Natural History, and Evolution of the Subfamily Peloniinae O (Coleoptera: Cleroidea: Cleridae). Part IX. Taxonomic revision of the New World genus Muisca S Opitz, Pitz Pinola Weston Linzer biologische Beiträge 2018 2018-07-27 50 1 587 653 (GORHAM, 1883) [178,372,344,365] Insecta Cleridae Muisca GBIF Animalia Coleoptera 34 621 Arthropoda species octonotata  D i a g n o s i s: Within the  M. octonotatagroup, two punctiform black spots are found at the center of the elytral disc in specimens of  M. octonotataand in those of M. testaspilota.However, in  M. octonotataspecimens the spots aligned in a straight line, across the elytron. The spots are more obliquely positioned in specimens of M. testaspilota. R e d e s c r i p t i o n: Size: Length 7.5 mm; width 2.1 mm. Form: As in Fig. 145. Color: Light castaneous, except each elytron with 4 black spots, 2 near the humeral angle, 2 at middle of disc. Head: Basal funicular antennomeres filiform, distal funicular antennomeres progressively shorter towards capitulum, capitulum about as long as combined length of funicular antennomeres, capitular antennomeres 9 and 10 oblong triangular ( Fig. 21), antennomere 11 obovate; frons wider than width of eye (EW/FW 25/33). Thorax: Pronotum (Fig. 61) oblong (PW/PL 87/105), side margin with welldeveloped tubercle, disc subglabrous at middle; elytral asetiferous punctation striate to distal 2/3 rd(EL/EW 280/80). Abdomen: Pygidium subquadrate, distal margin sublinear; posterior margin of 6 thvisible sternite broadly concave; aedeagus phallobase wide in basal half, tapered to lobe apices, phallic apex ( Fig. 114) triangular. V a r i a t i o n: Size: Length 4.0-7.0 mm; width 2-3.0 mm. Other than body size, the available specimens are quite homogeneous. N a t u r a l h i s t o r y: Specimens were collected throughout the year at altitudes ranging from 700 to 2100 meters; some with a black light, but most in a Malaise trap. Montane cloud forests seem to harbor many of these beetles.  D i s t r i b u t i o n ( Fig.95): I examined 231 specimens: Costa Rica:  Provincia de Guanacaste, Cacao Biological Station, 10°55'38''N 85°27'7'',  11-VII-2000, on vegetation, J. Ashe, R. Brooks, Z. Falin;  Southeastslopes of Volcán Cacao,  21-29-V-1991,  1000-1400 m, collector not noted; idem,?-  V-1990; idem,?-  VII-1989-?-  III-1990, collector not noted; idem,  VI-1990, collector not noted; idem,?-  VII-1989, collector not noted; idem,  25-IX-11-X-1990,  1000-1400 m, C. Chaves;  Southwesternslope of Volcán Cacao,?-  V-1991, Elfin Rainforest, collector not noted; idem,  21-29- V-1991, F. Araya; idem,?-XI-XII-1990, A. Guadamuz; idem,?-XI-XII-1989, R. Blanco& C. Chavez; idem,?-  VII-1989, collector not noted;   9 kmS Sta. Cecillia,?-  VI-1991, P. Rios,  700 m;  Parque Nacional Guanacaste,  6-26-VI-1994,  1600 m, K. Taylor;   Provincia de Puntarenas, Las Cruces Biological Station, 8°47.14'N 87°52.58'W, 40-  V-2004, 1330 m, J. S. Ashe, Z. Falin, I. Hinojosa;  Monteverde,?-  XII-1990, E. Bello; idem, Campbell’sbullpen,  20-V-1993, Steve Lingafelter; idem,?-  IX-1992, 1040 m, Z. Fuentes; 23-VIII-13-IX-1995, 1670 m, E, Navarro; idem,  9-VII-1989. MV light, D. G. Furth;  Estación Biológica Monteverde, 10°19'40'' N 84°49'08''W,  9-VI-2001, 1540 m, at light, R. Anderson; Estación Biológica Las Alturas,  2 kmNE Alturas, 08°56'56'' N 82°50'01''W,  10-VII-1999, 1520 m, upper montane cloud forest, R. Anderson;  Monteverde Reserve,  23-VIII-1987, at light, H. & A. Howden;  Monteverde, Hotel Belmar,  6-IX-1998, C. W.& L. O’Brien;  Monteverde,  4-6-VI-1980, E. Giesbert;  Reserva de Monteverde,  27-29-V-1979,  5000 feet, J. M. & B. A. Campbell; idem,  4-6-VI-1980, E. Giesbert; idem,  27-II-1987, E. Giesbert;  Estación Pittier,  13-IX-1995, 1670 m, E. Navarro;  Las Tablas,  1.4 kmSW Cerro Gemelos,  28-VII-7-VIII-1995, 1670 m, E. Navarro;  Monteverde,  1-20-IX-1982,  1400 m, C. Nagano& M. Hayes; idem,  4-6-VI-1980, J. E. Wappes; idem,  24-VII-1990,  1500 m, black light, Meredith& Powell; idem,  20-V-1985, J. Doyen; idem,  19-V-1985,  1300 m, J. Chemsak; idem,  18-VIII-1987, 1500 H. & A. Howden; idem,  21-V-1979, H. & A. Howden; idem,  4-VI-1979,  1400 m, H. & A. Howden; idem,  24-V-1985, A. J. Gilbert; idem,  5-VII-1986, R. D. Cave;   Provincia de Cartago, Rio Aquiares, near Santa Cruz,  9 kmNW Turrialba,  15-V-1985,  1500 m, black light, J. Powel& O. Opler;  Parque Nacional Tapanti, 9°45'.41''N 83°47'.5''W,  19- VII-2000, 1150 m, J. Sahe, R. Brooks, Z. Falin;   Provincia de San José,  7 kmN Varablancaat Cascada de la Paz,  24-VI-1992, B. Ratcliffe, M. Jameson;   2.4 kmENE San Gerando de Rivas, Cloudbridge Res. Cloudbridge Northtrail, 9°28.87'N 83°34.17'W,  11-VI-2004,  2000-2100 m, J. S. Ashe, Z. Falin. Panamá:  Provincia de Chiriquí, 4 kmS Jorge Arauz Biological Station, collection date not noted, Steve Lingafelter;  Finca Hartmann, Trailcoffee plantation, 8°.87685N 82°.73639W,  2-V-2013, 4853 feet, N. Franz. S. Flynn, Y. Aguirre; idem, 8°.86141N 82°.22829W,  28-V-2013, 6932 feet, N. Franz.S. Flynn, Y. Aguirre; idem, 8°.86141N 82°.74322W,  30-V-2013, 4853 feet, N. Franz.S. Flynn, Y. Aguirre;  Fortuna Forest Reserve, Devil’s Chin, 4.2 kmS of STRI Ranger Station, 8°.69016N 82°.22829W,  25-V-2013, 4166 feet, N. Franz. S. Flynn, Y. Aguirre;  FortunaSTRI Station, 8°.73464N 82°.24015W,  26-V-2013, 3775 feet, S. Flynn;  Reserva La Fortuna, Estación Biológica, 08°43'18''N 82°14'17''W,  4-9-V-1999,  3900 feet, A. Gillogly& J. B. Woodley; idem,  27-28-VI-1996, 1150 m, Gillogly& Schaffer;   4 kmE Boquete, Valle Palo Alto, 08°43'18''N 82°14'17''W,  29-VII-1999, J. Schaeffer;  Volcánde Chiriquí, collection date not noted,  4000-6000 feet, Champion;  Las Lagunas,  4 kmW Hato del Volcán, 24-V-73, 1360 m, H. Stockwell;  Fortuna, 08°44'N 82°15'W,  22-I-1977, H. Wolda;   4.7 kmN Vallede las Minas,  3-8- VII-1997, J. Huether;  Hartmans Finca,  4-7-VII-1997, Wappes& Morris;  Totumas Lodge,  19-25- VI-2011,  1900 m, J. B. Heppner;  Mount Totumas, NW Volcán,  24-30-V-2014, 1900, J. B. Hepner;  Alto Lino, near Boquete,  18-V-1978, O’Briens& Marshall;  Fortuna, 82°15'W, 08°44'N,  16-V-1978, O’Briens& Marshall;  Las Lagunas,  22-VII-1976, W. E. Clark;   16 kmN Hato del Volcán,  10-VIII-2000, beating roadside vegetation, intermix of lianas & verdure, W. Opitz;  Las Lagunas,  27-IV-1973, G. Ekis;  La Fortuna area, hydrological trail,  9-VI-1995,  1100 m, wet montane cloud forest, R. S. Anderson;  Las Lagunas,  4 kmW Hato del Volcán, 08°51'N 82°36'W,  9-23-V-1977, 1720 m, H. & A. Howden;   Provincia de Veraguas, Alto Oledra, Santa Fe,  22-23- V-2014, J. B. Heppner;  Specimensare deposited in ACMT, AMNH, ASUC, BMNH, CSCA, CNCI, CMNH, EMEC, FSCA, JMLC, JNRC, MNHN, USNM, WFBM, and WOPC. 2849222470 2000-07-11 J. Ashe & R. Brooks & Z. Falin Costa Rica 10.927222 Cacao Biological Station 34 621 231 Provincia de Guanacaste 2849222414 [280,1026,1230,1252] 1991-05-21 1991-05-29 1991-05-21 Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222528 1990-05 1991-05-29 1991-05-21 Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222502 [321,756,1254,1276] 1989-07 1990-03 1989-07 Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222487 [767,1082,1254,1276] 1990-06 1990-03 1989-07 Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222313 [223,584,1278,1300] 1989-07 1990-03 1989-07 Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222518 [601,1082,1278,1300] 1990-09-25 1990-10-11 1990-09-25 Chaves Costa Rica 1200 Volcan Cacao Southeast 34 621 1 Guanacaste 2849222447 [223,964,1302,1324] 1991-05 Elfin Rainforest Costa Rica Volcan Cacao Southwestern 34 621 1 Guanacaste 2849222526 1991-05-21 1991-05-29 1991-05-21 Araya Costa Rica Volcan Cacao Southwestern 34 621 1 Guanacaste 2849222330 [399,727,1326,1348] 1991-05-21 1991-05-29 1991-05-21 A. Guadamuz Costa Rica Volcan Cacao Southwestern 34 621 1 Guanacaste 2849222316 1991-05-21 1991-05-29 1991-05-21 R. Blanco & C. Chavez Costa Rica Volcan Cacao Southwestern 34 621 1 Guanacaste 2849222346 [302,638,1350,1372] 1989-07 1991-05-29 1991-05-21 R. Blanco & C. Chavez Costa Rica Volcan Cacao Southwestern 34 621 1 Guanacaste 2849222490 [648,1082,1350,1372] 1991-06 Rios Costa Rica 700 9 km S Sta. Cecillia 34 621 1 Guanacaste 2849222377 [223,796,1374,1396] 1994-06-06 1994-06-26 1994-06-06 K. Taylor Costa Rica 1600 Parque Nacional 34 621 1 Guanacaste 2849222312 2004-05 J. S. Ashe & Z. Falin & Hinojosa Costa Rica 8.785666 Las Cruces Biological Station 13 -87.876335 34 621 1 Puntarenas 2849222403 [321,645,1422,1444] 1990-12 E. Bello Costa Rica Monteverde 34 621 1 Puntarenas 2849222306 1993-05-20 Steve Lingafelter Costa Rica Campbell's 34 621 1 Puntarenas 2849222417 [336,686,1446,1468] 1992-09 Fuentes Costa Rica Campbell's 34 621 1 Puntarenas 2849222495 [223,637,1470,1492] 1989-07-09 Furth Costa Rica Campbell's 34 621 1 Puntarenas 2849222455 2001-06-09 R. Anderson Costa Rica 10.327778 Estacion Biologica Monteverde 21 -84.818886 34 621 1 Puntarenas 2849222339 1999-07-10 R. Anderson Costa Rica 8.948889 Alturas 21 -82.83361 34 621 1 Puntarenas 2849222423 [324,903,1542,1564] 1987-08-23 Howden Costa Rica Monteverde Reserve 34 621 1 Puntarenas 2849222554 1998-09-06 C. W. & L. O'Brien Costa Rica Hotel Belmar Monteverde 34 621 1 Puntarenas 2849222424 [615,969,1566,1588] 1980-06-04 1980-06-06 1980-06-04 E. Giesbert Costa Rica Monteverde 34 621 1 Puntarenas 2849222372 1979-05-27 1979-05-29 1979-05-27 Campbell Costa Rica 1524 35 622 Reserva de Monteverde 34 621 1 Puntarenas 2849222555 [802,1082,214,236] 1980-06-04 1980-06-06 1980-06-04 Giesbert Costa Rica 1524 Reserva de Monteverde 35 622 1 Puntarenas 2849222486 [223,487,238,260] 1987-02-27 1980-06-06 1980-06-04 Giesbert Costa Rica 1524 Reserva de Monteverde 35 622 1 Puntarenas 2849222350 [497,939,238,260] 1995-09-13 E. Navarro Costa Rica Estacion Pittier 35 622 1 Puntarenas 2849222558 1995-07-28 1995-08-07 1995-07-28 E. Navarro Costa Rica Las Tablas 35 622 1 Puntarenas 2849222361 1982-09-01 1982-09-20 1982-09-01 C. Nagano & M. Hayes Costa Rica 1400 Monteverde 35 622 1 Puntarenas 2849222461 [527,824,286,308] 1980-06-04 1980-06-06 1980-06-04 Wappes Costa Rica 1400 Monteverde 35 622 1 Puntarenas 2849222399 1990-07-24 1980-06-06 1980-06-04 Meredith & Powell Costa Rica 1500 Monteverde 35 622 1 Puntarenas 2849222338 [536,795,310,332] 1985-05-20 1980-06-06 1980-06-04 Doyen Costa Rica 1500 Monteverde 35 622 1 Puntarenas 2849222352 1985-05-19 1980-06-06 1980-06-04 Chemsak Costa Rica 1300 Monteverde 35 622 1 Puntarenas 2849222309 [316,703,334,356] 1987-08-18 1980-06-06 1980-06-04 Howden Costa Rica 1300 Monteverde 35 622 1 Puntarenas 2849222348 [712,1027,334,356] 1979-05-21 1980-06-06 1980-06-04 Howden Costa Rica 1300 Monteverde 35 622 1 Puntarenas 2849222322 1979-06-04 1980-06-06 1980-06-04 Howden Costa Rica 1400 Monteverde 35 622 1 Puntarenas 2849222551 [579,857,358,380] 1985-05-24 1980-06-06 1980-06-04 Gilbert Costa Rica 1400 Monteverde 35 622 1 Puntarenas 2849222469 1986-07-05 1980-06-06 1980-06-04 Cave Costa Rica 1400 Monteverde 35 622 1 Puntarenas 2849222542 1985-05-15 J. Powel & O. Opler Costa Rica 1500 Santa Cruz Rio Aquiares 35 622 1 Cartago 2849222333 2000-07-19 J. Sahe & R. Brooks & Falin Costa Rica Parque Nacional Tapanti 35 622 1 Cartago 2849222357 1992-06-24 B. Ratcliffe & M. Jameson Costa Rica Cascada de la Paz 35 622 1 San Jose 2849222531 2004-06-11 J. S. Ashe & Z. Falin. & Steve Lingafelter Panama 2050 9.481167 Jorge Arauz Biological Station 13 -83.5695 Cloudbridge Res. Cloudbridge North 35 622 1 Chiriqui 2849222376 2013-05-02 N. Franz. S. Flynn & Y. Aguirre Panama Trail Finca Hartmann 35 622 1 Chiriqui 2849222560 2013-05-28 Franz. & Flynn & Y. Aguirre Panama Trail Finca Hartmann 35 622 1 Chiriqui 2849222477 2013-05-30 Franz. & Flynn & Y. Aguirre Panama Trail Finca Hartmann 35 622 1 Chiriqui 2849222425 2013-05-25 N. Franz. S. Flynn & Y. Aguirre Panama Fortuna Forest Reserve Ranger Station Devil's Chin 35 622 1 Chiriqui 2849222510 [223,966,646,668] 2013-05-26 S. Flynn Panama Station Fortuna 35 622 1 Chiriqui 2849222549 1999-05-04 1999-05-09 1999-05-04 A. Gillogly & J. B. Woodley Panama 1189 8.721666 Estacion Biologica 21 -82.23805 Reserva La Fortuna 35 622 1 Chiriqui 2849222303 [316,774,694,716] 1996-06-27 1996-06-28 1996-06-27 Gillogly & Schaffer Panama 1189 8.721666 Estacion Biologica 21 -82.23805 Reserva La Fortuna 35 622 1 Chiriqui 2849222378 1999-07-29 J. Schaeffer Panama 8.721666 Valle Palo Alto 21 -82.23805 35 622 1 Chiriqui 2849222305 Champion Panama 1524 Volcan 35 622 1 Chiriqui 2849222509 H. Stockwell Panama Las Lagunas 35 622 1 Chiriqui 2849222446 [324,763,766,788] 1977-01-22 H. Wolda Panama 8.733334 Fortuna 1300 -82.25 35 622 1 Chiriqui 2849222301 1997-07-03 1997-07-08 1997-07-03 J. Huether Panama Minas 35 622 1 Chiriqui 2849222379 [420,865,790,812] 1997-07-04 1997-07-07 1997-07-04 Wappes & Morris Panama Hartmans Finca 35 622 1 Chiriqui 2849222335 2011-06-19 2011-06-25 2011-06-19 J. B. Heppner Panama 1900 Totumas Lodge 35 622 1 Chiriqui 2849222471 2014-05-24 2014-05-30 2014-05-24 NW Volcan & J. B. Hepner Panama Mount Totumas 35 622 1 Chiriqui 2849222349 [300,825,838,860] 1978-05-18 O'Briens & Marshall Panama Boquete Alto Lino 35 622 1 Chiriqui 2849222498 1978-05-16 O'Briens & Marshall Panama 8.733334 Fortuna 1300 -82.25 35 622 1 Chiriqui 2849222536 [539,904,862,884] 1976-07-22 W. E. Clark Panama Las Lagunas 35 622 1 Chiriqui 2849222375 2000-08-10 Opitz Panama 16 km N Hato del Volcan 35 622 1 Chiriqui 2849222493 1973-04-27 G. Ekis Panama Las Lagunas 35 622 1 Chiriqui 2849222438 1995-06-09 Anderson Panama 1100 La Fortuna area 35 622 1 Chiriqui 2849222410 1977-05-09 1977-05-23 1977-05-09 H. & A. Howden Panama 8.85 Las Lagunas 1300 -82.6 35 622 1 Chiriqui 2849222434 2014-05-22 2014-05-23 2014-05-22 J. B. Heppner Panama Santa Fe Alto Oledra 35 622 1 Veraguas 2849222327 ACMT, AMNH, ASUC, BMNH, CSCA, CNCI, CMNH, EMEC, FSCA, JMLC, JNRC, MNHN, USNM, WFBM, WOPC Panama Specimens 35 622 1 Veraguas